The genus Macrocanace, known only from the sub-Antarctic islands of New Zealand, contains only two described species. They are M. littorea from Antipodes Island and M. australis (Hutt.) from Campbell Island. One new species was collected by the 1950 expedition, and this is described below under the name of M. antipoda. The three known species of the genus Macrocanace can be separated readily by the following key. Key to Species of Macrocanace Tonnoir and Malloch 1. Post verticals present; wings without clear spots 2 Post verticals absent; wings with clear spots littorea (Hutt.) 2. Second antennal segment with strong bristles; legs greyish black antipoda sp. nov. Second antennal segment without strong bristles; legs reddish brown australis (Hutt.) Macrocanace littorea (Hutton). (Text-Fig. 2, Figs. 6, 8, 9.) A more complete description than that given by either Hutton or Tonnoir and Malloch is given below. It is based on the type specimen and the 104 specimens collected by the expedition. Male and Female. General. A dark grey fly having brown wings with lighter areas. Body length 4-6 mm. Wing length 3.5-5.0 mm. Head. Antennae black; third segment disc-shaped and covered with fine black pile; second segment with one distinct bristle and accompanying short, stout hairs on distal margin of segment; first segment with few short hairs. Front half or over half width of head at vertex; predominantly black but bronze in some lights. Ocelli present. Ocellar triangle slightly raised above level of frontalia and with several small hairs. Post verticals absent. Parafrontals black merging to light grey at orbits. The three fronto-orbital bristles evenly spaced, anterior bristle the shortest, posterior bristle directed somewhat laterally over the eye. Few short black hairs lateral to fronto-orbitals. Two verticals. Face grey, sometimes reddish but light grey in some lights. Parafacials bare. Strong vibrissa present and two or occasionally one or three other strong bristles posterior to vibrissa and directed more dorsally. A row of small hairs extending from vibrissa and associated bristles to posterior ventral region of head, which is much swollen and where two or three long but slender bristles occur. Occiput grey, slightly concave dorsally and with small scattered hairs laterally. Bucca bare and reddish. Eyes bare. Proboscis reddish grey, palps darker. Thorax. Dark grey dorsally, lighter ventrally. Five indistinct brown longitudinal bands on mesonotum. Four strong dorsocentrals; one humeral, two notopleurals; one supra-alar; two post-alars; mesopleural bristle with two or three small hairs about it, a row of hairs extending from sternopleural ventrally and ending in four or five large hair-like bristles situated anteriorly to the mesothoracic coxa. Legs grey. Distinct apical bristles absent on tibia. Wings brown with clear areas. First longitudinal vein with strong setulose hairs commencing near junction of Rs and extending to costa. Halteres yellow with proximal region grey. Abdomen. General colour dark grey. Hairs on all segments but more numerous and larger on posterior segments.