
Art. XXXI.—Notes on Regalecus pacificus, a new Species of Ribbon Fish from the New Zealand Seas.
Plate VII.
[Read before the Philosophical Institute of Canterbury, 15th December, 1876.]
Mrs. George Oram, of New Brighton, kindly informed me on the evening of May the 7th of this year (1876), that in the morning of that day a gigantic frost-fish had been found on the beach near the hotel, to which it had been brought, and that she would be very glad if I would come and inspect it. The next morning I proceeded there, and found the specimen in question, with a few minor exceptions, in a splendid state of preservation; and as that lady has kindly presented it to the Museum, I am not only able to offer a description of this remarkable fish, but my friend Dr. L. Powell, F.L.S., has also prepared a careful drawing, and offers in an appendix a few remarks on its anatomy and on the results of a microscopical examination of its so-called scales.

On measurement I found this beautiful fish to be 12 feet 5 inches long, having an average breadth of 13.5 inches and a thickness of 3.5 inches. The whole skin is covered with a coating looking as if it consisted of frosted silver, and which adheres to the fingers very easily. The brilliancy of this silvery appearance is still more prominently brought out by numerous tubercles which stand above the skin, and by a number of irregular black lines and spots in the anterior portion of the body.
If we add to this the fine metallic iridescence of the head, which possesses at the same time a crest of bright red spines on the top, it is not too much to say that the form and colouring of this species make it the most beautiful fish ever obtained on the coasts of New Zealand.
The capture of these greatly compressed, sword-like, deep-sea fishes occurring at such long intervals, and of which so little is known, has always excited the curiosity of the public and the deep interest of the naturalist; and cases when specimens of this genus were secured in England and on the continent of Europe were generally noted in the publications of the day.
On examining the literature dealing with this genus, I find that this specimen is new to science, and I therefore propose to describe it here under the specific name of Regalecus pacificus. It is very different from the Regalecus described by Mr. W. T. L. Travers, which towards the middle of October, 1860, was stranded on the beach near the entrance of Nelson harbour, as “from the lower lip of that specimen depended a large number of rigid slender barbules about sixteen inches long and of a brilliant red colour.”*
The description given in that standard work proves that very little is known of these occasional visitors from the deep seas, as there are six species enumerated, of which most have been described from single and generally mutilated specimens only.
Regalecus Pacificus, Haast.
B. 6 D. 9/223 P. 12
The length of the head is 7.75 inches, and height behind the eye 6.75 inches, consequently the length of the head is one-nineteenth of the total. The height of the body is—
| 2 feet from snout | 13.25 inches |
| 3 " 9 inches from snout | 13.75 |
| 4 " " 11 at anus | 13.25 |
| after which it tapers gradually to the tail. | |
| 3 feet from tail | 9.50 |
| 1 " | 6.25 |
[Footnote] * Günther's Catalogue of Acanthopterygian Fishes, Brit. Mus., III. 307.

Consequently the greatest height of the body to the length is nearly one eleventh. The greatest thickness of the body along the line of the vertebæ, 3 feet 6 inches from the snout, is 3.5 inches, gradually it increases to 4 inches at 4 feet 2 inches from the snout, a thickness it sustains for 4 feet more, after which it gradually diminishes till 3 feet 6 inches from the tail it has dwindled to 2.25 inches. The ventral edge of the body is broader than the dorsal one. Colour generally silvery, with a tinge of lavender. When the fine silvery coating is still attached, the tuberculated bands look lighter than those on which the tubercles are very minute. On the other hand, when this coating is rubbed off alike, the smooth bands look the lightest. In the latter case the tuberculated bands appear steel-coloured, on which the tubercles stand prominently with a silvery white lustre. On both sides are a number of blackish irregular stripes and blots, mostly elongate in the anterior portion, gradually decreasing and ceasing near the anus. The whole skin is also covered with dark rings, mostly of a round form, inclosing a lighter space. They are generally 1.5 inch in diameter and stand about one inch from each other.
Snout truncated with a very protractile upper jaw and a vertical cleft of the mouth. No teeth. The eye, which has a somewhat oval form, the largest diameter being in the horizontal direction, is 1.25 inch broad, or a little more than ⅙ of the length of the head. The horizontal diameter of the pupil is .52 inch. The three operculums, the supra-orbital, suborbital, and prefrontal, as well as the remarkable maxillary bones, possess a radiated structure on their surface, which is covered with a very thin skin, breaking with the least touch.
The frontal and nasal bones are smooth and covered with skin of a dark steel colour, showing the same iridescence as the former.
The cheeks, the supra-scapular and temporal bones are covered with tuberculated skin which has also the same dark bluish colour as the rest of the head.
The dorsal fin consists of two portions, of which the first nine spines form a crest. These spines enlarge at their termination to a lobe, as shown by the two only perfect ones when the fish was obtained; they cover a space of 2.5 inches. The first of these spines is broken off at 3 inches from the base; it is the stoutest of the whole series.
No. 2 is considerably thinner and 7 inches long. It is one of the complete ones. The next three spines (3, 4, and 5) were all broken off at 4 to 6 inches and were nearly as thick as the first. From here they get thinner, the thickness of the seventh having only the thickness of the second. This spine which is entire, is 7.75 inches long, and has, like the second, a lobe at its termination. The eighth is still thinner and broken

off one inch from its base, and there is only a fragment of the ninth, which is not thicker than one of the rays of the dorsal fin proper. All of these spines which have minute hooks directed upwards on their anterior and posterior edges are united with each other by a small membrane about .45 inch high. They had, like the two ventral rays, a red colour, very bright in their upper portion when the fish was first obtained, which, however, gradually faded to a dull light pink.
A quarter of an inch behind the last spine the dorsal fin proper begins, consisting of a great number of slender rays, which do not project beyond the margin of the thin membrane by which they are united. These rays stand at a distance of three-quarters of an inch from each other. At the beginning they are about one inch high, gradually increasing in length, till about one foot from the beginning of this dorsal fin they stand 2.5 inches high, continuing so to within 3 feet from the end of the tail; they after this gradually get smaller, till 5.5 inches from the end of the tail, they cease altogether. Here at the termination the last four rays stand not only above the membrane, but they are also thicker than all the preceding ones. The pectoral fin is one inch broad at the base; 2.5 inches high along the first ray. It also had a reddish hue, but not so bright as the crest and the ventral rays.
In a straight line below the hind edge of the pectoral fin stands a single strong ventral ray on both sides, of which, however, only 3 inches are preserved, the rest being broken off.
I was informed by the finder that it had been at least 15 inches long, but he did not observe any enlargement at the termination. I believe that, when the fish was first seen, the anterior portion of this remarkable ventral ray with the lobe had been already broken off during the efforts the fish made to regain the sea.
Tail without any caudal fin, being formed by a number of spines, getting gradually thinner and tending to a point.
Anal vent 4 feet 1 inch behind the ventral spines.
The lateral line is formed by a number of elongated smooth scales, which lie in a groove. It starts from the posterior end of the supraorbital, gradually descending, so that at the anus it is situated only 3.5 inches above it. Four longitudinal ridges which are cut off obliquely by the lateral line extend along the whole length of the body, they are covered with small round tubercles and are about three-quarters of an inch broad, the space between them having nearly the same breadth. This space is also covered with tubercles, which, however, are very minute. Below the latter the intermuscular septa are situated. The skin below the lateral line is almost smooth for about an inch, after which the tubercles appear again, getting

gradually more numerous and larger, till we reach the ventral line, where they become very large and well raised, possessing in the posterior half a small hook turned backwards.
Postscript, 22nd December, 1877.—The following account taken from a New Zealand newspaper about the middle of July, 1877, shows that another specimen of Regalecus has been washed ashore on the west coast of this is and. From the short description in that paragraph it appears that the fish in question is either a specimen of Regalecus pacificus, or at least belongs to a nearly allied species, the main difference being that the first portion of the dorsal fin in the west coast specimen has ten spines instead of nine as in the former. It is also interesting to observe that each of the two strong ventral rays was two feet long, but no data are given from which we can conclude that they were intact:—” An extraordinary fish was picked up on the Little Waimangaroa beach, Karamea District, by Mr. Alexander McDonald, on Tuesday last, of which the following description is furnished by the Westport Times:—Length, 14 feet 4 inches; girth, 2 feet 7 inches; one dorsal fin extending from head to tail; from the top of the head there extended ten feelers, each one foot in length, and two similar appendages, about two feet in length, grew underneath the jaw, resembing in appearance the feelers of a lobster; on each side of the head there was one small fin, and the gills of the fish resemble those of a turbot; the body was of a bright silvery hue and covered with fine scales, the shape of the body being more flat than round; the eyes resembled those of a cod-fish. When found it had only recently been stranded, as it was not quite dead. This is probably some large species of frost-fish.”

