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Volume 27, 1894
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[Read before the Wellington Philosophical Society, 14th November, 1894.]

Plates I.–VII.

I Find it necessary to make some remarks concerning the principle upon which I have acted for several years with regard to the erection of new species, the comparison of varieties, or the identification of species already known; because, in the course of correspondence with different other students of Coccids, as well as in their published writings, it seems that they accept canons of procedure which do not recommend themselves to me as sufficient. In the first place, I may say that the bent of my inclination is towards the restriction of the number of species rather than towards their extension. Looking upon classification as a means to an end, I consider that the very first, and probably only, requisites for it are clearness and convenience. For this reason, neither the fancied claims of priority, nor the desire to uphold some preconceived theory, nor even the wish to be the publisher to the world of some new-thing, ought to weigh with us at all. Personally, I disclaim any position but that which some people nowadays affect to despise—the position of a classifier, the laying-down of a stratum of facts upon which, some day, when we really do reach the stage of being able to say we know something, theories and doctrines may be built up. Therefore my aim is first at clearness and convenience; and with this object a careful avoidance of needless subdivision seems necessary. It is better, when a specimen is observed, to see how closely it approaches to some known form than to search for points upon which it may be separated—better to look upon a few distinguishable features, if possible, as only variable characters (making the specimen a “variety”) than to cumber science with a new “species” as if these features were of real organic importance. I know, and every student of Coccids knows, that external appearance of an insect may alter frequently, from being cramped for room or from having much free space: therefore I would lay small stress upon mere size. Take the genus Lecanium: there are a number of so-called species which have been set up by different authors almost, if not quite, entirely on account of their varying size: climate,

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food-plant, conditions of life, are in this case ignored. Again, we all know how deceptive a thing colour is: what is green to one man may be blue or red to another. Ornithologists and lepidopterists have invented thousands of new “species” on account of some different colour in a feather or in a wing-spot. But many cases are known in Coccid study where, even on the same twig, amongst perhaps a hundred individuals, five different “species” could be so made because there may be five variations in colour. Again, in the case of climate and food-plant: if we were to imagine that an insect in a tropical country on, say, a palm, must necessarily be for that reason a different species from one in a temperate country on a Fagus, it would be the simplest thing in the world to bring up the number of Coccid species to that of the Hymenoptera, ransacking every dictionary in every language to find names for them. A friend of mine once wrote that he could not agree to make two Planchonice identical “because they lived on such widely-separated orders of plants as Coronilla and Bambusa.” But there are dozens of Coccids which are multivorous if not omnivorous: for example—Aspidiotus nerii, A. aurantii, Mytilaspis pomorum, Ctenochiton viridis, Lecanium hesperidum, Planchonia fimbriata, several Dactylopii, Icerya purchasi, &c. You cannot predicate that an insect living on one plant will not live equally well on another, or in another locality. Taking, therefore, these four points—size, colour, locality or climate, and food-plant—I would very seriously deprecate the notion that any of them (I will go so far as to say that all of them) should be sufficient to induce the erection of a new species. Convenience demands that they should be all mentioned in a description, as a guide to other students or as information to the public, so that anybody could know where at least to search for particular insects. Beyond that I look on them as of little or no value.

The following pages contain notes concerning many species already reported from various parts of the world, and also descriptions of more than twenty insects which, after full consideration, seem to be new. Some, such as the Australian form of Eriococcus buxi, or the Sandwich Island form of Chionaspis biclavis, I have set down as merely varieties of the original types. The others, although seeming to be quite distinct, do not present such extraordinary and interesting features as those reported in my former papers. But the discovery in Australia of representatives of the genus Ctenochiton (hitherto confined to New Zealand) seems to be not entirely unimportant in its bearing on the distribution of animal forms and the connection between the two countries.

With regard to Ceronema, the new genus which I am here proposing to establish, it may be remarked that, whilst the

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secretion of all Coccids appears to be chemically the same, yet it is sometimes convenient to employ for separation the solidity or the loose texture of the resulting covering. The “wax” of Ceroplastes, the cotton of Dactylopius, and the fibrous (as it seems at first sight) puparium of Mytilaspis, can all, when microscopically examined, be found to be really similar in their essential character. Still, it is convenient to distinguish (for example) Lichtensia from Pulvinaria, or Ctenochiton from Inglisia, to some extent by the form, position, or structure of the secretion. I say “to some extent” because if possible the main distinction of a genus or a species ought to rest upon an organic character of the insect itself rather than upon its external covering. Ceronema differs from Ceroplastes and Ctenochiton in the loose texture of the test, and from the latter also in the absence of a fringe: but the secreting pores of the insect itself (in the species known so far) also differ from those of any other Lecaniodiaspid.

The species herein named Prosopophora prosopidis is close enough to P. dendrobii Douglas to be perhaps hereafter considered as only a variety; but for reasons given in its description I leave it for the present as distinct.

It appears from the synoptical list published in this volume that I am responsible for more than two hundred new species and varieties of Coccids reported from Australasia and the Pacific, besides a large number originally named by other writers and mentioned in my papers. I am sure that future years will produce a very much longer list, for the study of these insects is even yet only in its infancy; but there will be, I hope, no need for any one to make any important or subversive corrections in the work which has been so far accomplished.