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Volume 7, 1874

III.—Botany.

Art. XLVII.—On the Flowering Plants and Ferns of the Chatham Islands.

Plates XII—XV.

[Read before the Wellington Philosophical Society, 21st November, 1874.]

The present list of the Chatham Island plants is compiled from the collection in the herbarium of the Colonial Museum at Wellington. Nearly the whole of this collection has been made by Mr. H. H. Travers in the course of two visits paid to those islands in the years 1866 and 1871. A few specimens have been added to the collection at different times by other parties visiting those islands. With few exceptions all the plants in this list are also common to New Zealand, and will be found described in Hooker's Handbook of the New Zealand Flora, a work which is generally in the hands of every person taking any interest in the subject. Short descriptions will be added of the few not described there.

This list, however, is only published provisionally till Baron F. von Mueller, of Melbourne, who has undertaken the task, finds time from more important labours to furnish a complete analytical list of the whole.

An analysis of the present list shows the total number of plants indigenous to the Chatham Islands of the classes Dicotyledons, Monocotyledons, and Filical Alliance to be 205 species, belonging to 129 genera, and divided among the three classes as follows:—

Gen. Sp.
Dicotyledons 72 109
Monocotyledons 37 49
Ferns and Allies 20 47
129 205

Of this number 191 species are common to New Zealand. One species, Leucopogon richei, is also common to Australia, leaving only 13 species peculiar to the Chatham Islands. The latter are marked with a * in the list.

Dicotyledons.
Ranunculaceæ.
  • Ranunculus plebeius, Br.

  • rivularis, Banks and Sol.

  • acaulis, Banks and Sol.

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Cruciferæ.

  • Nasturtium palustre, DC. var. terrestre.

  • Cardamine hirsuta, Linn., var. a. debilis.

  • Lepidium oleraceum, Forst.

Violarieæ.

  • Viola cunninghamii, Hook. fil.

  • Hymenanthera latifolia, Endl. var. chathamica (F. v. Mueller, ‘Veg. Chatham Islands,’ but not described there).

A small pale-barked shrub-tree in bush. Leaves shortly petiolate, oblong ovate, acuminate at both ends, 2—3 inches long, 3/4—1 inch broad, serrate, strongly reticulated on both surfaces; coriaceous; flowers fascicled on the branches, peduncles decurved, 1–5th inch long, with two minute bracts near the middle, calyx-lobes obtuse, ciliolate, not spreading; petals linear oblong, twice as long as the sepals; anthers erect, connate into a fringed membranous tube. Fruit not seen.

This var. has also been collected by Dr. Hector in the Upper Rangitikei country, with large lanceolate leaves 3—4 inches long, sharply serrate, the venation on both surfaces strongly marked and very coriaceous.

Caryophylleæ.

  • Stellaria parviflora, Banks and Sol.

  • Colobanthus billardieri, Fenzl.

Malvaceæ.

  • Plagianthus divaricatus, Forst.

  • betulinus, A. Cunn. Nat. name—Houhere. 10—15 feet high.

Lineæ.

  • Linum monogynum, Forst.

Geraniaceæ.

  • * Geranium traversii, Hook. fil. (Pl. XIII. f. 2.)

Rhamneæ.

  • Discaria toumatou, Forst.

Sapindaceæ.

  • Dodonœa viscosa, Forst.

Anacardiaceæ.

  • Corynocarpus lœvigata, Forst.

Coriarieæ.

  • Coriaria ruscifolia, Linn.

Leguminosæ.

  • Sophora tetraptera, Aiton, var. grandiflora.

Rosaceæ.

  • Potentilla anserina, Linn., var. b.

  • Acœna sanguisorbœ, Vahl.

  • ” “ var. a.

In addition to the species, which is common both to New Zealand and the Chatham Islands, this well-marked var. has been collected by Mr. H. H. Travers on both visits. Characterised by its larger size, very silky habit, and

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long bristles on the calyx. It is often found among scrub, growing 6—8 feet high; stems rigid upright.

Crassulaceæ.

  • Tillœa moschata, DC.

Droseraceæ.

  • Drosera binata, Labill.

Halorageæ.

  • Haloragis alata, Jacq.

  • Myriophyllum elatinoides, Gaud.

  • pedunculatum, Hook. fil.

  • Callitriche verna, Linn.

Onagrarieæ.

  • Epilobium confertifolium, Hook, fil., var. a.

  • rotundifolium, Forst.

  • junceum, Forst.

  • pubens, A. Rich.

  • billardierianum, Seringe.

  • pallidiflorum, Sol.

Ficoideæ.

  • Mesembryanthemum australe, Sol.

  • Tetragonia implexicoma, Miquel, var. chathamica.

Widely spreading, or frequently climbing, frutescent leaves broadly rhomboid, occasionally broadly lanceolate, pedicels axillary, solitary, or geminate, downy, longer than the flower, lobes of the calyx unequal in size, semi-lanceolate, inside intensely yellow, stamens 12—16, styles 3—4, fruit 3—4-celled, spherical, red, 3—4-seeded, without wings or teeth. (F. Muell. l. c.)

Umbelliferæ.

  • Hydrocotyle asiatica, Linn.

  • moschata, Forst.

  • Crantzia lineata, Nutt.

  • Apium australe, Thouars.

  • Oreomyrrhis colensoi, Hook. fil.

  • * Aciphylla traversii, F. Muell. (Gingidii sp. F. Muell. l. c.; Hook. f. Handb. N.Z. Flora ii. 729).

Plant small, 10in. high, rather flaccid, smooth, leaves all radical, pinnate, 2in. long, numerous, sheath 2in. long, denticulate, 1/2in. broad, with membranous margins, leaflets 5—7, striate and pungent, scape stout, striate, bearing at top one bract and seven involucral leaves similar to the radical leaves, of 5—7 leaflets, and seven long, peduncled, densely capitate, globular umbels of fruit, peduncles unequal, 1 1/2—2 inches long, grooved, umbels lin. in diameter, compound, involucral, leaves of fruit, linear, subulate, 1/3in. long, carpels with five wings, the central dorsal broadest.

Aciphylla lyallii, Hook. fil.

monroi, Hook. fil.

Collected on both visits.

* Ligusticum dieffenbachii, Hook. fil. (Gingidium dieffenbachii, F. Muell. l. c. 17). Daucus brachiatus, Sieb.

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Araljaceæ.

Panax crassifolia, Dene & Planche. (Hedera crassifolia, A. Gray; F. Muell, l. c.)

This species, so remarkable in New Zealand for its varied leaf-form, shows in the Chatham Island plant another well marked variety.

In the varieties of New Zealand the leaves, whether one or three foliolate, are much elongated, narrow, and reflexed downwards for several years before flowering; afterwards they become shorter, broader, and erect.

In the variety under notice the leaves are erect from the first, before flowering being 3—10 inches long, 1—1 1/2 inches broad, and with a few deep serrations at the tips. Mixed with these in the earliest stage are a few small obcordate leaves, lin. long, 3/4in. broad. After the plant commences to flower, the leaves diminish in size and become entire. The fruit is more elongate in this variety than in the varieties of New Zealand, resembling most that of Panax lineare, Hook. fil., from Dusky Bay, South Island.

Corneæ.

Corokia buddleoides, A. Cunn.

Native names—Whakataka and Hokataka; grows 15—20 feet high.

Rubiaceæ.

Coprosma baueriana, Endl.

Nat. name—Karamu. This species is found in the bush as a small tree, 10—12 feet high.

Coprosma robusta, Raoul.

cunninghamii, Hook. fil.

propinqua, A. Cunn.

Nat. name—Mingimingi; 3—6 feet high, according to Mr. Travers, forming a considerable portion of the bush, and from its dense close growth almost impenetrable.

Coprosma acerosa, A. Cunn.

Compositæ.

Olearia angustifolia, Hook. fil., var. (Olearia operina, Hook f. l. c. 731; Eurybia operina, Forst. F. Muell. l.c.)

Leaves very variable in size, obovate-acute to ovate-obtuse, 1—3 inches long, 1/2—1 inch broad, very coriaceous. Peduncles none or short, with a few lax bracts. Flowers large, purple; single flowering, in succession over a lengthened period. Achenes ribbed, silky. Pappus robust; purple on the upper half, pale yellow on the lower. More closely allied to O. angustifolia than to O. operina.

*Olearia semidentata, Decaisne, (Pl. XIV).

Described by Mr. H. H. Travers as a beautiful shrub 1—3 feet high, having masses of dark purple flowers, and generally growing in damp places.

The leaves of young plants are very close set and narrow, differing in this

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Myosotidium Nobile, Hook.

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1.Veronica Chathamica. n.s. 2.Germanium Traversii, Hook.fil

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Olearia Semidentata, Decaisne.

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respect from young plants of O. operina and O. angustifolia, to both of which it is allied.

*Olearia traversii, F. Muell.

Nat. name—Akeake; 20—30 feet high, and sometimes 2 feet in diameter. This may be considered as the only valuable timber tree in the islands, being durable and not subject to the attacks of insects.

  • Brachycome sinclairi, Hook. fil.

  • Cotula coronopifolia, Linn.

  • *(Myriogyne) featherstonii, F. Muell.

  • lanata, Hook. fil.

  • Craspedia fimbriata, DC.

  • Gnaphalium bellidioides, Hook. fil.

  • filicaule, Hook, fil.

  • luteo-album, Linn.

  • involucratum, Forst.

  • collinum, Labill.

  • Erechtites prenanthoides, DC.

  • scaberula, Hook. fil.

  • quadridentatus, DC.

  • Senecio lautus, Forst.

  • " Forst., var. radiolatus (S. radiolatus, F. Muell., l. c.).

A broader leaved var. of this species than that from the Chatham Islands has since been collected by Mr. Travers on the Brothers Rocks, Cook Strait. The leaves are remarkably thick and fleshy, and only show membranous, as stated in the Handb. N.Z. Flora, p. 734, after drying.

  • *Senecio huntii, F. Muell. Nat. name—Rautine. (F. Muell., l. c.)

  • Taraxacum dens-leonis, Desf.

  • Sonchus oleraceus, Linn.

  • Wahlenbergia gracilis, A.DC.

  • Lobelia anceps, Thunb.

  • Pratia macrodon, Hook. fil.

Ericeæ.

*Cyathodes robusta, Hook. fil. (C. acerosa, Br., var. b. latifolia, Flora N.Z. i., 163, and Veg. Chatham Islands, 43).

  • Leucopogon richei, Br. (F. Muell., l. c. 45).

  • Pentachondra pumila, Br.

  • Dracophyllum scoparium, Hook. fil., Flora Antarct. i. 47 (D. urvilleanum, A. Rich, var. d. Handb. N.Z. FI. i. 182; D. latifolium, Hook. fil., var. ciliolatum, l. c. ii. 736).

A small tree 20—30 feet high. Specimens in Mr. Travers' collection show the young leaves flat, 8—9 inches long, and 3/4 inch broad, becoming reduced in size before flowering to 2—3 inches long, very narrow, involute, and ciliate on the lower margin in all stages; both sizes may be found on the same plant, the larger form on the lower branches only.

In the young state the fascicles of large broad leaves bear a considerable resemblance to those of D. latifolium, A. Cunn., and D. menziesii, Hook. fil., neither of which are found on the islands.

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This plant, from its large leaves when young and ciliate margins, has good claim to be re-established as a distinct species, though latterly it has only been considered as a var. of D. urvilleanum.

It is also found in the Lake District of Otago, where it was at first supposed to be D. menziesii, Hook. fil., never having been found in flower, although the leaf transition was noticed on small plants.

Dracophyllum rosmarinifolium, Forst.

A small shrub 3—8 inches high. In Mr. Travers' collection rooted specimens 3—4 inches high are found in flower. It agrees in every respect with the above species, except in sometimes having 1—4-flowered spikes, instead of 1—2-flowered spikes, as described by Dr. Hooker (Handb. N.Z. Flora, 183).

Myrsineæ.

*Myrsine chathamica, F. Muell.

Shrubby, erect, much branched. Leaves 1 1/2—2 1/2 inches long, obovateoblong, obtuse or emarginate, pale and reticulated on both surfaces (Dr. Hooker, Handb. N.Z. FI. 736). The fruit is described by Baron F. von Mueller as purplish, spherical, one-seeded, size of a large pea. To this may be added: Flowers in capitate lateral fascicles; calyx, 4-lobed; petals, 4, pale greenish, spotted red, ciliate on the margins, erect; stamens, 4, adherent to the petals; anthers large; stigma sessile, capitate. Dr. Hooker thinks this may be a large state of M. urvillei, and specimens of the latter, with large, nearly flat leaves, found in New Zealand, approach it very closely, except in size of fruit.

  • Myrsine nummularia, Hook. fil.

Primulaceæ.

  • Samolus littoralis, Br.

Gentianeæ.

  • Gentiana pleurogynoides, Griseb.

Boragineæ.

  • *Myosotidium nobile, Hook. (PI. XII.)

Convolvulaceæ.

  • Convolvulus sepium, Linn.

  • soldanella, Linn.

  • Dichondra repens, Forst.

Solaneæ.

  • Solanum aviculare, Forst.

  • nigrum, Linn.

Scrophularineæ.

  • Veronica salicifolia, Forst. (V. forsteri, F. Muell. var. salicifolia, l. c. 46.)

  • *dieffenbachii, Benth.

  • *chathamica, n. sp. (V. forsteri, F. Muell. var. elliptica, l. c. 46.)

  • PL XIII. f. 1.

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A. small prostrate rambling shrub. Branches wiry, 1/3 inch diameter, pubescent. Leaves spreading, irregular in size, sessile, 1/3—1 1/2 inches long, 1–5th—1/2 inch broad, obovate-oblong or ovate-oblong, acuminate, entire, flat, scarcely coriaceous. Racemes few or many, 1—1 1/2 inches long, subterminal at the ends of the branches and axillary to the uppermost leaves, orbicular or ovate. Flowers numerous, closely set; peduncles 1/2 inch long, pubescent; pedicels and lanceolate bracts equal in length, pubescent; sepals 1/8—1/6 inch long, linear lanceolate, ciliate; corolla large, dark purple, tube short; capsule imperfect. This beautiful shrub has been cultivated by Mr. Travers in his garden in Wellington, where the profusion of its dark purple flowers and prostrate habit has proved a most showy addition to those plants adapted for the ornamentation of rock-work or earth banks.

Verbenaceæ.

  • Myoporum lœtum, Forst.

Labiata.

  • Mentha cunninghamii, Benth.

Chenopodiaceæ.

  • Chenopodium triandrum, Forst.

  • glaucum, Linn.

  • Atriplex patula, Linn.

  • billardieri, Hook fil.

  • Salicornia indica, Willd.

Polygoneæ.

  • Polygonum minus, Huds., var. decipiens.

  • Muhlenbeckia adpressa, Lab.

Thymeleæ.

  • Pimelea arenaria, A. Cunn.

Euphorbiaceæ.

  • Euphorbia glauca, Forst.

Urticaceæ.

  • Urtica australis, Hook fil.

  • Parietaria debilis, Forst.

Piperaceæ.

  • Piper excelsum, Forst.

Monocotyledons.

Orchidaceæ.

  • Earina mucronata, Lindl.

  • Acianthus sinclairii, Hook. fil.

  • Corysanthes macrantha, Hook. fil.

  • Microtis porrifolia, Spreng.

  • Pterostylis banksii, Br.

  • banksii, Br., var. b.

  • micromega, Hook. fil.

  • Caladenia bifolia, Hook. fil. (Chiloglottis traversii, F. Muell., l. c., 51.)

  • Chiloglottis cornuta, Hook. fil.

  • Thelymitra longifolia, Forst.

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Irideæ.

  • Libertia ixioides, Spreng.

Naiadeæ.

  • Triglochin triandrum, Mich.

  • Potamogeton natans, Linn.

  • Ruppia maritima, Linn.

Liliaceæ.

  • Rhipogonum scandens, Forst.

  • Astelia cunninghamii, Hook. fil.

  • grandis, Hook. fil. (Trans. N.Z. Inst., IV., 245, Mr. Kirk on N.Z. Asteliads).

  • Phormium tenax, Forst.

Palmeæ.

  • Areca sapida, Sol.

Junceæ.

  • Juncus planifolius, Br.

  • bufonius, Linn.

  • novœ-zealandiœ, Hook. fil.

  • Luzula campestris, DC.

Restiaceæ.

  • Leptocarpus simplex, A. Rich.

  • *Sporadanthus traversii, F. Muell., n. gen. and n. sp.

Cyperaceæ.

  • Schœnus axillaris, Hook. fil.

  • Scirpus triqueter, Linn.

  • Eleocharis gracilis, Br., var. c. radicans.

  • acuta, Br., var. platylepis.

  • Isolepis nodosa, Br.

  • aucklandica, Hook. fil.

  • Desmoschœnus spiralis, Hook. fil.

  • Cladium glomeratum, Br.

  • Uncinia rupestris, Raoul.

  • Carex appressa, Br.

  • Forsteri, Wahl.

  • trifida, Cavan.

  • lambertiana, Boott.

Gramineæ.

  • Hierochloe redolens, Br.

  • Dichelachne crinita, Hook. fil.

  • Agrostis œmula, Br.

  • Arundo conspicua, Forst.

  • Danthonia semi-annularis, Br.

  • Deschampsia cœspitosa, Palisot.

  • Trisetum subspicatum, Palisot.

  • Poa breviglumis, Hook. fil.

  • foliosa, Hook. fil., var. a.

  • " " var. b.

  • Festuca littoralis, Br.

Cryptogamia.

Filices.

  • Gleichenia dicarpa, Br.

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Olearia Angustifolia, Hook.fil.var.

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  • Cyathea dealbata, Swartz.

  • medullaris, Swartz.

  • Dicksonia squarrosa, Swartz.

  • Hymenophyllum bivalve, Swartz.

  • dilatatum, Swartz.

  • javanicum, Spreng. (H. crispatum, Wallich, Synopsis Filicum.)

  • demissum, Swartz.

  • flabellatum, Labill.

  • Trichomanes reniforme, Forst.

  • venosum, Br.

  • Lindsaya linearis, Swartz.

  • Adiantum hispidulum, Swartz.

  • affine, Willd. (A. cunninghamii, Hook. fil., Synopsis Filicum.)

  • Hypolepis tenuifolia, Bernh.

  • distans, Hook.

  • Pellœa rotundifolia, Forst.

  • Pteris aquilina, Linn., var. esculenta.

  • scaberula, A. Rich.

  • incisa, Thunb.

  • Lomaria procera, Spreng., var. d.

  • dura, Moore.

  • lanceolata, Spreng.

  • discolor, Willd.

  • alpina, Spreng.

  • Asplenium obtusatum, Forst.

  • var. g. lucidum.

  • var. g. lucidum.

  • falcatum, Lam.

  • bulbiferum, Forst.

  • flaccidum, Forst., var. d.

  • Aspidium aculeatum, Swartz.

  • oculatum, Hook.

  • capense, Willd. (A. coriaceum, Swartz, Synopsis Filicum).

  • Nephrodium decompositum, R. Br., var. b.

  • Polypodium grammitides, Br.

  • punctatum, Thunb., var. b. (P. rugulosum, Labill., Synopsis Filicum).

  • pennigerum, Forst.

  • serpens, Forst. (P. rupestre, R. Br., Synopsis Filicum).

  • pustulatum, Forst.

  • billardieri, Br.

  • Schizœa fistulosa, Labill.

  • Ophioglossum vulgatum, Linn., var. b. costatum

  • Botrychium ternatum, Swartz, var. dissectum (B. cicutarium, Sw., Synopsis Filicum.

Lycopodiaceæ.

  • Lycopodium varium, Br.

  • billardieri, Spreng.

  • densum, Labill.

  • laterale, Br.

  • scariosum, Forst.

  • volubile, Forst.

  • Tmesipteris forsteri, Endl.

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Art. XLVIII.—Observations on the different Modifications in the Capsules
of Mosses, with, reference to the Dispersion of their Spores.

Plate XXII.

[Read before the Otago Institute, 12th October, 1874.]

The fructification of mosses consists of a variously shaped capsule, or sporangium, generally placed on a stalk, which is sometimes short, but generally much elongated (Plate XXII., fig. 13). Through the centre of the capsule rises a pillar of cellular tissue called the columella (fig. 15), and it is round the columella, between it and the wall of the capsule, that the spores, or seeds, are formed.

It is evident that if these spores were liable to be all blown away together by the first puff of wind that occurred after they were ripe, or to be all knocked out by the first drop of rain that fell upon the capsule, they would have but little chance of being widely dispersed. If also the spores escaped and were blown away in moist weather they could not travel far, for they would stick to the first leaf or stone that they were blown against. If, on the other hand, there were some means of holding the spores fast in wet weather, and letting them escape when the air was dry, and then only by a few at a time, so that some might be driven off in one direction, some in another, it is evident that they would then be scattered as far and as wide as possible.

But as mosses inhabit very different stations, some living among long grass or in swamps, others on exposed walls; some on trees, others under water; some in sheltered ditches, and others on mountain peaks, different combinations will be required by different species. For example, those species that grow among long grass require a long fruit-stalk to elevate the capsule above the grass and so let it be exposed to the wind; while this would be detrimental to those mosses that live in exposed situations, and these generally have the fruit-stalk so reduced in length that the capsule is buried in and protected by the perichætial leaves, unless the species possesses some other and more perfect way of preventing its spores being blown away; and I propose to offer a few observations on the different approaches towards perfection in attaining this object that are found in various genera and species of mosses.

I have already said that the spores are contained in a capsule placed on the top of a fruit-stalk. In by far the larger number of mosses this capsule is furnished with a lid, or operculum (fig. 14), which falls off when the spores are ripe, and the mouth of the capsule is then seen to be either naked (fig. 1), or furnished with numerous teeth and cilia, called the peristome (fig. 14). The

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fruit-stalk is sometimes straight, but often curved, so that the capsule is inclined or pendulous (fig. 13), with the mouth turned downward and thus protected from rain. In some mosses the same effect is obtained by the capsule itself being cernuous, while the fruit-stalk is straight.

In order to give some method to the remarks that follow I shall divide the mosses into (1) those in which the capsule alone is used; (2) those in which the peristome is used; and (3) those in which the columella is used, combined either with the operculum or with the peristome.

1. Mosses in which the capsule alone is used.

In the lowest forms, the family Phascei, the capsule decays and bursts open irregularly so that the spores are quite exposed, except in those species where the capsule is immersed in the perichætial leaves. But it must be remarked that in Phascum cuspidatum, which is the only species in which I have been able to make the observation, the perichætial leaves open when wet and close when dry, so that the protection thus afforded can be but slight. They have, however, very few spores in a capsule, Phascum alternifolium only having about sixteen, and P. serratum about one hundred, so that it would hardly be necessary to protect so few from being all blown away together. These mosses are all inhabitants of the plains, and grow on ditch banks and other sheltered situations; but there is another family, the Andreœeœ, which lives on exposed rocks on mountains, and yet has the capsule not much more complicated than in the humble Phascei. In this family when the spores are ripe the capsule divides longitudinally into four or more parts, the upper extremities of which adhere together. These parts bend downward when dry, and allow the spores to escape through the open fissures (fig. 8b), but on the air getting damp they straighten and close up the capsule again quite tight (fig. 8a), and thus prevent the spores being blown away in wet weather.

The first step towards a more complicated state of things is when the capsule, instead of bursting through decay, is furnished with an operculum, which falls off when the spores are ripe, thus leaving them to some extent protected by the vase-like form of the capsule. Some genera of mosses, such as Braunia, Physcomitrium, and Aulacopilum, have no further provision against all their spores being knocked out at once; but in others, as Hedwigia and Leptangium, * the capsule is immersed in the leaves. In the genus Sphagnum, when the capsule is ripe, the operculum and spores are said to be driven off with violence to a distance of six or seven inches. The next step is the formation of a flexible ring of very hygroscopic cells, called the annulus (fig. 14), interposed between the mouth of the capsule and the operculum, as in Œnectangium and Calomnion, which protects in some measure the spores from

[Footnote] * Leptangium, is said by Dr. Hooker to have the mouth of the capsule closed in by a membrane. I have had no opportunity of examining it personally.

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the wind; while in some species of Gymnostomum, as well as in Cadomnion, a further provision is made by the walls of the capsule being much thickened at the mouth, which reduces the size of the opening. In most of those genera in which the apparatus for protecting the spores is more perfect this annulus falls away * after it has performed its office of throwing off the operculum from the capsule, but in the genera just mentioned it is persistent.

In Œnectangium also the spores are very minute and numerous, which gives evident facilities for their far and wide dispersion

2. Mosses in which the peristome is used.

I come now to that large division of mosses which have a series of teeth, called the peristome, arranged round the mouth of the capsule. In its most complete state this peristome consists of an external row of sixteen conical teeth, and an internal folded membrane divided above into processes sometimes with cilia between each (fig. 7). The inner peristome is, however, variously modified, and is often altogether absent. The outer peristome consists normally of sixteen teeth, which are sometimes split half way down, as in Dicranum, Fissidens, etc. Sometimes they are divided to the base so as to make their number thirty-two, as in Trichostomum and Tortula. In rare cases also two teeth are joined together so as to reduce their number to eight, as in the tropical genus Octoblepharum, while occasionally they are reduced to four, as in Tetraphis.

The peristome also is very variously developed in different mosses. In some, such as Weissia, Didymodon, Conomitrium, etc., the teeth are short and fragile and soon break off. In Trichostomum, Encalypta, etc., they are stronger; while in most genera they are strong and answer admirably their purpose of preventing the too easy escape of the spores.

These mosses can be divided into five groups in each of which the peristome is used in a different way.

(a.) The first group, which is represented in New Zealand by the genera Dicranum, Dicranodontium, Campylopus, Racomitrium, Leucobryum, Grimmia, Ceratodon, Conostomum, and Symblepharis, have the peristome long and well formed, and the teeth stand nearly erect round the mouth of the capsule with

[Footnote] * Symblepharis is an exception.

[Footnote] † Brachyodus might be placed here, for its peristome being shorter than the annulus, it cannot in any way affect the dispersion of the spores.

[Footnote] ‡ I have not, however, been able to trace any connection between the abundance or rarity of a moss and the size of its spores. For instance, the spores of Hypnum cupressiforme and H. rutabulum are nearly twice the diameter of those of H. prœlongum and H. confertum, while H. triquetrum and H. purum hold an intermediate position.

[Footnote] † In Racomitrium aciculare the teeth are slightly hygroscopic, spreading when dry, but erect and slightly incurved when wet. I have noticed that in species a sticky substance is found amongst the spores, which is drawn out into lines like cobwebs between the teeth, and by this means the spores are often stuck on to the inner side of the peristome.

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their points touching or interlaced, so as to form a kind of cage over the mouth which protects the spores from the wind, but allows them to be blown away in small quantities at a time, and by high winds only. In Conostomum (fig. 5), which inhabits exposed rocks on mountains, this cage is further strengthened by the tips of the teeth cohering together, and this moss may be said to be the type of the group. All these mosses have strong persistent peristomes well calculated to resist the wind, consequently we find that many of them inhabit mountains or subalpine regions, and several grow upon exposed rocks, while in some species of Grimmia only is the capsule immersed in the perichætial leaves. The peristome however is not able to protect the spores from the wet, and to accomplish this the greater part have the mouth of the capsule inclined or bent downwards either through the fruit-stalk being curved, or the capsule being cernuous or drooping. I must here remark that the fruit-stalk of Campylopus is hygroscopic, being curved and burying the capsule among the leaves in dry weather, while when the air is moist standing more erect, an anomaly for which I can offer no explanation. In Seligeria recurvata the fruit-stalk straightens when dry.

(b.) In the second group, which is represented in New Zealand by Dawsonia superba and the genera Trichostomum and Tortula, the peristome consists of numerous filiform, slightly hygroscopic teeth, which are either distinct from one another, or united at the base into a membrane. In Trichostomum, and Dawsonia superba (fig. 11) the teeth are nearly straight, but in Tortula (fig. 9) they are twisted into a spiral, which in damp weather twists tighter and closer over the mouth of the capsule. The mosses in this group are preeminently inhabitants of the plains, their peristome being too weak and fugacious to withstand the boisterous winds of the mountains. Tortula possesses the strongest peristome, and consequently we often find it growing on walls and other exposed places, while Trichostomum and Didymodon generally grow on the ground or in sheltered hollows. All have the fruit-stalk elongated, and the capsule erect or nearly so.

(c.) In the third group the teeth are all more or less hygroscopic, opening when dry and closing over the mouth of the capsule when wet. In some, e.g., Phyllogonium (fig. 10) and Blindia, the movement is small and the teeth when wet never get beyond an erect position; in others, e.g., Fissidens and Cryphœa, they continue the movement until they have attained a horizontal position, and then curl their points inward; in others again, such as Orthotrichum, Zygodon (fig. 3), Eremodon and Fabronia, the movement is still further continued until their backs are adpressed against the outer sides of the capsule. In this group the peristome is beautifully adapted for preventing the spores being dispersed in wet weather, but there is nothing to prevent them being all blown away in one direction by the first strong wind, and we find therefore that they are chiefly inhabitants of the plains, and generally

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grow in forests and sheltered places. With the exception of some species of Orthotrichum the capsule is always exserted, and in Macromitrium it is sometimes much elongated. Fissidens is the only genus with the mouth of the capsule inclined or directed downward.

(d.) In the fourth group the peristome is double, the inner one being composed of a membrane more or less divided in the upper part. The outer peristome is hygroscopic, closing tight over the mouth of the capsule when moist, and spreading outward with the tips of the teeth incurved when dry. The inner peristome is not hygroscopic, but on the opening of the outer peristome the interior projections, or “trabeculæ”, as well as the points of the teeth, get entangled in the perforations and cilia of the inner peristome and drag it open, often quitting their hold with a jerk which spirts out the spores to some little distance, thus answering the same purpose as the elaters of the Hepaticœ. This group is a very large one and includes the New Zealand genera of Bryum, Cladomnion, Isothecium, Hypnum (fig. 7), Rhizogonium, Hypopterygium, Racopilum, Hookeria, and others. These mosses are chiefly inhabitants of plains and forests; in nearly all the species the capsule is inclined, while in some, e.g., Bryum and Mnium, it is pendulous (fig. 13), so that its mouth is directed straight downward. All have the fruit-stalk elongated except Fontinalis, a northern genus that lives in water. In this moss (fig. 6) the outer peristome is hygroscopic, and the inner is converted into a conical membrane regularly perforated with square holes, which prevents the spores being washed ont too quickly.

(e.) The fifth group consists of the genus Funaria alone, which stands apart from all other mosses as far as the apparatus for the dispersion of the spores is concerned. In this moss the peristome is double, the points of the teeth of the outer one being connected by a disc, which in time falls away. The inner consists of narrow teeth, which might almost be called cilia. Both are hygroscopic, and curl outward when dry after the rupture of the disc. The capsule also is pendulous on a hygroscopic fruit-stalk, which, however, when either curling or uncurling, always keeps the mouth of the capsule directed downwards, and thus protects it from the rain. We thus see that this moss combines the advantages of the first and third groups. At first its strong united peristome enables it to distribute its spores in small quantities to high winds only, while afterwards the mouth of the capsule opens wide, and allows, on the first dry day, all those spores that still remain to be blown away, and so prevents them from being wasted through failing to escape from the capsule. This is perhaps the most perfect apparatus possessed by any moss, and we cannot wonder at its almost cosmopolitan distribution.

3. Mosses in which the columella is used.

These mosses may be divided into two groups, which use the columella in connection with the operculum or the peristome respectively.

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(a.) In some European mosses, such as Gymnostomum curvirostrum, Pottia truncata (fig. 1), P, heimii, and Stylostegium cœspiticum, the operculum adheres to the columella, which elongates when the spores are ripe and lifts up the operculum. Some mosses even which have peristomes have also the operculum adhering to the columella, such as Dissodon hornschuchii, from Europe and North America; and in Climacium dendroides, also from Europe, the operculum is lifted up in dry weather, but when the air is moist the columella contracts again and closes up the mouth of the capsule.

(b.) In the European genus Cinclidotus (fig. 2), which lives in water, the columella is exserted and the filiform teeth of the peristome are twisted round it, thus forming a cage as in Fontinalis, but on quite a different principle. When the capsule gets dry or old the columella shrinks and breaks off the upper part of the peristome, thus liberating those spores that had previously failed to make their escape.

In Splachnum (fig. 12) the columella is large, exserted, and dilated at the top. In moist weather the teeth fold over the mouth of the capsule and touch the dilated apex of the columella, but in dry weather they bend back until they are adpressed against the outer side of the capsule.

In our species of Dawsonia the filaments are, as I have said, free, but in some Australian species these filaments, or cilia, are connected with the top of the columella. In Wardia, from the Cape of Good Hope, the peristome consists of an irregularly fissured membrane which adheres to the top of the columella, and Scouleria, from North America, has thirty-two laciniated teeth which are also connected with a process on the top of the columella, (Berkely).

But it is in the Polytrichaceœ that the columella is put to the greatest use for protecting the spores. In the genus Polytrichum (fig. 4) the top of the columella is spread out into a thin flat drum-like membrane that extends over the whole orifice of the capsule, and is connected at its edge with the mouth of the capsule by thirty-two or sixty-four short processes formed of many threads, like those in Dawsonia, soldered together, so that the spores can only escape through the apertures between these processes, and consequently only a few are likely to be blown out at a time. The genus Lyallia, from India, has a similar membrane over the mouth of the capsule, but at length the columella contracts within the capsule, and detaches a circular portion from the centre of the drum, thus releasing those spores which have failed to make their escape. As this membrane extends over the whole of the mouth of the capsule, it entirely excludes the rain when the capsule is erect, which position is therefore in this case better than an inclined one, and is that which nearly every species of the family assumes. The species are equally distributed between the plains and the mountains.

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Art. XLIX.—Description of a new Species of Senecio.

[Read before the Auckland Institute, 29th June, 1874.]

In the spring of 1868, while journeying up the course of the Kaueranga River, at the Thames Gold-fields, I observed in several localities a white-rayed Senecio, evidently differing in a marked degree from any known species. Unfortunately, the specimens collected on this occasion were accidentally lost before I had an opportunity of carefully examining them; and although I revisited the locality in 1870, it was at too advanced a period of the year to procure flowers. Last year, however, I was fortunate enough to again find the plant, not only in the district where I originally discovered it, but also in other portions of the gold-field. It may thus be described:—

Senecio myrianthos, n. sp. A slender sparingly branched shrub or small tree, with purplish black bark. Young leaves and branches covered with a thin buff tomentum. Leaves four to five inches long, membranous, ovate or ovate-lanceolate, acute or acuminate, generally unequal at the base, sharply and coarsely doubly serrate, when adult glabrous above, but covered beneath with a thin silvery-white closely appressed tomentum. Primary veins few, conspicuous on both surfaces, forming large areoles. Petioles one to two inches long. Panicles terminal, leafy, large, often over two feet long, but narrow for the length; peduncles and pedicels slender, with a short and thick covering of purplish brown glandular hairs. Ultimate bracts narrow subulate, or almost filiform. Heads numerous, campanulate, 1/3 inch long. Scales of the involucre about eight, scarious, linear obtuse, glabrous or nearly so. Ray florets four to six, ligules broad, very short, not 1/8 of an inch, white; disc florets about the same number, campanulate above the middle, five-toothed. Stigmas exserted; anthers slightly tailed. Pappus hairs in a single series, white, prominently denticulate, slightly thickened at the apex. Ripe achenia not seen.

Habitat.—Kaueranga River, Karaka, Tararu, and Puru Creeks; and other localities at the Thames. It is probably not uncommon throughout the Cape Colville peninsula.

A most charming plant, covered when in bloom with large panicles of deliciously sweet-scented white flowers. From the other species of Senecio, native to New Zealand, it can easily be distinguished by the slender habit; thin, membranous leaves; the peculiar indumentum on the branches of the inflorescence; the scarious, nearly glabrous scales of the involucre; and the short ligules to the ray florets. The beauty of the flowers will doubtless obtain it a place in both Colonial and European gardens.

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Art. L.—On the Fertilisation of Acianthus and Cyrtostilis.

[Read before the Auckland Institute, 1st June, 1874.]

1. Acianthus sinclairii.

In examining the fertilisation of this plant, we do not find contrivances so remarkable and unique as those that obtain in Pterostylis, or in other of the New Zealand Orchids; on the contrary, the mode employed is simple, and presents few features of interest. Yet, if the completeness of any method of fertilisation is to be judged of by the results obtained, as undoubtedly it should be, we must regard that of Acianthus as one of the most perfect of the many different modes in use among our Orchids.

The flowers, varying in number from one to twelve, are minute, and of an inconspicuous appearance. The lip, which is horizontally spread out in front of the flower, or slightly deflexed, is ovate-lanceolate in outline, and greatly concave, so as to form a kind of bucket. At its base it is furnished with two large glands, and the margins and point are also plentifully studded with minute fleshy papillæ. The column is somewhat curiously shaped. At first it is erect, but towards the summit suddenly arches over the lip, and is much thickened and expanded. The anther is terminal, two-celled, each cell possessing two pollinia, which are deeply bilobed, so as to resemble a horse shoe in shape. The stigma is a deep circular hollow situated just below the anther; and, by the arching of the upper part of the column, hangs directly over the lip. The rostellum is placed on its upper margin. It consists of two triangular projections, which at first are cellular, but ultimately resolve into masses of viscid matter, covered with an extremely delicate membrane. As the flower expands, the connection of these projections with the rest of the column becomes very slender, so that at last they can be detached by a comparatively slight touch, leaving the upper margin of the stigmatic chamber quite plane.

Long before the flower opens, each lobe of the anther splits gradually from base to apex, allowing the included pollinia almost to touch the upper part of the rostellate points. The pollinia then emit a number of excessively delicate thread-like projections, which reach the rostellum, and become firmly attached to it. So that, in a fully expanded flower, each set of pollen-masses is quite free from its anther cells, but they are firmly attached by their bases to their respective rostella, neither of which can be removed without bringing away the pollinia.

The glands at the base of the lip secrete nectar, which is stored up in the cavity just in front of them. From this circumstance alone we might surmise

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that the flowers would be frequently visited by insects, and a little observation soon shows this to be a fact. On a warm sunny day it is almost impossible to watch a bed of this Orchid for any length of time without seeing numerous Diptera flitting from flower to flower, busily engaged in robbing them of their sweets.

If we now call to mind the manner in which the column arches over the lip, we can easily see that an insect crawling into the flower to get at the supply of nectar can hardly avoid touching one of the points of the rostellum, ranging almost directly over it; if it did so, the delicate exterior membrane would be at once ruptured, and the viscid mass firmly glued to the insect's back. Thus, on withdrawing from the flower, the visitor would carry away with it not only the portion of the rostellum which it had touched, but also the attached pair of pollinia. These (from each pollinium being nearly subdivided into two) would form four little projections standing rigidly erect on the back of the insect; and consequently, when conveyed to another flower, can hardly fail to strike the overhanging stigmatic chamber, which is sufficiently viscid to detach a portion, at least, of the pollinia from the body of the insect, thus ensuring the fertilisation of the flower.

As I have several times seen insects remove the pollinia, and on one occasion also seen a pollen-mass left on the stigma, there can be little doubt that fertilisation is conducted on this plan. That insect aid is absolutely required is proved by the fact that the pollinia remain in their cells, and never reach the stigma, when the plant is covered up or allowed to expand its flowers in a room. But, under natural conditions, the flowers are so frequently visited that the pollinia are generally removed directly after the opening of the blossoms; while the large proportion of capsules produced is good evidence of the completeness with which the visitors perform their duties. Out of eighty-seven flowers, borne on fourteen plants, no less than seventy-one matured capsules, and of those that had failed to do so, many were imperfect ones situated at the summit of the panicle, and probably incapable of producing seed. Another set, from a different locality, had borne forty-seven flowers, of which no less than forty-four had ripened capsules.

The fact that almost every perfect flower produces a capsule, is in remarkable contrast to what occurs in several other genera of our Orchids. For instance, Pterostylis is fertilised on a plan much more complex, and the co-adaptation of the various parts of the flower is so complete that almost every insect that fairly enters the flower must remove the pollinia, which is not the case in Acianthus. Yet, from some reason, probably from the want of sufficient attraction, the flowers are comparatively seldom visited, and consequently few capsules produced. In my account of the fertilisation of this genus, (Transactions of the N. Z. Institute, Vol. V, p. 356.) I have

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estimated that about one quarter of the flowers produce capsules; but from subsequent observations I am now convinced that the number is much less. Corysanthes offers a case of imperfect fertilisation even more singular. In all the species the proportion of capsules produced is very small, and large patches can often be found that have failed to mature a single one. As an illustration, a bed of Corysanthes triloba, in a favourable situation for the visits of insects, expanded, during the last season, over two hundred flowers: yet of this large number only five succeeded in ripening capsules. We must be cautious, though, in assuming that the imperfect fertilisation of these plants is of much real disadvantage to them. In many districts Pterostylis trullifolia is quite as abundant as Acianthus; while the less general distribution of the species of Corysanthes is probably due to their organization not being so well adapted to a variety of conditions and habitats, rather than to the scarcity of seed produced. In their special localities they are often abundant.

2. Cyrtostylis oblonga.

The great resemblance that this plant bears to Acianthus, induced me to suppose that its fertilisation would be conducted on the same plan, and this appears to be the case. We find in Cyrtostylis, as in Acianthus, the lip horizontally spread out, secreting abundance of nectar; the column arching over it; the points of the rostellum hanging downwards, with the pollinia firmly fastened to their upper margins; together with other contrivances, all apparently co-ordinated, so that an insect, having once entered the flower, can hardly avoid attaching itself to the pollen-masses, and removing them on its departure.

On comparing the flowers of the two plants, we at once find a difference in the structure of the lip. In Acianthus this organ is concave, for the purpose of storing up nectar to serve as an attraction for insects: in Cyrtostylis it is narrow, and quite plain; but the same end is attained by allowing the nectar slowly to trickle down each side of the midrib. The secreting glands at the base of the lip are much smaller than in the former species, while the papillæ on the margins and points are totally wanting. The column agrees with that of Acianthus in most features, but is broadly winged on each side. This may be of use as a protection to the stigma, or perhaps the projections serve as guides for the proper withdrawal and insertion of the pollen-masses, No difference worth mention is found in the stigma, or rostellum; and the mode of attachment of the pollinia to the latter organ appears to be precisely the same in both plants. In the shape of the pollen-masses themselves, however, we find a marked divergence, for instead of being nearly subdivided, as in Acianthus, they are simply falcate in shape. They are laterally much compressed, and extremely friable.

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Notwithstanding the minuteness of the flowers, they are frequently visited by insects, chiefly minute species of Diptera. The pollinia, however, are not removed with the same regularity and precision as in Acianthus, nor is such a large proportion of capsules produced. I find, though, that specimens from some localities give very discordant results in this respect, although as a rule there can be no doubt that the proportionate number of capsules matured is much less thon in Acianthus.

I have made no observations on the fertilisation of the only other species of Cyrtostylis (C. rotundifolia) native to New Zealand. The difference between the two plants is so slight (if indeed it is sufficient to allow a specific distinction being maintained) that I can entertain no doubt but that, on investigation, the mode of fertilisation will be found to be the same for both species.

Art. LI.—On Pterostylis squamata in New Zealand.

[Read before the Auckland Institute, 27th July, 1874.]

For the discovery of this singular plant in New Zealand, we are indebted to the late Dr. Sinclair, so well known for his successful researches into the natural history of this country. Specimens collected by him in some locality near Auckland were forwarded to Sir W. Hooker at Kew, about twenty-four years ago; and accordingly we find the plant described in the first volume of the Flora Novæ-Zealandiæ. Nearly the same description is reproduced in the more recently issued Handbook, but with no additional habitats attached. Unfortunately, no precise record appears to have been preserved of the station in which it was found; and consequently, on Dr. Sinclair's decease, the species became entirely lost to New Zealand botanists. Since then, although the vicinity of Auckland has been repeatedly searched, no traces of its presence have been observed, nor, until a few months ago, had it been found elsewhere in the colony. As the plant must undoubtedly be considered one of the rarest members of the New Zealand flora, and as it is structurally one of the most interesting, it seems not out of place to put the fact of its rediscovery on record.

During a recent visit to the Thames, while examining the low clay hills which skirt the river immediately behind the native settlement of Kapu, I observed a solitary Pterostylis, that on inspection proved to be the long-lost P. squamaia. On a further search two more specimens were found, but both of these were barren. Unfortunately the time at my disposal was not sufficient to enable me to make a complete examination of the district, and to

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ascertain definitely if the plant is abundant or rare in this, the only locality in New Zealand of which we have now any certain knowledge of its occurrence. It seems most probable, though, that it will prove to be far from common although isolated specimens will doubtless be met with.

Pterostylis squamata can be distinguished from its congeners in New Zealand by the very obvious character of the lateral sepals being turned downwards, instead of being erect. This distinction has been made use of to divide the genus into two sections; the first, Antennœa, to which the majority of the Australian and New Zealand species belong, having the sepals erect; the second, Latochiuis, which includes our plant, having them deflexed Another excellent character is afforded by the labellum, which is filiform and hangs out of the mouth of the flower. It is also clothed throughout its whole length with copious golden-yellow hairs, except at the apex, where it bears a curious large purple gland, which has been aptly compared to the head of a nail.

Although Pterostylis squamata is rare and local in New Zealand, it is comparatively common in Tasmania, and is also found in Victoria and South-Western Australia. In this respect it agrees with Epacris purpurascens and Leucopogon richei, both of which are abundant Australian-plants, and both of which are confined to limited areas in New Zealand.

It is worth remarking that the affinity existing between the temperate Australian and New Zealand floras, easily recognised except in a few anomalous instances, is perhaps more clearly shown in the Orchideæ of the two countries, than in any other order of equal extent. Thus, in examining the distribution of the eighteen genera found in New Zealand, we find that no less than sixteen also occur in Australia, while the two remaining ones, although peculiar to New Zealand, are yet closely related to Australian forms. This result, striking in itself, becomes more so if we look closely into it, when it appears that out of the sixteen, six are absolutely peculiar to the two countries, and six others, although possessing outlying species in the Indian Archipelago or Pacific Islands, yet plainly have their head-quarters in Australia and New Zealand. Pterostylis is a good instance of this. It has twenty-four species in Australia, and seven in New Zealand. One of the Australian species extends into New Caledonia, but beyond this the genus is endemic in the two countries.

In the sixth volume of the Flora Australiensis, recently issued, I observe that Mr. Bentham considers the true P.squamata of R. Brown to be a form only of the P. rufa of the same author; and the plant to which the specific name of squamata has been, in many instances, applied by later botanists, is referable to a species long ago described by Dr. Lindley. If this view should meet with the general acceptation of botanists, as seems likely, our plant will in future bear the name of Pterostylis barbata, Lindley.

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Art. LII—A Description of some new Species of Gymnostomum.

[Read before the Wellington Philosophical Society, 29th August, 1874.]

1. Gymnostomum patulum, n. s.

Caulis brevis. Folia sicca crispata anguste lanceolato-linearia Basi vix latiora margine subaicem erecta, nervo virido in acumen producto e cellulis minutis rotundatis ohlorophyllo repletis basi parallelogrammis laxioribus areolata; perichætiala longiora erecta linealia. Theca inpedunculo brevi ellptica, orificio membrana clauso, operculo oblique subuli-rostro.

Stems short. Leaves crisped lanceolato-linear scarcely wider at the base, the margin erect under the apex, nerve green excurrent, point aristate, areolæ formed of minute rounded cells filled with chlorophyl, towards the base more lax and four-sided, the perichœtial leaves longer, erect. Fruit-stalk no longer than the perichætial leaves. Capsule elliptic, the mouth closed by a membrane, operculum oblique with an awl-shaped beak.

2. Gymnostomum knightii, n. s. Schimp. in litt.

Caulis perbrevis. Folia linguœformia carinata, margine e basi ad apicem recurva, ubique e cellulis minutis pellucidis rotundatis areolata, nervo crasso infra apicem evanido. Theca in pedunculo medio erecta oblonga sicca cylin-drica, operculo rostrato, et thecæ orificium membranâ non clausum.

Stem very short. Leaves tongue-shaped, keeled, margins recurved from base to apex. Areolæ minute, round, pellucid, nerve stout, vanishing below the apex. Capsule oblong, erect, mouth not closed by a membrane. Operculum beaked.

3. Gymnostomum calcareum var. intermedium.

Plantæ densissime cœspitosœ gracillimœ, inferne ferrugineæ. Folia minuta erecta crispula lanceolato-linearis mutica obtusa concta, abrupte evanidinervia, margine minute crenulata et circum apice erecta, e cellulis basi rectangular-ibus pellucidis superne rotundatis parvis papillosis areolata. Theca ovali-cylindrica; ore e cellulis transverse angustatis pluriseriatis formato.

Stems densely cæspitose, slender, ferruginous below. Leaves minute, erect, crisped when dry, linear-lanceolate, concave, obtuse, with a mucro. Margin minutely crenulate, erect round the blunt apex; nerve ceasing below the apex. Capsule cylindrical when dry, the mouth formed by several series of narrow cells.

This moss differs greatly from Seligera calcarea, Br. et Sch., to which Hooker refers it. The latter has a yellow-red coloured peristome with broad flat teeth, stout bars, without a median line.

Explanations of Plate XXVIII.

1.

Gymnostomum patulum.

Plant nat. size, and capsule enlarged.

  • a, b. Leaves enlarged.

  • c, d, e. Portion of leaves greatly enlarged.

  • 2. Gymnostomum knightii.

Capsule enlarged.

  • a, b. Leaves enlarged.

  • c, d. Portion of leaves greatly enlarged.

  • 3. Gymnostom calcareum var. intermedium.

Capsule enlarged.

  • a, b. Leaves enlarged.

  • c. Portion of leaf greatly enlarged.

  • 4.Gymnostomum sulcatum.

Plant enlarged.

  • a. Operculum and calyptra enlarged.

  • b, c, d. Leaves enlarged.

  • e. Portion of leaf greatly enlarged.

  • 5.Gymnostomum areolatum.

Plant enlarged.

  • a. Leaf enlarged.

  • b. Portion of leaf greatly enlarged.

  • 6.Gymnostomum angustatum.

Capsule enlarged.

  • a, b. Leaves enlarged.

  • c. Portion of leaf greatly enlarged.

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4. Gymnostomum sulcatum, n. s.

Dense cæspitosum erectum gracillimum innovando-ramosum, foliva dense conferta minutissime papillosa carinata crispula madefacta patentiaerecta lanceolata apiculata evanidinervia, margine plana v. parum recurva, e cellulis parvis incrassatis areolata. Theca ovalis in pedunculo brevi erecta sulcato-striata, operculo conico suboblique-rostrato.

Crowded, erect, very slender, branched. Leaves crowded, minutely papillose, keeled, crisped (when moistened patentia-erect), lanceolate, apiculate, nerve vanishing, margin flat or slightly recurved. Areolæ small. Capsule oval, erect, striate, on a short fruit-stalk. Operculum conical with an oblique awl-shaped beak.

5. Gymnostomum areolatum, n. s.

Gregarium brevissimum, folia concava laxiuscula crispula madefacta erecto-patentia elongato-oblonga v. subspatulata apiculata v. acuminata, ubique e cellulis amplis pellucidis tenuissime papillosis superne rotundato-angulatis v. quadratis areolata, nervo ad apicem fere extimum perdurante. Theca ovata in pedunculo brevi erecta, operculo conico suboblique rostrato.

Plants crowded, stem very short. Leaves concave, lax, crisp, when moistened erecto-patent, elongato-oblong or subspatulate, apiculate or acuminate; areolæ throughout large, pellucid, papillose, towards the point rotundato-angular, nerve extending to the apex. Theca ovate, erect, on a short fruit-stalk, operculum conical with an oblique beak.

6. Gymnostomum angustatum, n. s.

Plantæ humiles. Folia madefacta patentia curva papillosa linealia obtusa, margine erecta, nervo dorso prominente infra apice evanido. Theca erecta ovalis; operculum e basi conica in rostrum subulatum continuatum.

Plants smaill. Leaves (when moistened, spreading, curved), papillose, linear, obtuse, margin erect, nerve prominent at the back, vanishing below the apex. Theca erect, oval, operculum with an awl-shaped beak continued from the conical base.

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Art. LIII.—Description of some New Zealand Lichens.

Plate XXIII.

[Read before the Wellington Philosophical Society, 21st November, 1874.]

1. Phlyctis egentior, Nyl.

Thallus tenuissimus, livido-cineraceus, sat determinatus. Apothecia innata parva flavo-fusca rotundata aut difformia, margine thallino pulverulento cincta, disco plano brunneo sæpius pulverulento. Sporæ incolores falcatæ 7—9-septatæ, longit. 0–062 mm., crassit. 0–007 mm.

Ad cortices arborum.

Thallus very thin, livido-cineraceous, somewhat determinate. Apothecia innate, small, yellowish-brown, rounded or misshaped, bound by a powdery thalline margin. Spores colourless, falcate, 7—9-septate, length 0.062 mm., breadth 0–007 mm.

Differs from Phlyctis andennis and P. brasilensis in the colour of the thallus, and probably, by the powdery thalline margin with which the apothecia are bound.

2. Philyctis neo-zelandia, Nyl.

Thallus crustaceus, uniformis, albus, tenuis vel crassus, late expansus, rimosus, determinatus. Apotheca minuta, innata, maculiforrai-difformia, concaviuscula, carnea, excipulo proprio nullo—nisi marginie thallino spurio extus pulverulento—cincta. Sporœ incolores, vermiformi-cylandraceæ, spiraliter curvatæ, 7—11-septat, long., 0.005 millim.Thick.

Ad cortcicem arborum.

Thallus crustaceous, uniform, white, thin or thick, widely expanded, rimose, determinate. Apothecia minute, innate, concave, flesh-coloured, without a proper excipulum, surrounded by a pulverulent thalline margin. Spores colourless, spirally curved, 7—11-septate, 0.07 millim. long., 0.005 millim. thick.

On bark.

3. Lecidea stellulata, Taylor.

Thallus albidus, tenuissimus, areolato-diffractus, hypothallo nigro manifesto inter areolas. Apothecia sessilia, nigra. Sporæ fuscæ vel nigræ, uni-septatæ, longit. 0.01 mm., crassit. 0.006 mm.

Ad saxa.

Thallus white, very thin, areolate-diffractus, hypothallus black, visible between the areolæ. Apothecia sessile, black. Spores brown or black, 1-septate, 0.01 millim. long., 0.006 millim. thick.

On rocks.

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3*. Lecidea parasema, Ach.

L. parasema, Ach. (Nyl. in Lich. Scand. p. 217), described in Handbook N.Z. Fl., p. 584, I have not succeeded in collecting in New Zealand, and it is most likely that the common lichen, L. myriocarpa, DC., has been mistaken for it. This latter plant is common in New Zealand, although not described in the Handbook. The thallus is ashy-green, thin, subgranular, and determinate. The apothecia small and margined. Spores uniseptate, brown, 0.013 millim. long, and 0.008 millim. broad. Hypothecium black or obscurely brown.

Syn. L. punctata, Schœr. Exs. 197, and L. punctata g. denudata, Schœr. Exs. 529.

It is true that our lichen agrees with those in the collections of Leighton from Great Britain, and others from Germany and Scandinavia, under the name of L. parasema, but in every specimen that I examined of those collections the character agrees with L. myriocarpa, to which species in fact they belong.

4. Lecidea radomma, Nyl. in litt.

Thallus cinereus, granulosus. Apothecia atra, superficialia, mediocria, plana, margine promiente. Sporœ octonœ, incolores, ellipsoideæ, biloculares, plerumque tubulo loculos jungente, longit, 0.025 mm., crassit. 0.015 mm. Stratum hypothecii subhymeniale fuscum, stratum excipulare album.

Ad cortices arborum.

Thallus ashy, granular. Apothecia black, superficial, medium size, flat, with prominent margin. Spores eight in each ascus, colourless, ellipsoid, bilocular, cells united by a tube, 0.025 millim. long, 0.015 millim. thick. Subhymeneal stratum brown, excipular stratum white.

On bark of trees.

Differs from L. leucoplaca,Chev., in the excipulum being white, and the spores mostly with a connecting tube between the cells.

5. Lecidea marginiflexa.

Montague considers this plant to be L. tuberculosa, Fée (Flora N.Z., p. 299). This is a mistake. Fée's lichen has multilocular spores, while L. marginifexa has uniseptate spores and only one spore in each ascus.

Lecanora parella var. implicata.

Differs from the typical form in the thallus being neither granular nor warty, but rugose. In this it resembles a corticular specimen collected by J. P. Norlin at Tavaastia Asikkala. The disk is more coloured than usual in Buchanan's specimen, and is convex or flat in the full grown apothecia, as is sometimes the case in L. tartarea. Specimens are seen in all stages between slightly rugose and warty. One, which Nylander has named L. thelotremoides, is clearly intermediate between L. implicata and L. parella.

Syn. Lecanora implicata, Stirton.

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6. Lecidea fusco-lutea, Dickson.

Thallus albus, granulosus. Apothecia magna, sordide lutea vel liventia, plana, prominente marginata. Ascis unam sporam oblongam grandissimam minutissime muricatam foventibus.

Thallus white, granular. Apothecia large, dull yellow or liver-coloured, disk flat with a prominent margin. Asci with one large muricated spore, 0.188 millim. long, 0.052 millim. thick.

The Lecidea fusco-lutea of Acharius has bilocular spores, the cells connected by a tube. See Schœrer Exs. 215 (L. ferruginea b. leucorrhœa)

7. Lecidea leucoplaca, Chev.

Thallus cinereo-virescens, effusus, granulatus vel granulato-rugosus. Apothecia exserta, disco plano et margine prominente, stratum hypothecii subhymeniale et stratum excipulare nigra, lineo ochro-fusco disjuncta. Sporœ 8næ, incolores, ellipsoideæ, biloculares,longit. 0.025 mm. crassit. 0.012 mm.

Syn. L. premnea, Fries. L. grossa, Nyl. (in litt.) L. maculosa, Stirton L. melastegia, Nyl. L. melaclina, Nyl. Leighton Exs. 125 ! 90 !

Ad cortices arborum.

Thallus ashy-green, effuse, granular or granulo-rugose. Apothecía exserted, with flat disk and prominent margin, the subhymeneal stratum of hypothecium and excipular stratum black and separated by a yellowish-brown line. Spores 8 in each ascus, colourless, ellipsoid, bilocular, 0.025 millim long, 0.012 millim. thick.

I have examined specimens in Mr. Buchanan's collection—whence Dr. Stirton's lichens were obtained—and find no difference between L. maculosa, Stirt., and L. leucoplaca, Chev. The measurements of the spores given by Dr. Stirton would answer for L. parasema, Nyl. Lich. Scand.

8. Lecidea versicolor, Fée, var. subtuberculosa, Knight.

Thallus subtuberculosus, pallido-stramineus. Apothecia sparsa, fusca, marginata, margine concolore. Sporœ pallido-lutaE, uniseptatæ, ovatæ, longit. 0.024 mm. Hypothecium pallido-flavidum.

Ad cortices.

Thallus subtuberculose, pale straw. Apothecia scattered, brown, margined; margin concolorous. Spores pale yellow, 1-septate, ovate, 0.038 millim. long, 0.024 millim, thick. Hypothecium pale yellow.

On bark of trees.

It is not unlikely that L. taitensis, Mont., and L. versicolor, Fé, are the same plant. If so the varieties in New Zealand are (1) L. taitensis with rugose thallus, and spores 0.075 mm. in length, and margin of apothecium much darker than the ashy-brown disk; (2) L. versicolor with pale border and brown disk, spores .05 mm. in length, hypothecium white, and thallus tuberouloso

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isidiose; and (3) L. taitensis var. subtuberculosa, with smoother thallus, the disk and margin of the apothecium brownish-black, spores 0.033 mm. in length, and hypothecium pale yellow.

In L. marginiflexa the asci contain only one spore in each and that linear oblong and bilocular. In L. fusco-lutea, Dicks., also, the asci contain only one spore in each, but that is linear oblong or cylindrical, and minutely muricated.

8c. Lecidea versicolor, Fée.

Thallus albus, granulato-inæqualis vel granuloso-isidiosus. Apothecia disco fusco, et margine pallido, tumido, undulato. Sporœ uniseptatæ, ovatæ, incolores, longit. 0.05 mm., crassit. 0.023 mm. Hypothecium album.

Ad corticem arborum.

Thallus white, granulose, unequal, or granular-isidiose. Apothecia with brown disk, and pale tumid wavy margin. Spores 1-septate, ovate, colourless, 0.05 millim. long, 0.023 millim. thick. Hypothecium white.

9. Lecideas taitensis, Mont.

Thallus cineraceus vel eborinus, graulatus vel verrucoso-rugosus, passim rimulosus. Apothecia disco fumeo, et margine fusco-atro, tumido, undulato. Sporœ pallido-lutœ vel incolores, magnæ, ovatæ, incurviusculæ, uniseptatæ, longit. 0.075 mm., crassit. 0.035 mm. Stratum hypothecii subhymeniale incolore, perithecium fuscum.

Ad corticem arborum.

Thallus greyish or cream-coloured, granulate or warty rugose, here and there rimulose. Apothecia greyish-brown, and margin brownish-black, tumid, wavy. Spores pale yellow or colourless, large, ovate, little incurved, 1-septate, 0.075 millim. long, 0.035 millim. thick. Hymeneal stratum of hypothecium colourless; perithecium brown.

On bark.

10. Lecanora contigua, Ach., var. meiospora, Nyl.

Thallus albus vel cineraceus, areolatus, lineâ fuscâ limitatus. Apothecia sessilia, convexa, fusco-nigra, immarginata. Sporæ, incolores, ellipsoideæ, longit. 0.014 mm., crassit. 0.008 mm. Hypothecium fusco-nigrum.

Ad saxa et supra terram argillaceam.

Thallus white or ashy-white, areolate or cracked, bounded by a dark line. Apothecia sessile, convex, brownish-black, immarginate. Spores colourless, ellipsoid, 0.014 millim. long, 0.008 millim. thick. Hypothecium brownish-black.

On rocks and clayey banks.

11. Lecidea melanotropa, Nyl.

Thallus fractus, granuloso-inæqualis vel granuloso-rimosus, in aliquot

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exemplis sublævis, albidus vel pallido-cinereus vel cinereo-flavus. Apothecia nigra vel cæsio-pruinosa vel cæsio-pallentia, plana, demum convexa; margine obtuso, ceraceo-pallido, quandoque subconcolori. SporoE incolores, ellipsoideæ, interdum incurvæ, sæpius utroque apice acutiusculæ, longit. 0.014 mm., crassit. 0.007 mm. Hypothecium incolor.

Ad cortices arborum.

Thallus cracked, unequally granular, or granuloso-rimose, sometimes rather smooth, white, or pale-ashy or ashy-yellow. Apothecia black or bluish pruinose, at length convex; margin broad, waxlike, pale, now and then of one colour with the disk. Spores colourless ellipsoid, sometimes incurved, often rather acute at either end, 0.014 millim. long, 0.007 millim. thick. Hypothecium colourless.

The character of the thallus and the colours of the apothecia vary greatly, but nevertheless nearly all the variations of the latter may be found in one and the same plant. The thallus varies according to the nature of the bark it grows on. A variety named by Dr. Nylander (in a letter) Lecidea cœsio-pallens, Nyl., is best treated as a distinct variety.

12. Lecidea planella, Nyl. Syn. Lich. Novæ-Caledoniæ, p. 45.

Thallus cinereo-virescens, effusus, tenuissimus vel evanescens, et sæpius vix ullus. Apothecia adpressa vel peltata, albida vel carneo-pallida vel carneo-lutea vel carneo-cinnabarina, sæpius undulato-difformia; margine paulo dilu-tiore, evanescente, (latit. 0.1 mm. ad 0.25 mm.). Sporœ octonæ, uniseriales, incolores, fusiformes, uniseptatæ, longit. 0.008 mm., crassit. 0.002 mm. Para-physes mediocres, apice clavato. Hypothecium incolor.

Ad cortices fruticium et filicium et supra muscos.

Syn. Bœomyces pertenuis, Stirton. Biatora cinnabarina, Bab., Flora N.Z., p. 300.

Thallus ashy green, effuse, very thin, frequently scarcely any. Apothecia appressed or peltate, white or pale fleshy-coloured or fleshy-yellow or fleshy-cinnabar, often undulate and irregular in outline, margin somewhat paler, evanescent, (diameter 0.1. ad 0.25 mm.). Spores 8 in each ascus, in one row, colourless, fusiform, 1-septate, 0.008 millim. long, 0.002 millim. thick. Paraphyses club-shaped, apex nearly as broad as ascus.

On bark of shrubs and ferns, and on mosses.

I have no doubt that this is the Biatora cinnabarina of Babington (Flora N.Z., p. 300), and certainly the Bœomyces pertenuis of Stirton. After examining specimens from Mr. Buchanan's collection, and numerous specimens in my own, I have met with no instance of spores with four cells in one case; there is an appearance of four divisions, but no more than one true septa. The lichen agrees in every particular with Nylander's description. The spores in plate 129 C., Flora N.Z., should be uniseptate.

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13. Lecidea plesia, Knight.

Thallus ochroleucus, tenuis, areolo-rimosus. Apothecia nigra, plana, mediocria, elevato-marginata, margine flexuosa vel subanguloso-difformia. Sporœ lineari-clavatæ, 4—5-septatæ, incolores vel diluto-fuscentes, longit. 0.032 mm., crassit. 0.011 mm. Hypothecium nigrum.

Ad saxa.

Thallus pale ochrey colour, thin, divided into areolæ by chinks. Apothecia black, plane, medium size; margin raised, flexuose or subangularly deformed. Spores linear, clavate, 4—5-septate, colourless or pale brown, 0.032 millim. long, 0.011 millim. thick. Hypothecium black.

Differs from L. contigua in colour of thallus and the multilocular, lineari-clavate spores; from L. sabuletorium, whose spores are elongate fusiform.

14. Lecidea allotropa, Nyl.

Thallus continuus, tenuissimus, cineraceus. Apothecia sessilia, planiuscula, demum convexa, immarginata, nigra, sat parva. Sporœ oblongæ, 3-septatæ, longit. 0.015 mm., crassit. 0.006 mm. Stratum hypothecii subhymeniale incolore, stratum inferiore album.

Thallus continuous, very thin, greyish. Apothecia sessile, flat, at length convex, immarginate, black, rather small, spores oblong, 3-septate, 0.015 millim. long, 0.006 mm. thick. Subhymeneal stratum of the hypothecium colourless, the infeirior white.

14.* Pannaria periptera, sp. n.

Thallus monophyllus, plumbeus, orbicularis, e laciniis linearibus ferme adpressis flabelliformibus multifidis radiantibus compositus, centro concreto. Apothecia rubro-fuscentia, plana, margine duplici cincta, (excipulo a thallo crenulo marginato). Sporœ incolores, ellipsoideæ, simplices, longit. 0.013 mm., crassit. 0.0075 mm.

Ad cortices arborum.

Thallus monophyllous, lead colour, round, formed of linear, firmly pressed, fan-shaped, radiating laciniæ, coalesced in centre. Apothecia reddish-brown, flat, bound by double margin—the proper excipulum surrounded by a crenulate thalline margin. Spores colourless, ellipsoid, simple, 0.013 millim. long, 0.0075 mm. thick.

On bark of trees.

Named by Dr. Nylander (in a letter) Coccocarpia periptera. One of the most distinctive characters of the Coccocarpiœ is the adnate, immarginate apothecia; it would, therefore, be unsatisfactory to place among the species of that genus a lichen with two distinct raised margins to the apothecia—a thallodal one enclosing a sharply defined excipulum. It is true that like C. smaragdina the thallus is furnished with gonimia in lieu of gonidia; but

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both Pannaria and Coccocarpia agree in that character. As to our lichen being monophyllous, this is a very uncertain and questionable character. Many specimens of it can no more be considered monophyllous than P. rubiginosa, which latter has never been treated as a Coccocarpia, although its close ally C. plumbea, Nyl., is by Nylander. I agree with Dr. Lauder Lindsay that Nylander's arrangement, which separates P. rubiginosa from P. plumbea, is a most artificial and unnatural one (vide Trans. Roy. Soc. Edinb., vol. xxii part 1. p. 256).

15. Lecanora flavo-pallescens, sp. n.

Thallus tenuis, subgranulosus, flavo-albicans, interdum evanescens. Apothecia biatorina, disco fulvo plano tandem tumido, marginem pallidum levem demum excludente. Sporœ simplices, incolores, longit. 0.015 ad 0.02 mm., crassit. 0.01 ad 0.013 mm.

L. flavo-pallescens, Nyl. in litt.

Ad ligna. Grandibus sporis satis dignoscitur.

Thallus thin, subgranular, yellowish-white, sometimes evanescent. Apothecia biatorine, buff or tawny, disk flat, at length tumid, overlapping the pale smooth thalline border. Spores simple, colourless, 0.015 to 0.02 millim. long, 0.013 millim. thick. Closely allied to L. varia, if not a variety. The specimens with tumid apothecia resemble L. symmetrica, but differ from it and from all varieties of L. varia in the smooth thalline border of the apothecia, and especially in the much larger size of the spores, which in none of the varieties of L. varia in Leighton, Schœrer, and several German collections that I have examined, exceed 0.011 mm. in length. L. varia is a most polymorphous species. The numerous varieties may be arranged in four sets.

(1).

Those with larger apothecia crowded together, with bold granular flexuose or angular margin, growing on wood, viz:—Parmelia varia, Ehrh. var. vulgaris, Korb., Leight., Exs. 51; Lecanora varia var. pallescens, Schœr., Exs. 325.

(2).

Those with minute apothecia, with pallid powdery margin and flat disk, sometimes a little convex, viz:—Lecanora varia var. hypopta, Ach.; L. aitema, Ach.; L. apocrœa, Ach.; L. sarcopis, Ach.; Parmelia varia, Leight., Exs. 176.

(3).

Those with tumid apothecia, viz:—Lecanora varia var. symmetrica, and other varieties.

(4).

Lecanora varia var. polytropa, with its varieties, growing on stones.

16. Lecanora thiomela, Nyl.

Thallus crustaceus, citrinus, areolatus, areolis hypothallum fuscum obtegentibus, linea fusca limitatus. Apothecia minuta, cupulæformia, marginata, nigra, margine thallino citrino cincta. Sporœ fuscentes, ellipsoideæ, uni-septatæ,

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longit. 0.017 mm., crassit. 0.01 mm.

Ad saxa.

Thallus crustaceous, citrine, areolate, areolæ covering the brown hypothallus, bordered by a dark line. Apothecia minute, black, bound by a citrine-coloured thalline margin. Spores brown, ellipsoid, 1-septate, 0.017 millim. long, 0.01 millim. thick.

17. Ascidium melanosporum, sp. n.

Thallus crustaceus, verrucosus, eborinus. Apothecium hemisphericum, apice depresso-umbilicatum. Perithecium nigrum, globosum, apice poro pertusum. Nucleus paraphysibus tenuibus farctus. Thecæ mono-sporæ. Spora magna, fusiformis, multicellulosa vel muricata, demum fusca, opaca, longit. 0.18 ad 0.25 mm., crassit. 0.065 ad 0.09 mm.

Ad corticem arborum.

Thallus crustaceous, verrucose, cream-coloured. Apothecium hemispherical, depressed at the apex. Perithecium black, globose, opened by a pore at the apex. Nucleus with slender paraphysis. Theca 1-spored. Spore large, fusiform, many-celled or muricated, at length brown, opaque, 0.18 to 0.25 millim. long, 0.065 to 0.09 millim. thick.

On bark of trees.

Differs from A. domingense and A. cinchonarum, in the spores being irregularly cellular, much broader, and at length quite opaque. (Vide Tab. 1, fig. 20, Nyl. Prod. Flores Novo-Granatensis, Lichenes.)

18. Melaspilea metabola Nyl. Lich. Novæ Caledoniæ, p. 69.

Thallus cinereus, diffracto-rimosus vel areolatus, tenuissimus. Apothecia nigra vel fusca, mediocria, rotundato-lecideoidea (juniora punctiformia), convexa, immarginata, intus pallido-incoloria. Sporœ incolores, cylindraceæ, inferne magis attenuatæ et curvatulæ (sæpe in capillam productæ), murali-divisæ, longit. 0.075 ad 0.1 mm., crassit. 0.01 mm.

Ad cortices arborum.

Thallus ashy, cracked or areolate, very thin. Apothecia black or brown, medium size, rotundato-lecideoid—younger mere spots—convex, immarginate, pale within. Spores colourless, cylindrical, curved, attenuated below, often terminating in a hair, muriculate, 0.075 to 0.1 millim. long, 0.01 millim. thick.

On bark.

19. Squamaria perrugosa, Nyl.

Thallus cinereo-fuscus vel cinereo-rufescens, adpressus, tesselato-areolatus, areolis in centro depressis, versus peripheriam laciniato-divisus, laciniis pterygioideo-expansis. Cephalodia cerina vel ochraceo-rufescentia, demum in radios fissa. Apothecia ochro-badia, concava; excipulum proprium marginem

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thallinum cinctum. Sporœ uniseriales, incolores, ellipsoideæ, simplices evl perrarò biloculares, longit. 0.013 mm., crassit. 0.008 mm.

Ad saxa.

Thallus ashy-brown or ashy-red, closely adhering to matrix, tesselato-areolate, areolæ depressed in centre, divided towards circumference into laciniæ, laciniæ expanded, wing-like. Cephalodia dull yellow or yellowish-red, at length split into rays. Apothecia yellowish-brown, concave. Proper excipulum surrounded by thalline border. Spores in a single row, colourless, ellipsoid, simple, or very rarely 2-celled, 0.013 millim. long, 0.008 millim. thick.

Syn. Squamaria thaumasta, Stirton.

Section of apothecium shows the proper excipulum of a pale colour, bound by a thalline border.

20. Squamaria perrugosa, Nyl., var. neglecta, Knight.

Thallus cinereo-albescens, granulato-verrucosus, crustaceo-adnatus, centro confusus, versus peripheriam lobulo-crenatus vel pterygioideo-expansus. Apothecia, rosea, farinosa, margine thallino crasso, tumido, farinoso, obsolete crenato cincta. Cephalodia convexa, irregulariter fissa. Asci prælongi, angusti. Sporœ uniseriales, octonæ, ellipsoideæ, simplices, pellucidæ, longit. 0.027 mm., crassit. 0.013 mm.

Ad saxa.

Similis L. perrugosœ, sed sporis subgrandibus facile ab eo distinguitur.

Thallus greyish-white, granulate with warts, closely attached to matrix, centre confused, towards circumference lobulato-crenate, or outspread wing-like. Apothecia rose-coloured, farinaceous, bordered by a thick, swollen, obsoletely crenate margin. Cephalodia convex, irregularly split. Asci very long and narrow. Spores in one row, ellipsoid, simple, pellucid, 0.027 millim. long, 0.013 mm. thick.

On rocks.

Differs from the typical form in the larger size of the spores, and the characters of the thallus.

Lecidea campylospora, Stirton, is the Lecanora taitensis, Mont., as determined for me some years since by Dr. Nylander.

Lecidea maculosa, Stirton, is the L. leucoplaca, Chev.; L. premnea, Fries.; L. grossa, Nyl. in litt.; L. melastegia, Nyl. Leighton's Exs. 90 ! 125 !

Melaspilea amphorodes, Stirton, is the M. metabola, Nyl. Syn. Lich. Nov. Cal. p. 69.

Lecidea (Lecanora?) implicata, Stirton, is the Lecanora thelotremoides, Nyl. in litt.

Astrothlium prostratum, Stirton, is the A. ochrocleistum, Nyl. in litt.

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Note.—In reference to Dr. Stirton's description of certain New Zealand lichens, published in the Transactions of last year, page 235, I find that the Bœomyces pertenuis, Stirton, is the Lecidea planella, Nyl., described in Dr. Nylander's Synopsis Lichenum Novæ Caledoniæ, p. 45.

Psoroma implexa, Stirton, is the P. sphinctrina var. pholidotoides, Nyl. Journ. Linn. Soc. p. 244. A careful examination of specimens of Dr. Stirton's lichen in the collection of Mr. Buchanan discloses no difference between them and Dr. Nylander's description. It grows on bark or trees, not on rocks as (inadvertently?) stated in Dr. Stirton's description. It is to be observed that the same specimen may exhibit here and there simply a broad black boundary line, and in other parts coarse black dendritic radiating fringe.

Psoroma arthrophyllum, Stirton. This is a coarse variety of P. subpruinosum, Nyl. Like many common lichens the P. subprinosum has many varieties. They all agree in one respect—their spores are spherical or oval, with no tendency to be pointed at the ends. In the same ascus are often found both descriptions of spores intermixed. In such instances it may be that the spherical appearance is due to the spore lying transversely in the ascus, when it would be seen “end on,"—but I am not sure that such is ever the case.

Squamaria thaumasta, Stirton. This is the Sq. perrugosa, Nyl., of which a full and excellent description is given by Dr. Nylander in the Linnean Journal above quoted. The scales of the thallus instead of being umbate are most frequently somewhat depressed in the centre. The spores are strictly uniserial and simple. Very rarely an uniseptate spore may be met with. This lichen is closely allied to Sq. gelida, Linn.

These and most other lichens cannot be satisfactorily studied without the aid of a microscope. The dimensions and figure of the spores, the distinction between gonidia and gonimia of Nylander, and other differences, can only be determined by a good microscope. Dr. Hooker in the Handbook of the N.Z. Flora, especially alludes to the excellent specific characters afforded by the spores, but it would, he adds, take many months of microscopic study to ascertain accurately the size of the spores of New Zealand lichens. He has in numerous instances given references to Dr. Nylander's works where the dimensions are given. I have taken the measurements of the spores of most of the New Zealand lichens, and find them to agree with those stated by Dr. Nylander in his work on the Lichens of Scandinavia.

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Description of Plate XXIII.

Fig.

1. Phlyctis egentior, Nyl. Section of thallus and apothecia. a, 7—9-septate spores.

2.

Phyctis neo-zealandia, Nyl. Section of thallus and apothecia showing asci and paraphyses. a, Ascus and spores. b, 7—11-septate spores. c, do. do. of different form.

3.

Lecidea stellulata, Taylor. Ascus and spores, a, 1-septate spores.

3* Lecidea myriocarpa, DC. Section of portion of thallus with apothecia. a, Section of hymenium with ascus and paraphyses. b, 1-septate spore.

4.

Lecidea radomma, Nyl. Section of thallus and apothecia. a, Ascus and spores. b, Ascus and spores, c, Bilocular spore.

5.

Lecidea marginiflexa, Taylor. Section of portion of thallus with apothecia. a, 1-septate spore. b, 1-septate spore.

6.

Lecidea fusco-lutea, Dickson. Section of portion of thallus with apothecia. a, Muricated spore, b, Muricated spore.

7.

Lecidea leucoplaca, Chev.

Section of portion of thallus with apothecia.

a, Ascus with spores. b, do. do. c, 1-septate spore. d, do. do.

8.

Lecidea versicolor, Fée, var. subtuberculosa, Knight. Section of portion of thallus with apothecia. a, 1-septate spore. b, do. do. 8c, Lecidea versicolor, Fée, 1-septate spore.

9.

Lecidea taitensis, Mont.

Section of portion of thallus with apothecia.

a, Ascus with spores, b, 1-septate spore. c, do. do.

10.

Lecanora contigua, Ach., var. meiospora Nyl. Section of apothecia. a, 1-septate spore. b, Spores, different forms.

11.

Lecidea melanotropa, Nyl. Section of thallus and apothecia. a, 1-septate spore. b, spores, different forms.

12.

Lecidea planella, Nyl. Section of portion of thallus with apothecia. a, Ascus with spores. b, Spores. c, Ascus with spores.

13.

Lecidea plesia, Knight. Section of thallus and apothecia. a, 4—5-septate spores. b, Ascus and spores. c, Spores.

14.

Lecidea allotropa, Nyl. Section of apothecia. a, Ascus with spore.

14* Pannaria periptera, sp. n. Section of portion of thallus with apothecia. a, Ascus with spores. b, do. do. and paraphysis. c, Spores.

15.

Lecanora flavo-pallescens, sp. n. Section of thallus with apothecia. a, Ascus with spores. b, Spore.

16.

Lecanora thiomela, Nyl. Section of portion of thallus with apothecia. a, Ascus with spores. b, do. do. c, Spore. d, do.

17.

Ascidium melanosporum, sp. n.

Section of apothecium.

a, Spore. b, do.

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18.

Melaspilea metabola, Nyl. Section of portion of thallus with apothecia. a, Ascus with spores. b, Spore.

19.

Squamaria, perrugosa, Nyl. (young state.) Section of portion of thallus with apothecia. * Section, of apothecia. a, Ascus with spores. b, do. do. c, do. do.

20.

Squamaria perrugosa, Nyl., var. neglecta, Knight. Ascus with spores.

Art. LIV.—Description of a new Lichen (Stereocaulon buchanani).

Plate XXV.

[Read before the Wellington Philosophical Society, 21s November, 1874.]

A Few days ago I received from Mr. J. Buchanan of the Colonial Museum, Wellington, a lichen which presents peculiarities of internal organization of sufficient importance to warrant a special notice. As these peculiarities are somewhat anomalous, the absence of spermogonia in the specimens is so far a matter of regret, inasmuch as its generic place is rendered to a certain extent doubtful.

Of late these secondary organs of fructification have been occupying considerable attention, and the modifications in size and shape of the spermatia, as well as the comformation and appearance of the spermogonia themselves, have served, in doubtful cases, as a means of discriminating genera, more especially Lecanora from Lecidea.

The lichen in question occupies an isolated position, and presents affinities to three genera, viz:—Gomphillus, Stereocaulon and Bœomyces.

In the extreme length and tenuity of its spores, the former dimension reaching almost the limit, in this respect, amongst lichens; in the tenacious stiff gelatine which prevades the hymenium, and almost conceals the paraphyses until the application of liq. potassæ, which softens if it does not dissolve this gelatine; as well as in the presence of longitudinal cavities apart from the thecæ, this lichen shows considerable affinities to the curious and anomalous genus Gomphillus. On the other hand the apothecia, in their external conformation and, more especially, in the fact that occasionally very short stipites are present, coated with granules, betray a manifest relationship to Stereocaulon, with which I have meanwhile united it. This relationship is brought out all the more through the variety of St. condensatum, Hffm., where the apothecia are found also sessile on mosses, and have at first sight much the aspect of those of a Lecidea.

During the current year I detected on Ben Lawers, Perthshire, a form of

[Footnote] * Dated at Glasgow, 21st September, 1874.

[Footnote] * Dated at Glasgow, 21st September, 1874.

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this variety of St. condensatum, where the apothecia do not betray the slightest traces of stipites, and which, in the absence of intermediate states, I am strongly inclined to elevate to the rank of a species, more especially as the spores are shorter and thicker than any belonging to this species hitherto seen.

In St. strictum, Bab., as well as in one or two others, are seen spores, very much resembling those of the present lichen, although not more than half their length. The reactions by means of iodine on the hymeneal gelatine also correspond generally to those seen in species of the same genus.

The relationship of this lichen to Bœomyces is less obvious, and, but for Dr. Nylander's assertion, (Synop. p. 175) and a certain primâ facie resemblance, I would not have mentioned this genus as analogous, indeed in the light of the present plant I can assert, with considerable confidence, that Gomphillus has much stronger affinities to Stereocaulon than to Bœomyces.

In plants such as the present it is extremely foolish to dogmatize in the matter of classification, and we are taught the lesson that nature is not to be cramped and confined by any such well defined limits as our modern classifiers would fain lead us to believe.

Stereocaulon buchanani, Strn.

Thallus tenuis effusus, cinerascens vel cinereo-virescens, e gonidiis conglomeratis mediocribus et filamentis irregularibus, fere omnino constans; apothecia majuscula (latit. 1—2 millim.), sessilia vel elevato-sessilia, interdum conglomerata, fusca (junioribus pallidis et nonnihil turbinatis), convexa et immarginata, intus fuscescentia, textura tenaci cornea. Sporœ 8næ, in thecis cylindricis, longissimæ, filiformes, 40–50-septatæ, longit. 0.16–0.22 millim., crassit. circa 0.003–0.004 millim.; paraphyses haud distinctæ. Gelatina hymenialis iodo leviter cærulescens (apicibus thecarum intensius tinctis) dein rufescens.

Thallus thin granular cinerascent or cinereo-virescent effuse, composed almost entirely of conglomerated gonidia of a medium size and an irregular fibrous stroma; apothecia rather above average sessile or elevato-sessile fuscous, in a younger state much paler and then somewhat turbinate, convex and immarginate, fuscescent within and of a tenacious horny texture; spores 8, in cylindrical asci, very long and filiform, 40–50-septate, and measuring 0.16–0.22 by 0.003–0.004 of a millim., paraphyses not distinct. Hymeneal gelatine slightly cærulescent with iodine, the apices of the asci more deeply tinted, the whole assuming afterwards a rufescent tint.

Notes on Stereocaulon buchanani, Stirton.

The valuable report of Dr. Stirton on Buchanan's plant has been brought

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Stereocaulon Buchanani, Stirton.

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under my notice white it was passing through the press. The following notes—with the analysis made some years since—will be found useful. The plant has been named by Dr. Nylander (in a letter)Lecidea subglobosa

Dr. Stirton's follow pretty closely parts of Nylander's desription of the genus Gomphillus, Nyl. There is, however, an essential difference in the structure of the thallus from that of the Gomphillus, which is overlooked by Dr. Stirton, but which is strongly in support of his view that the plant should be classified under the genus Stereocaulon. In Buchanan's lichen the thallus is made up of gonimia, as in Stereocaulon, instead of gonidia as in Gomphillus. Nylander's description of the genus Gomphillus would apply if “gonimia” was substituted for “gonidia,” thus: “Thallus tenuissimus,e gonimiis [gonidiis] mediocribus sphericis elementisque filamentoso-irregularibus gelatinose conglu-tenatis constans.” The paraphyses are indistinct only from their exquisite fineness and compactness; not, I think, from the “tenacious stiff gelatine which pervades the hymenium.”

The filaments which bind the gonimia together are few and scattered. The spores are 0.160 millim. long, and 0.003 millim. broad.

Plate XXV.—Stereocaulon buchanani. Fig.1,Section of apothecium with portion of thallus. 2, Gonimia. 3, Gonimia, with filaments. 4, Spores.

Art. LV.—On the Occurrence of Hæmatoccus sanguineus on the Wool of
a dead Sheep.

Plate XXIV.

[Read before the Philosophical Institute of Canterbury, 2nd April, 1874.]

That group amongst the Algæ, which was formerly included under the term Protococcus, contains several species, which by their occurrence together in enormous numbers suffuse with a characteristic colour the substance upon which they grow.

To this section of plants belongs the so-called “red snow,” which consists of simple roundish microscopic red cells, by which the ice and snow of alpine and polar regions are coloured red; another species sometimes grows in swamps and canals imparting to the water a similar hue; while to another, luxuriating in certain portions of the ocean, the Red Sea owes its name.

There are also species which form a red covering upon mosses or rocks; others again grow upon vegetable products used by man, for instance, fruit and bread—an occurrence which has given rise to remarkable errors of even historical importance as to their origin and nature.

Similar Algæ appear also occasionally upon animal bodies. Thus I found

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upon the wool of a dead sheep in the New Zealand Alps, in the neighbourhood of the West Coast road, a species which, as far as microscopic examination has hitherto shown, is Hœmatococcus sanguineus.

This plant, forming blood-red spots upon the wool, consists of globular cells, invisible to the naked eye. Viewed under the microscope, the globules exhibit a golden-yellow colour, whilst the fibres of the wool are scarlet-red. This colour without doubt is derived from the red pigment of the cells, which has been pressed out from some cause impregnating the fibres.

The portion of the wool which was not exposed directly to the light has a green appearance, and this colour is most probably of the same origin as the red.

An analogous example of the change of colours is furnished by the plant which forms the so-called “red snow” mentioned above. Whilst the contents of the cells in a certain stage of their development are purple red, they change to a green colour in another; and one of the Algæ which imparts to the surface of the water a blood-red appearance, exhibits this colour only when subjected to the direct rays of the sun; the higher the light, the more intense the colour appears. At other times the plant is green.

A specimen of wool (sent to the Canterbury Museum by Messrs. Studholme Brothers), showing a green-colour and taken from a living sheep, was shown to me by Dr. Haast, but this may be due to some infiltration of some mineral colour, although it has all the appearance of having been produced by a similar cause. It is true that all the fibres are uniformly coloured, but the Algæ originally growing amongst the wool may have been destroyed, leaving only the traces of their former existence in the fibres of the wool. It is desirable that further specimens presenting this uncommon appearance should be collected by persons having the opportunity of doing so.

Plate XXIV.—Fig. 1, Fibres of wool and Alga magnified. 2. Alga still more magnified.

Art. LVI.—On some of the Naturalized Plants of Otago.

[Read before the Otago Institute, 7th April, 1874.]

Mr. Kirk has recently called my attention to the fact that no list has yet been drawn up of the naturalized plants either of Dunedin or Otago. It is to be regretted that steps have not been taken, ere this, to watch and record the introduction of all foreign plants, as otherwise it becomes in course of time almost a matter of impossibility, on finding a plant of cosmopolitan distribution, to say, with any degree of certainty, whether it is indigenous or not. We have lately had experience of this difficulty, in the case of one of the commonest of weeds—Polygonum aviculare—which has given rise to so much discussion

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Haematococcus Sanguineus

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between Messrs. Travers and Kirk. For the future the evil may be to a great extent avoided, by cataloguing all the non-indigenous plants hitherto collected, and constantly supplementing this record by the addition of every fresh discovery. It is quite unnecessary to point to the uncertainty which exists with regard to a large proportion of the British flora as to its origin. In this country there are some plants to which doubt may yet attach, as to the time of their introduction and the place from which they were originally carried.

Perhaps there is no method by which foreign plants are introduced in such quantity and variety as by the direct importation of agricultural and other seeds from Britain and other countries where there is much cultivated land. Every one conversant with rural matters in England or on the Continent, knows that, even where high cultivation is the rule, hundreds of species of weeds contest for possession of the soil, along with the legitimate crop of the husbandman. The meadows and pastures teem with their own peculiar weeds, and the seeds of these are certain to be present in greater or less quantity in all grass seeds collected. English seedsmen are in the habit also of purchasing largely from foreign importers, particularly heavy seeds, as clovers, timothy, rape, etc., so that seed brought to this country from any part of Britain is nearly sure to contain a great deal that is not bargained for, even when most carefully selected. By this means also, the weeds of Europe, America, India, etc., are distributed far and wide over our Colonial Empire, and, as is too often the case, to the serious detriment of the country which imports them. As a case in point, I may mention that, in 1869, a farmer in Southland imported from Messrs. Lawson and Sons of London and Edinburgh—a firm which has made the cultivation of grasses their speciality—a quantity of selected grass seed for laying down in permanent pasture. The assortment comprised twenty-one species of plants, but besides all that were included in the list making their appearance in due time, no less than thirty-seven species of weeds were found among them, including the clover-dodder (Guscuta trifolii), and other comparatively rare plants.

In the Transactions of the N.Z. Institute, vol. II., p. 131, Mr. Kirk has drawn up a very elaborate series of divisions for the proper classification of naturalized plants. One of the most rapid agencies for the diffusion of such plants, and which affects horticultural as well as agricultural and accidentally introduced species, is that of floods. The Taieri plain furnishes a good example of this, plants from most diverse localities and introduced by totally different agencies, being there found in close proximity to each other.

In the accompanying list, I have to some extent followed the plan adopted by Mr. Kirk, using two or three similar contractions, and dividing them into similar series according to their method of introduction, viz:—

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  • (a.) Horticultural. (Hor.) Escapes from, or remains of gardens.

  • (b.) Agricultural. (Agri.) Escapes from, or remains of field cultivation.

  • (c.) Accidentally. (Acc.) Introduced by animals, or unintentionally by man's agency.

  • (d.) Uncertain. (Unc.)

List of Naturalized Plants found in Otago.

Ranunculaceæ.

1.

Ranunculus acris, (L.) Acc. Common in rich pastures and cultivated bush land.

2.

" repens, (L.) Acc.

Berberidaceæ.

3.

Berberis vulgaris, (L.) Hor. Abandoned gardens in the Taieri Plain.

Papaveraceæ.

  • 4. Papaver rhoeas, (L.) Acc. In one or two spots in cultivated land, and near Dunedin in gardens.

Fumariaceæ.

5.

Fumaria officinalis, (L.) Acc. In cultivated land.

Cruciferæ.

6.

Cheiranthus cheiri, (L.) Hor. Escaped from gardens, and gone wild in a few dry and stony localities.

7.

Nasturtium officinale, (R. Br.) Hor. or Acc. Possibly introduced as a salad plant, but now abundant in streams.

8.

Barbarea vulgaris, (R. Br.) Acc. Certainly the introduced plant, now a common wayside weed.

9.

Brassica oleracea, (L.) Agri. or Hor. On Otago Peninsula and Taieri Plain.

10.

" rapa, (L.) Hor. and Agri. Establishing themselves as weeds in some cultivations.

11.

" napus, (L.) Agri.

12.

Sinapis arvensis, (L.) Acc. Fields and roadsides.

13.

Cochlearia armoracia, (L.) Hor. Abandoned gardens.

14.

Lepidium campestre, (R. Br.) Acc. Roadsides.

Resedaceæ.

15.

Reseda suffruticulosa, (L.) Hor. As a weed in some gardens in Dunedin.

Violaceæ.

16.

Viola tricolor, (L.) Acc. In arable lands.

17.

” “ var. arvensis (Murr.). Acc. In arable lands.

Caryophyllaceæ.

18.

Silene gallica, (L.) Acc. or Hor. Abundant on the Forbury Head, Dunedin.

19.

Lychnis githago, (Lam.) Acc. In arable and grass lands.

20.

Stellria media, (Wither.) Acc. Common weed.

21.

Cerastium glomeratum, (Thiul.) Acc. In arable land and roadsides.

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22.

Cerastium vulgatum, (L.) Acc. Common weed.

Geraniaceæ.

23.

Geranium molle, (L.) Acc. Roadsides, fields, and meadows.

Linaceæ

24.

Linum usitatissimum, (L.) Agri. Fields and waste ground.

Leguminosæ.

25.

Ulex europœus, (L.) Agri. or Hor. Selfsown from gardens and hedgerows. Have become a pest in some pastures.

26.

Sarothamnus scoparisis, (Koch.) Agri. or Hor.

27.

Medicago sativa (L.) Agri. Occurs in pasture in Southland, but seems to make no progress.

28.

Medicago lupulina, (L.) Agri. Meadows.

29.

Trifolium pratense, (L.) Agri. "

30.

" medium,(L.) Agri." increasing fast.

31.

" repens (L.) Agri. In pastures, and in fact everywhere from sea-level to 1,500 feet.

32.

Trifolium minus, (Sm.) Agri. Very abundant in pasture, etc.

33.

Lotus corniculatus, (L.) Agri. In pastures.

34.

" major, (Scop.) Agri. Pasture in Southland.

36.

Anthyllis vulneraria, (L.) Acc. Introduced with grass seeds into Southland.

37.

Vicia tetrasperma, (Mœnch.) Acc. Fields and roadsides.

38.

" sativa, (L.) Agri. or Acc. Fields and pastures.

39.

Onobrychis sativa, (Lam.) Agri. Fields and meadows, but does not increase.

Rosaceæ

40.

Fragaria vesca, (L.) Hor. Woods, banks, and pastures.

41.

Rubus idœus, (L.) Hor. Old gardens.

42.

" fruticosus (?.) Hor. Roadsides, etc.

43.

Rosa rubiginosa, (L.) Hor.

44.

" canina, (L.) Hor.

Grossulariaceæ.

45.

Ribes grossularia, (L.) Hor. Old gardens and in bush.

Umbelliferæ.

46.

Apium graveolens, (L.) Hor. Escapes from gardens.

47.

Pitroselinum sativum, (L.) Hor.

48.

Pastinaca sativa, (L.) Hor.

49.

Daucus carota, (L.) Hor.

Caprifoliaceæ.

50.

Sambucus nigra, (L.) Hor. In a few spots in bush.

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Rubiaceæ.

51.

Sherardia arvensis, (L.) Acc. Pastures, arable land and roadsides.

Dipsacaceæ.

52.

Knautia arvensis, (Coult.) Acc. In a meadow at Roslyn, Dunedin.

Compositæ.

53.

Bellis perennis, (L.) Acc. Pastures.

54.

Achillœa millefolium, (L.) Acc. Pastures in Taíeri Plain.

55.

Anthemis nobilis, (L.) Hor. Meadows and roadsides.

56.

Matricaria parthenium, (L.) Hor. Meadows.

57.

Chrysanthemum leucanthemum, (L.) Acc. Pastures.

58.

" segetum, (L.) Acc. ".

59.

Artemisia absinthium, (L.) Hor. Spreading from gardens with great rapidity.

60.

Senecio vulgaris, (L.) Acc. Fields, waysides, etc.

61.

Centaurea cyanus, (L.) Acc. Fields in Taieri Plain.

62.

Carduus lanceolatus, (L.) Acc. Abundant everywhere.

63.

Chichorium intybus, (L.) Hor. Fields on the Taieri.

64.

Taraœacum dens-leonis, (Desf.) Acc. Fields and waysides.

65.

Sonchus oleraceus, (L.) Acc. Abundant weeds.

66.

" asper, (Hoffm.)

67.

"arvensis, (L.) Acc. Cultivated land.

68.

Crepis virens, (L.) Acc. Fields and roadsides.

Convolvulaceæ.

69.

Cuscuta trifolii, (Bab.) Acc. Introduced into Southland from Britain with clover seed in 1869.

Boraginaceæ.

70.

Echium vulgare, (L.) Acc. In meadow in Southland.

71.

Lithospermum arvense, (L.) Acc. Fields and roadsides in Taieri Plain.

Scrophulariaceæ.

72.

Digitalis purpurea, (L.) Hor. Escaped into the bush in many localities.

73.

Linaria purpurea, (Mill.) Hor. Near Dunedin.

74.

Rhinanthus crista-galli, (L.) Acc. In meadows.

75.

Veronica chamœdrys, (L.) Acc. Pastures in Taieri Plain.

76.

"serpyllifolia, (L.) Acc. Pastures and waste ground.

Labiatæ.

77.

Mentha viridis, (L.) Hor. Abundant on banks of streams.

78.

Ajuga reptans, (L.) Acc. Covering great parts of the town-belt round Dunedin.

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Primulaceæ.

79.

Primula vulgaris, (L.) Hor. In neighbourhood of gardens.

80.

Anagallis arvensis, (L.) Acc. Dry open banks and roadsides.

Plantaginaceæ.

81.

Plantago lanceolata, (L.) Acc. These ribworts are all found as meadow and wayside weeds, often associated together.

82.

" media, (L.) Acc.

83.

" major, (L.) Acc.

Polygonaceæ.

84.

Rumeœ crispus, (L.) Acc. An exceedingly abundant weed.

85.

" acetosella, (L.) Acc. Abundant everywhere, and most difficult of eradication.

86.

Polygonum aviculare, (L.) Acc. Indigenous or introduced. An exceedingly common weed.

87.

" convolvulus, (L.) Acc. A weed in many gardens in Dunedin.

Euphorbiaceæ.

88.

Euphorbia peplus, (L.) Acc. A garden weed; common.

89.

" lathyris, (L.) Acc. or Hor. Only seen in the neighbourhood of Dunedin, where it occurs as a garden weed.

Urticaceæ.

90.

Urtica urens, (L.) Acc. or Unc. Some small patches occur in the Valley of the Leith, and appear to be increasing, though not so fast as might be expected.

Gramineæ.

91.

Anthoœanthum odoratum, (L.) Agri. All introduced as pasture grasses.

92.

Phleum pratense, (L.) Agri.

93.

Alopecurus pratensis, (L.) Agri.

94.

Holcus lanatus, (L.) Acc. Very abundant in fields, swamps, roadsides, etc.

95.

" mollis, (L.) Acc.

96.

Poa annua, (L.) Acc. Indigenous or introduced.

97.

" pratensis, (L.) Agri. Meadows in Southland.

98.

Briza media, (L.) Acc. or Hor. Waysides about Dunedin.

99.

Dactylis glomerata, (L.) Agri. Meadows and pastures.

100.

Festuca ovina, (L.) Agri. Introduced as pasture grasses.

101.

" " var. duriuscula, (L.) Agri

102.

" Pratensis, (Huds.) Agri.

103.

" "var. elatior, (Koch.) Agri.

104.

Serrafalcus mollis, (Parl.) Acc. In meadows and roadsides.

105.

Lolium perenne, (L.) Agri. Introduced as pasture grasses.

106.

" italicum, (A. Braun.) Agri.

The foregoing comprise all the naturalized plants as yet collected by me. There are still, however, a great many introduced grasses, and I have never examined the littoral and aquatic floras, so that no doubt the above list would

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require to be doubled or trebled to represent adequately the number of nonindigenous plants. I hope that this small contribution may be the means of inciting other members of the Institute and Field Club to make known any plants which they have discovered, and which are not mentioned by me.

Art. LVII.—On Mottled Kauri.

[Read before the Auckland Institute, 26th August, 1874.]

Regarding the origin of the rich mottled shading of some kauri trees, considerable diversity of opinion exists, some ascribing it to a timber disease, while others believe such trees to belong to a variety of the kauri. The slightest examination, however, seems to show that this is no variety, but a peculiarity due to an abnormal growth in the sap-wood and bark. The mottled kauri is rather a rare tree, but a young mottled kauri is rarer still.

If it were a variety we should expect to find the markings in the roots and stumps; on the contrary, it is believed those parts are never affected, and for some distance up the trunk the timber is often just like that of other kauri. Allowing then, that it is to an abnormal state of the growth of the tree we owe this rich and valuable furniture wood, an enquiry as to the operating cause may not be uninteresting.

On examining a portion of the outer timber and bark of a highly mottled tree such as the specimens now exhibited were cut from, we see the bark entangled in the sap-wood, being pitted and streaked in some places very deeply. The specimens are all sawn from a board showing bark and sap-wood, and little or no heart timber. The bark has mostly fallen away during the seasoning, but the action and growth of the mottling are very clearly shown by examining the sections across grain. The bark is shown to become entangled in the ligneous growth, and has been surrounded and retained, sometimes in cells, sometimes leaving only a trace of non-fibrous barky matter in thin flakes, causing shaded parts, broad or narrow, according as the timber is cut relative to their planes. Nearest the bark the mottlings are nearly pure bark, and as they are found in the tree away from the surface, they are observed to be more and more of a ligneous structure. A close study of the specimens will convey an impression of the connection between the bark and the markings, better than any written description can do, but it is to be observed that all around this, in some degree foreign substance, the true wood is shaded and marked in itself, just as it is to be observed in knotty timber, and as the timber is transformed to the state of heartwood, a deposit of gummy and ligneous matter seems to transform even the largest of the imprisoned flakes into veritable woody fibre.

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Any abnormal state of growth seems not to affect the health of the tree, as mottled kauri trees are often the largest and soundest. The beauty of this timber and its effect in furniture are often marred by injudicious selection. Not only do trees vary in their mottled character, but by judicious sawing a great variety can be obtained from one tree, and nothing has a worse appearance in furniture—however well made—than an indiscriminate loading of mouldings, framing, pilaster, and panels, with rich and heavy mottled kauri.

Art. LVIII.—Description of a new Species of Isoëtes.—

Plate XXV.

[Read before the Wellington Philosophical Society, 29th August, 1874.]

The genus Isoëtes has been added to the New Zealand flora since the publication of the Handbook, the first species having been discovered in the Waikato Lakes by Captain F. W. Hutton and myself in 1869. The plant which forms the subject of the present notice was discovered by Captain Hutton and Mr. W. T. L. Travers three years later in Lake Guyon, Nelson, at an altitude of 3,000 feet. I am indebted to Mr. Travers for specimens.

Not having the opportunity of referring to the recently described austral species of this genus, I can only venture to offer a brief diagnosis under the provisional name of I. alpinus, and indicate its chief points of difference from the lowland species, I. kirkii, Braun.

Isoëtes alpinus, n.s.

Fronds numerous, ten to fifty or more, stout, rigid 4” — 6” long; phyllopodes extremely dilated, with the border produced for some distance along the edge of the frond; sporangium broadly elliptic, 4-septate; ligule minute, triangular; macrospores globose, perfectly smooth, except the faint tricrural line; microspores ovate, closely tuberculated, faintly angled, longitudinal furrow obscure.

Hab.—Lake Guyon, Nelson, alt. 3,000 feet—Messrs. Hutton and Travers; Lake Pearson, Canterbury, if I am correct in identifying a very imperfect specimen collected by Dr. Berggren with the present plant.

Solitary specimens of Isoëtes alpinus often exhibit from sixty to seventy crowded fronds, radiating in a circle of from six to seven inches diameter. The central vascular portion of the frond is also largely developed.

Our plant is readily distinguished from I. kirkii by its robust habit and larger size: and especially by the smooth macrospores and tuberculated microspores.

Plate XXV.Isoëtes alpinus, nat. size.—Fig. 5, Phyllopodium containing macrospores. 6, Sporangium. 7, Transverse section of ditto. 8, Macrospores. 9, Microspores.

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Art. LIX.—On the Occurrence of Juncus lamprocarpus, Ehr., in New Zealand.

[Read before the Wellington Philosophical Society, 29th August, 1874.]

In the early part of the present year I discovered Juncus lamprocarpus, Ehr., growing in great abundance about Karori and other places in the vicinity of Wellington; and a few months later in the southern part of the province of Otago. About Wellington it occurs only in wet, swampy places, where it often forms a large portion of the herbage and is closely cropped by cattle; at Invercargill it is found not only in swamps, but in places where the soil is scarcely moist, even on the railroad track, a peculiarity which is possibly due to the greater amount of moisture in the atmosphere at the latter place.

Juncus lamprocarpus is allied to J. holoschœnus, Br., which it resembles in the jointed and compressed stems and leaves, but from which it differs in the much branched panicle, in the segments of the perianth being shorter than the capsule, and especially in the fascicles never being collected into lateral cymes.

When compared with British specimens, the New Zealand plant is seen to have the internal divisions of the leaf and stem more prominent, while both the inner and outer segments of the perianth are acute and much shorter in relation to the capsule.

Our plant may be expected to prove of not unfrequent occurrence in the South Island, although probably confined to the province of Wellington in the North.

The avidity with which it is eaten by cattle is doubtless one cause of its having been so long overlooked in this district.