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Volume 8, 1875
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Art. XXXIII.—Remarks on the Pselaphidæ (Coleoptera) of New Zealand.

[Read before the Auckland Institute, 16th August, 1875.]

On referring to Captain Hutton's Catalogue of the New Zealand Insecta, as published in the “Transactions of the New Zealand Institute,” under date the 11th November, 1873, it will be observed that no mention is made of this group of the Pseudotrimera, most probably because of its having been unknown to our entomologists at the time.

As the omission of an entire group of Beetles from our only available list, seems to me a matter of importance, I will endeavour to remedy the

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defect, by giving a brief account of the species at present known to science.

The family Pselaphidœ is represented in these islands by fifteen named species, placed in six genera, four of which appear to be peculiar to Australia and New Zealand, whilst the others (Pselaphus and Euplectus) are of wide distribution; and there can be no doubt, that this list will be greatly augmented from time to time as our entomologists make further discoveries which will render, it necessary, to establish several new genera.

Those which have been described are named as follows:—

  • Tyrus mutandus

  • Pselaphus pauper

  • Bryaxis inflata

  • " micans

  • " dispar

  • " deformis

  • " impar

  • " grata

  • Dalma pubescens

  • Sagola major

  • " prisca

  • " miscella

  • " parva

  • Euplectus convexus

  • " opacus.

The Beetles comprising this family are small, but remarkably handsome as compared with the greater portion of our Coleoptera; their characteristic colour is red, varying from orange to brown; and they may be readily distinguished from all our other beetles by their abbreviated elytra and cumbersome antennæ, the latter being usually terminated by a distinct club; but inexperienced collectors are apt to confound them with the Brachelytra, a mistake, however, which is easily avoided by attending to the joints of the tarsi, as well as the general outline of the body; the abdomen of the Pselaphidœ, though comparatively as much exposed as is the case with the insects belonging to Brachelytra is much less elongate.

The group is divided into the sub-families Pselaphini and Euplectini, and as an instance of the importance of a thorough investigation of our Coleoptera, I may quote from a paper read by Dr. Sharp, before the Entomological Society of London, in which it is asserted that the discovery of “the new genera Dalma, which is intermediate between the two genera Batrisus and Euplectus, indicates that the division of the Pselaphidœ into two main groups can scarcely be maintained.”

I have found these insects in various parts of this Province amongst vegetable refuse, under boards and stones, in splintered stumps of trees, and in company with, if not actually joint inhabitants of the nests of, the Red Ant, at all seasons of the year; but although they occur under such varying circumstances, a collector will be a long time in obtaining many species, or even many individuals of any one species, as it is but seldom that more than one or two are met with at one time, and being inconspicuous as to size, they are very likely to be overlooked altogether.

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I now deposit in the Museum a typical specimen of this group, for reference by such entomologists as may not be acquainted with them, but as even a complete typical collection of the group in this Museum would not promulgate a knowledge of these interesting insects beyond that narrow sphere, I take the liberty of attaching copies of the descriptions of the fifteen species already enumerated, and of asking the members of this Institute to aid me in pressing on the “Board of Governors of the New Zealand Institute,” the advisability of reprinting them in the “Transactions.” If some such course is not pursued, what encouragement will be held out to our entomologists to persevere in their researches beyond the mere selfish gratification of enriching their private collections with a number of nameless beetles? And moreover, how are we to avail ourselves of the joint labours of our collectors, and those eminent entomologists who place their valuable services at our disposal in describing and naming our recently discovered insects, unless the course I venture to recommend be adopted? Surely it cannot be expected of our entomologists, that after expending a considerable amount of time and money in bringing to light our indigenous fauna, and inducing the more skilled European entomologists to name and describe those unknown to science, that they should also, if desirous of communicating such acquired knowledge, supplement their labours by personally transcribing the printed descriptions for each of our colonial collectors.

I will now conclude this paper by expressing a hope, that this year's volume of “Transactions” will afford satisfactory replies to these queries in the shape of a re-print of such descriptions, in the form of an appendix or otherwise, as the Board of Governors of the Institute may deem most beneficial to the interests of science.

Additional Notes, Descriptions, Etc.

Descriptions of two new genera of Pselaphidæ.

Dalma, nov. gen. Corpus sat elongatum, subdepressum. Palpi maxillares breves, articulo 2° basi gracile, apice abrupte fortiter incrassato, articulo 3° parvo subtriangulare, articulo ultimo crasso, securiforme-ovali, longitudine articuli 2;. Caput mediocre, nullo modo rostrato-deflexum, tuberculis frontalibus evidentis, sat distantibus. Antennæ breviusculæ, apice fortiter clavatæ, 11-articulatæ, basi distantes. Prothorax cordatus. Prosternum magnum; coxæ anteriores robustæ modice exsertæ. Trochanteres intermedii breves, ut femoris apex cum coxa articula est. Coxæ posteriores

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prominentes basi fere contiguæ. Abdomen sat elongatum, minus deflexum, marginatum, segmentis ventralibus sex, quorum primo vix conspicuo. Pedes robusti modice elongati, tarsis unguiculo unico valido.

Sagola, nov. gen. Labrum, broad and transverse; its front margin forming a gentle curve, the sides being more advanced than the middle. Mandibles without teeth on their inner edge, with the basal portion very thick; the apical portion abruptly curved inwards; elongate, slender, and acuminate. Maxillæ with the lobes distinct, short, but with long pubescence; their palpi short, four-jointed; first joint abruptly curved in the middle, second joint rather longer than the first, rather narrower at the base than at the extremity, twice as long as broad; third joint short, about as long as broad; fourth joint oval, broader than the preceding joints, about twice as long as broad; its extremity a little truncate, and furnished with a very minute appendage. Mentium large, rather broader than long, quadrate, but with the anterior margin forming a slight double curve, being a little produced and acuminate in the middle. Labial palpi short, stout, two-jointed; second joint shorter than and not quite so thick as the first joint. Paraglossæ prominent, extending about as far as the extremity of the labial palpi.

Antennæ, eleven-jointed, elongate, and rather stout, not clubbed; the apical joints being but little thicker than the basal ones, separated at their point of insertion by their broad, flattened, contiguous, frontal tubercles. Head short, not in the least rostrate. Eyes moderately large. Prosternum rather large, front coxæ slender, moderately prominent.

Mesosternum elongate. Middle coxæ large, only partly embedded in their cavities; separated only by a thin lamina of the mesosternum. Femoral portion of hind coxæ prominent and conical, contiguous at their base; their trochanters moderately large; but the apex of the femur almost attains the coxa. Legs elongate, simple; tarsi much shorter than tibiæ, with two well-developed unguiculi. Hind body elongate, strongly margined at sides; the dorsal and ventral plates equal to one another, with five visible segments, both above and below, but with a well-developed additional basal segment visible on dissection, the ventral plate of which is horny, the dorsal plate membranous. Body pubescent, general form elongate, sub-depressed, very staphylinous-like.

This genus appears to be extremely close to Faronus; but the species possess a well-marked process of mesosternum, separating the middle coxæ, of which there is no trace in Faronus lafertei; the frontal tubercles, also, are more approximate in Sagola, so that the distance between the antennæ at their insertion is less than in Faronus lafertei.

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Copies of descriptions of new species of Pselaphidæ from New Zealand.

Tyrus mutandus, n. sp. Rufescens, antice angustatus, sat dense setosus, impunctatus; pedibus elongatis; elytrorum striâ suturali basi foveolato. Long. 1 ¾ M.M.

Antennæ, longer than head and thorax, reddish yellow, first and second joints about equally stout; first, longer than second; second, about as long as broad; third, fourth, and fifth, about equal in length, each a little longer than broad; joints six, seven, and eight, slightly shorter than the preceding joints, especially the eighth; ninth joint, stouter and longer than the eighth, about as long as broad; tenth joint, slightly broader and a little shorter than the ninth, not quite so long as broad; eleventh joint, stout, oval, as long as the two preceding ones, but stouter than they are. Head, narrow, not half the width of the elytra, the antennæ approximate at their insertion, the tubercules contiguous, but separated by a well-marked channel, on each side near the eye is a distinct fovea; the vertex is elevated, smooth, and shining. Thorax, longer than broad, only about half as broad as the elytra, much narrowed in front, behind the middle with a very deeply-impressed curved line, which terminates on each side in a deep, but illdefined impression; it has no punctures, but is clothed with a short upright pubescence. Elytra, bright reddish-yellow, much narrowed at the shoulders, each with a sutural stria which is very deeply impressed at the base, and outside this a short, deep, and broad humeral impression; they have no punctuation, but are clothed, especially about the sides, with a long, fine, upright, pale pubescence. Hind body short and convex, pubescent, the first segment only slightly longer than the second. Legs long and slender, the claws of the tarsi small.

Pselaphus pauper, n. sp. Rufo-castaneus, nitidus; capite medio impressione magnâ; prothorace elongato, impressione basali curvatâ bene distinctâ; elytris striâ discoidali sat profunde impressâ. Long. corp. 2 1/3 mm.

Obs.—P. lineata, King, peraffinis; prothoracis impressione magis profundâ, ejusque parte basali nitidâ, elytro-rumque striâ discoidali bene distinctâ, differt.

Antennæ, longer than head and thorax, the ninth joint but little thickened. Head, with the channel between the frontal processes rather broad, and terminating between the eyes in a deep impression, which is continued backwards along the vertex. Thorax, not more than half the width of the elytra, longer than broad, not much dilated in the middle, in front of the base with a deep curved impression, the part behind this shining like the rest of the upper surface. Elytra, longer than the thorax,

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much narrowed at the shoulders, each with a sutural, and a very distinct curved discoidal stria; they are quite shining and furnished with a few fine curved hairs. Hind body rather densely set with very fine depressed hairs.

Bryaxis inflata, n. sp. Pilosa, nitida, rufescens, capite prothoraceque picescentibus; vertice foveis duabus magnis; prothorace basi trifoveolato, foveis lateralibus magnis, sulco curvato profundo conjunctis; elytris abbreviatis, apice utrinque fortiter sinuatis, estriatis; abdomine balde convexo; Metasterno brevissimo. Long. corp. 2 ½ mm.

Mas., abdomine segmento 2° ventrali apice medio leviter emarginato, seg. 4° basi tuberculo parvo, 6° leviter impresso.

Antennæ pilose, rather stout, fifth joint distinctly longer than the contiguous ones; ninth joint hardly broader than the eigth; tenth transverse, nearly twice as broad as the ninth; eleventh joint large, a little broader than the tenth, distinctly pointed at the extremity. Apical joints of maxillary palpi stout. The part of the head in front of the antennæ distinctly rostrate; the upper surface of the head with two very large pubescent foveæ between the eyes. Thorax subglobose, in front of the base with a very deep curved impression, terminating on each side in a large fovea, and in its middle impressed with a small and not very distinct fovea. Elytra not longer than the thorax, rounded at the sides, and greatly narrowed at the base; convex, without striæ or humeral impression, but emarginate on each side at the extremity. Hind body very convex, all its dorsal segments about equal in length.

Bryaxis micans, n. sp. Rufescens, nitida, impunctata, setis elongatis, erectis parce vestita; capite fronte depressâ, vertice bifoveolato; prothorace elongato, simplice; elytris striâ suturali minus distinctâ, discoidali nullâ. Long. corp. 1 ¾ mm.

Mas., antennis articulo 5° magno, intus acuminato, articulis 9-11 distortis; metasterno medio impresso; trochanteribus anterioribus spinâ tenui; abdomine segmento 2° ventrali ante apicem tuberculis duobus, apice setiformibus subito recurvis. Fem., incog.

Mas., antennæ rather stout, first joint elongate, quite as long as the three following joints together, these scarcely differing from one another; fifth joint elongate, inwardly projecting and angulate; joints nine, ten, and eleven forming a distorted club; the tenth joint is broader than the ninth; but has its base cut away on one side, and its apical portion projecting; the eleventh joint is large, and it also is irregularly formed; its base being broad and oblique, and the articulation not in the middle,

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but on one side. Head depressed in front, so that the antennal tubercles are distinct and between the eyes with two distinct foveæ. Thorax narrow and elongate, longer than broad, the sides prominent in the middle, the base margined; on each side, behind the projecting part of the thorax, and obscured by it, there is a not very easily seen fovea. Elytra longer than the thorax, with a fine sutural stria, but otherwise without striæ or depressions. The whole of the upper surface is shining and impunctate, but bears some long, sparing, fine hairs.

Bryaxis dispar, n. sp. Piceo-rufa, nitida, setis elongatis tenuissimis parcius vestita; vertice bifoveolato, fronte depressâ; prothorace simplice, latitudine haud longiore; elytris striâ suturali distincta, discoidali nullâ. Long. Corp. 21/3 mm.

Mas., antennis 10—articulatis, articulis duobus ultimus estus, concavis; trochanteribus anterioribus spinâ tenui elongatâ armatis; abdomine segmento 2° ventrali ante apicem processis tenuibus duobus leviter recurvis insigne.

Mas., antennæ longer than head and thorax; first joint scarcely so long as the two following together; fifth joint longer, but scarcely stouter than the contiguous ones; eight joint small, scarcely so large as the seventh; ninth joint large, cut away on one side, so as to leave the apical portion prominent on that side; eleventh joint large, much broader in one direction than in the other, and with one of the two broad faces impressed or concave. Head with the front much depressed in the middle, and the vertex with two large foveæ. Thorax much narrower than the elytra, about as long as broad, the sides dilated a little in front of the middle, and on each side there is an indistinct fovea behind the dilated part. Elytra much longer than the thorax, with a deep and distinct sutural stria; but without other impressions. Legs long and rather slender. The whole of the upper surface is shining and impunctate, and bears some long, fine hairs.

Bryaxis deformis, n. sp. Fem., rufescens, nitida, setis elongatis tenuissimis parcius vestita; capite quadrifoveolato (foveis frontalibus antice minus discretis); prothorace simplice; elytris striâ suturali profundâ, discoidali nullâ; predibus quatuor anterioribus deformibus (tibiis extus curvatis). Long. corp. 2 mm.

This species closely resembles the Fem. of B. dispar, but has the antennæ shorter and stouter, and has two foveæ in the frontal depression, which appear quite distinct and separate when viewed from above; but less so when looked at from the front. The four front tibiæ are extremely remarkable, as from the middle to the extremity they are much bent outwards.

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This form is so remarkable that I at first supposed the legs were deformed; but, after a careful examination, I have concluded that it is more probably natural. Except for the characters mentioned above, the insect closely resembles the Female of B. dispar.

Bryaxis impar, n. sp. Rufescens, nitida, glabra, vertice bifoveolato; clypeo antice transversim impresso; elytris striâ suturali minus profundâ. Long. corp. 1 ½ mm.

Mas., antennis 10-articulatis, art. 9° maximo; metastorno late sed parum profunde impresso; abdomine segmento basali ventrali apice bituberculato.

Fem., antennis 11-articulatis, art. 9°, 10° que transversis; metasterno abdomineque simplicibus.

Antennæ, stout and short (except for the two terminal joints in the male; the basal joint short, its visible part not longer than the second joint; the ninth joint in the male excessively developed, longer than broad, and on the inside it is a little cut away at the extremity, and the tenth joint in the same sex is only about half the bulk of the ninth; in the female, the seventh and eighth joints are extremely small; the ninth joint is also very short, but much broader than the eighth; and the tenth joint, which is also short, and very transverse, is considerably broader than the ninth, the eleventh joint being comparatively large. The head is smooth and shining; it has in the frontal depression two indistinct foveæ, and the vertex has also two very small foveæ. The thorax is about as long as broad, smooth, and shining, without impressions or foveæ. The elytra are very elongate, quite smooth and shining, and show only on each a single fine sutural stria. The hind body is very short and deflexed; the legs are slender.

Bryaxis grata, n. sp. Rufescens, nitida, fere glabra; antennæ in utroque sexu 11-articulatæ, articulis penultimis parvis; clypeo antice æquali haud impresso; prothorace ante basin lineâ curvatâ impressâ, medio desinente; elytris striâ suturali distinctâ, plicâque intra-humerali obsoletâ; capite subtus medio lineâ longitudinali elevatâ valde discretâ; pedibus minus elongatis. Long. corp. 1 2/3 mm. (vix).

Mas., vertice bifoveolato; metasterno late impresso; abdomine segmento 2°, 6° que transversim foveolatis (segmento 5° medio omnino carente).

Femina, vertice æquali; metasterno abdomineque haud impressis.

Antennæ (only differing in the sexes in that those of the male are slightly longer than those of the female, with the first joint short, its visible part about as long as the second joint; third joint, more slender than, and about as long as the second joint; joints, fourth to tenth, bead-like, the

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tenth differing but little from the others; eleventh joint abruptly larger, obtusely pointed. Thorax about as long as broad, smooth and shining, without foveæ, but immediately in front of the base transversely depressed, the depression, however, leaving the middle untouched. Elytra elongate, nearly twice as long as the thorax, each with a well-marked sutural stria, and an indistinct intra-humeral impression. Hind body very short.

N. g. Dalma pubescens, n. sp. Obscure rufescens, nitidus sed pubescentia (præsertim in abdomine) obtectus; prothorace ante basin transversim impresso trifoveolatoque, medio antice minus profunde, lateribus utrinque profunde canaliculatis; elytris striâ suturali latâ et profundâ, basique profunde bi-impressis. Long. corp. 2 ½ mm.; lat. elytrorum fere 1 mm.

Mas., antennarum articulo nono maximo (undecimo paulo majore) intus apice foveâ magnâ impresso.

Fem., ant. articulo nono præcedente paulo majore.

Antennæ stout in the male, moderate in the female; about as long as head and thorax, basal joint only a little elongate; second joint stout, bead-like, about as long as broad; joints third to sixth short, bead-like; joints seven and eight in the male short and very transverse, in the female scarcely differing from the preceding joints; ninth joint in the female broader, but scarcely longer than the eighth; in the male extremely large, subquadrate and impressed on the inner side at the extremity; tenth joint short and transverse in both sexes; eleventh joint stout, obtusely pointed, moderately long; in the male slightly stouter than in the female. Head rather small (smaller in the female than in the male sex), considerably narrower than the thorax, the frontal tubercles quite distinct, short, flattened, and shining, rather widely separated; the vertex is elevated, and on each side has a fovea confluent in front with a frontal depression. The thorax is narrower than the elytra, not so long as broad, the sides rounded in front, and considerably narrowed behind; in front of the base is a deep transverse impression, which commences on each side in a large fovea, from which there proceeds forwards a longitudinal impression; on the middle of the transverse basal impression is placed a very large fovea or depression, from which a moderately distinct channel proceeds forwards, but does not reach the front of the thorax; the thorax is not punctured. The elytra are longer than the thorax, and are redder than the rest of the surface; they are impunctate, but each has a very distinct sutural stria, and outside this they are rather deeply impressed, the impression between divided into two by a well-marked, raised, longitudinal fold. The whole surface is covered with a

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fine yellowish pubescence, which is more distinct on the hind body than elsewhere.

Sagola major, n. sp. Rufescens, nitida, elytris rufis; prothorace transversim cordato; capite lato, angulis posterioribus leviter dilitatis. Long. corp. 2 ½ mm.

Mas., trochanteribus anticis prominulis acutis; abdomine segmento 6° ventrali tuberculis duobus elevatis. Fem., incog.

This species differs from S. prisca by its much broader form, by its more slender antennæ, the basal joint in particular of these organs being notably more slender, and by the more deflexed extremity of the hind body, as well as by the different characters of the male. The first visible dorsal segment of the hind body possesses a transverse band of glandular pubescence, which is wanting in the other species here described.

Sagola prisca, n. sp. Obscure rufa, elytris sanguineis, capite thoraceque parce, longius, abdomine dense pubescentibus; antennis crassiusculis, articulis quatuor penultimis leviter transversis; capite angulis posterioribus rotundatis. Long. corp. 2 ½ mm.

Mas., abdomine segmentis 3°, 4° que apice tuberculis duobus elongatis, 5° transversim depresso, apice emarginato.

Antennæ with the first joint stout and elongate, second joint small, sub-globular; third joint similar in shape to second, but still smaller than it; joints fourth to tenth differing little from one another; eleventh joint hardly as broad as the tenth, but a little longer than it, obtusely pointed. Head small and short, with two small foveæ on the vertex, and with a fine channel separating the short, flattened, frontal tubercles; this channel expanding a little behind, so as to appear as if it terminated in a very small fovea. Thorax subcordate, with a large quadrate impression on the disc behind the middle, and, close to each hind angle of this, a very small fovea, and with a larger fovea on each side. Elytra about one and a-half times as long as the thorax, a little narrowed towards the shoulders, each with a sutural stria, which towards the base is very deeply impressed, and between this and the shoulder with a coarse, elongate impression; this impression appears to be nearly divided into two near its base. The hind body is broad, and its exposed portion is slightly longer than the elytra.

Sagola misella, n. sp. Obscure rufa, elytris sanguineis; antennis articulis penultimis vix transversis; elytris abdomine multo brevioribus. Long. corp. 2 ½ mm.

Mas., a femina notis sexualibus externis vix distinguendus.

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This species is very closely allied to S. prisca, but is readily distinguished therefrom, by its much shorter elytra and metasternum; its antennæ are also more slender, and their fifth joint is notably thinner; the hind body is broader towards the extremity; and the remarkably conspicuous male characters of S. prisca are in S. misella entirely wanting.

Sagola parva, n. sp. Corpore antice fortiter angustato. Obscure rufa, elytris sanguineis; antennis sat gracilibus, articulis penultimis vix transversis; prothorace elongato, latitudine fere longiore; elytris abbreviatis, abdomine multo brevioribus, prothorace vix longioribus. Long. corp. 2 1/3 mm.

Very closely allied to S. misella, but with the head and thorax narrower, and the elytra a little shorter than in that species; the antennæ also are rather less developed than in S. misella, being both a little shorter and more slender.

Euplectus convexus, n. sp. Rufescens, pube brevi depressâ, dense vestitus; fronte profunde bisulcatâ; prothorace angustulo, basin versus impressionibus tribus magnis; elytris striâ suturali basi profunde impressâ; impressioneque intra-humerali bene distinctâ; antennis articulo ultimo acuminato. Long. corp. 2 1/3 mm.

Mas., pedibus omnibus incrassatis, tibüs posterioribus intus angulatis.

Antennæ, shorter than head and thorax, second joint not so long as first; joints third to ninth bead-like, differing little from one another, except that the ninth is a little broader than the others; tenth joint, short, rather strongly transverse, about twice as broad as the ninth; eleventh joint, large, broader than the tenth. Head, rather long and narrow, very deeply impressed between the frontal tubercles; from each side of the impression proceeds backwards a deep furrow, which terminates between the eyes as a fovea-like expansion. Thorax much narrower than the elytra, about as long as broad, much narrowed behind, with a very large impression behind the middle, which is connected on each side with a deep large fovea near the hind angles. Elytra distinctly longer than the thorax, with the sutural stria deeply impressed at the base, and with a rather large intrahumeral impression.

Euplectus opacus, n. sp. Rufescens, opacus, pube brevissimâ densius vestitus; antennæ breves; capite parvo, transversim impresso; prothorace basin versus impresso, disco canaliculato; elytris striâ suturali, alterâque subtili, discoidali, abbreviatâ, basi profunde impressis. Long. corp. 1 1/3 mm.

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Antennæ, shorter than head and thorax, second joint a little shorter than the first, subglobose; joints, third to eighth very small; ninth joint broader than its predecessors, transverse; tenth joint, broader than the ninth, strongly transverse; eleventh joint, stout. Head very short, a large portion of its upper surface occupied by a curved or angulated transverse impression. Thorax short, not so long as broad; in front of the base it has a deep curved impression, which is indistinctly expanded in the middle and on each side, and in front of this there is a longitudinal impression on the disc. Elytra, longer than the thorax, with a distinct sutural stria and a fine abbreviated discoidal stria, these striæ being deeply impressed or foveolate at their commencement. Legs, rather short.