
Art. XX.—Notes on New Zealand Land Planarians: Part III.*
[Read before the Philosophical Institute of Canterbury, 30th November, 1896.]
The following notes deal with a number of Land Planarians received from various parts of New Zealand during the past year. The majority were collected by myself on a trip to the west coast of the South Island in January, 1896, at Otira, Jackson's, Kumara, and Lake Mahinapua, near Hokitika. I am indebted to Sir James Hector for a large specimen of G. triangulata var. australis, from Nelson; to Mr. H. Suter for some very interesting specimens collected by himself at Dunedin; and to Mr. W. T. Locke Travers for valuable specimens from Wellington.
Geoplana sanguinea, Moseley.
Specimens of this widely-distributed species were met with in abundance under wood lying on the ground in paddocks between Lake Mahinapua and the sea-shore. I also found, a single specimen at Kumara Junction. There is some difficulty in distinguishing between this species and G. triangulata var. australis, especially after preservation in spirit, when the dark purplish-brown colour of the latter has faded, and the pale supra-marginal bands are no longer distinct. I cannot help suspecting that, while G. triangulata var. australis is truly endemic, G. sanguinea may, after all, have been introduced from Australia. Its great abundance on cleared ground on the West Coast in company with introduced slugs (Limax agrestis and Amalia gagates) is very suggestive.
Geoplana triangulate var. australis, Dendy.
I have received from Sir James Hector a large specimen preserved in alcohol which is evidently referable to this variety. It was collected at Nelson, and is, I believe, the specimen referred to by Sir James Hector in the “Transactions of the New Zealand Institute,” vol. xxv., p. 255. It is a singular fact that this unspeckled variety should occur both north and south of Christchurch, while in the Christchurch neighbourhood itself the speckled
[Footnote] * See Trans. N.Z. Inst., vol. xxvii., art. xvii., and vol. xxviii., art. xxii.

form, originally described as the type of the species, is extremely common and the other apparently entirely absent. The dark-brownish colour of the dorsal surface, with its narrow pale-coloured supra-marginal bands, is still very distinct in the Nelson specimen, even after a lapse of some years, and leaves no doubt in my mind as to the correctness of this identification.
Geoplana flavimarginata, n. sp.
Body very long and narrow, especially when crawling, when an average-sized specimen measures as much as 80mm. long by only 2mm. in breadth. In spirit the body is still long and narrow, flat below and convex above, about 40mm. long and 3mm. broad, with the peripharyngeal aperture 13mm. and the genital aperture 6mm. from the posterior extremity.
Dorsal surface in life very rich deep crimson or Indian red, with a narrow, sharply-defined, supra-marginal yellow band on each side, which contrasts strongly with the red. Anterior tip pink. Ventral surface yellow all over.
The eyes are numerous, but very minute and inconspicuous. The movements of the living worm are very active.
This species appears to be intermediate between G. sanguinea and G. triangulata var. australis. It is distinguished from the former by the very well defined yellow supra-marginal bands and by the narrower body, and from the latter by the much narrower body and the bright-red colour. From average specimens of both it is distinguished by its smaller size.
About a dozen specimens were sent to me alive by Mr. W. T. Locke Travers, of Wellington, who found them beneath a damp mat at the door of his greenhouse in May and June, 1896. As all the specimens agree closely with one another, I feel justified in distinguishing them by a new specific name. The locality suggests that the species may possibly be introduced.
Geoplana cucullata,* n. sp.
Body in life much flattened when at rest, but with no noticeable peculiarity in shape. In spirit, however, the shape is very characteristic: only about three times as long as broad, fattened on both surfaces, and with narrow crenate lateral margins; the anterior, tip strongly curved ventralwards, beaklike, with smooth incurved margin forming a kind of hood, deeply excavated beneath and bearing the eyes along its edge; the posterior extremity generally broadly rounded, sometimes tapering rather abruptly.
[Footnote] *Cucullata, hooded.

The largest specimen measures, in spirit, 20mm. in length by 7mm. in breadth. Another specimen measures, in spirit, 15mm. by 5mm., while in life when crawling it measured about 34mm. by 3mm.
The peripharyngeal aperture, in spirit, is situated slightly in front of the junction of the middle and posterior thirds of the body, and the genital aperture is about half-way between it and the posterior end.
The eyes are very distinct, but arranged only in single series, or approximately so, around the anterior margin, and continued sparingly backwards along the lateral margins.
Dorsal surface in life brownish-yellow minutely speckled with warm brown, with paler narrow margins and a very narrow, ill-defined, pale median line. Ventral surface nearly white, faintly and minutely speckled with pale brown, except at the anterior end. Anterior tip very pale yellow or brownish yellow.
This remarkable species is certainly nearly related to Geoplana latissima, described in Part II. of these notes, from Springburn, but is distinguished by the narrower body, the more pronounced hood, and especially the colour markings; perhaps also by the absence of the median notch in the posterior margin.
I collected six specimens near Lake Ma [ unclear: ] hinapua, under fallen logs.
Geoplana graffiti var. occidentalis, nov.
Body at rest flattened above and below, tapering gradually in front, more suddenly behind. Dorsal surface with a narrow median stripe of dull orange or buff, very minutely speckled with brown, or even with a median very narrow brown line almost denned in the middle of the orange. The orange band is followed on each side by a broad band of dark-purplish tint thickly dusted with minute specks of bluish lustre, and first fading and then intensified on its outer edge to form a narrow band of nearly black colour, followed by the pale grey or buff margin of the body speckled minutely with darker grey or brown. Anterior tip brownish red. Ventral surface light brown, sometimes with an ill-defined lighter median band, and minutely speckled all over with darker brown. Size of average specimen about 53mm. by 4mm. when crawling, and 36mm. by 6mm. in spirit; genital aperture 11mm. and peripharyngeal aperture 16mm. from the posterior end in the same specimen in spirit.
This western variety is distinguished from the eastern types of the species by its larger average size, more speckled character, and more strongly developed dark band above the light marginal band. Though difficult to express in writing,

the difference is sufficiently striking and constant in actual specimens to merit the use of a distinctive varietal name.
I found four specimens of this variety at Otira, and six near Lake Mahinapua.
Geoplana graffiti var. otiraensis, nov.
Body when at rest in life broad and much flattened, but convex above and somewhat triangular in section. When crawling, tapering very gradually in front, much less so behind. In spirit, flat or slightly concave below, more or less arched above, broader behind than in front, tapering very gradually to the narrow anterior extremity, more suddenly to the blunter posterior extremity.
A fair-sized specimen, in spirit, measures 33mm. in length by 6mm. in greatest breadth.
The peripharyngeal aperture, in spirit, is only very slightly behind the middle of the body, and the genital aperture only very slightly behind the junction of the middle and posterior thirds.
Dorsal surface in life dark-grey, intensified to black in the middle line, flecked all over with small whitish, greenish, or yellowish specks, varying in distinctness in different specimens. Anterior tip dark-grey. Ventral surface brown, minutely speckled with a darker tint, and with a paler-coloured narrow median band. Very narrow brown margins.
Two somewhat larger specimens, apparently of the same variety, have the pale median ventral and dark median dorsal bands either absent or very obscure, and one of them has an intensified supra-marginal band on each side of the dorsal surface.
Eyes numerous, and arranged as usual, but difficult to make out owing to the dark colour of the body.
This variety comes very near to G. graffii var. somersii, described in Part II. of these notes; but the present specimens are much larger, and differ slightly in pattern from those of the Springburn variety.
Five specimens were obtained at Otira, and one near Lake Mahinapua.
Geoplana moseleyi, Hutton.
I have received from Mr. Suter several specimens preserved in spirit which I have little doubt are referable to Hutton's species. They were collected at Dunedin, where Hutton's original type was also obtained. They agree very closely with Hutton's description, but the degree of development of the median dorsal stripe is variable, sometimes broad and sometimes very narrow, while the “interrupted narrow

black line” mentioned by Hutton as occurring near the margin of the stripe is also variable in its development.
I have long suspected that my G. graffii must be nearly-related to G. moseleyi, but in the absence of more detailed information as to the latter I did not venture upon an identification. I now find that the species, assuming my present identification of Mr. Suter's specimens to be correct, are indeed very near together, the only tangible distinction lying in the paler-coloured ventral surface of G. moseleyi, and the absence from the ventral surface of any distinct speckling and of the distinct pale median band which characterize G. graffii. G. graffii also occurs at Dunedin, as I have, previously recorded.
Perhaps in the future it may be desirable to unite the several varieties of G. graffii which I have described, together with G. iris and the Dunedin form described by Hutton, under the one name, G. moseleyi, which will then have to be regarded as a widely-spread species with numerous local varieties.
Geoplana Inæqualistriata, b. Dendy.
I have found several more specimens of this species in my garden near Christchurch since writing the last part of these notes. I also obtained one planarian at Jackson's, on the Teremakau, which may possibly belong to the species, but it is not quite typical, and in some respects resembles G. graffii.
Geoplana jacksoniana, n. sp.
Body at rest in life much flattened, slightly convex above, very broad, tapering equally in front and behind. Movements very sluggish. Dorsal surface rich-brown marbled with short longitudinal dashes of grey and dull-orange, the orange dashes more abundant and paler in the middle line and above the margins. Ventral surface pale reddish-brown very minutely dusted with darker grey, except below the margins, where the brown is more intense; with indications of a pale narrow median line. Anterior tip grey.
In spirit the body is strongly convex dorsally and concave ventrally, 43mm. long by 6.5mm. broad; genital aperture 15mm. and peripharyngeal aperture 21mm. from the posterior end. Eyes as usual, with crowded lateral patches.
This species appears to be nearly related to G. gelatinosa, of which it may prove to be a variety, but the specimens look so different when placed side by side that I do not care to unite them under a common name. The difference lies chiefly in the thicker body and elongated colour-dashes of G. jacksoniana, which is also considerably larger, and has the external apertures more anteriorly placed. The sudden

tapering of the anterior extremity in G. gelatinosa also seems to be distinctive, even in spirit. In spirit, also, the colour of the two specimens is very different, but, as this may be due to different degrees of fading, I say nothing about it at present.
The single specimen was obtained at Jackson's, on the Teremakau, under a sawn log at the battery, some way up the mountain-side.
Geoplana splendens, Dendy.
This handsome species was originally described from a single specimen obtained at Jackson's by my sister, and it gave me great pleasure to meet with it again both at Jackson's and Otira. It may be at once recognised, by the three bright-green stripes on the dark-brown background of the dorsal surface. I only found a single specimen at each of the above-mentioned localities.
Geoplana mariæ, Dendy.
I obtained one specimen of this species at Otira and four or five near Lake Mahinapua.
Geoplana subquadrangulata, Dendy.
I have now to record this common East Coast species from Otira, Jackson's, and Lake Mahinapua, on the West Coast, though it was not abundant at any of these localities. The Otira specimen was very dark-coloured, and had a faint green speckled lustre on the back, and only the narrow median dorsal stripe was recognisable.
I have also received varieties of this species collected at Dunedin by Mr. Suter.
Geoplana suteri, n. sp.
Body when crawling long and narrow; approximately oval in transverse section, flattened above and below; tapering gradually in front and behind. One specimen, when crawling, measured 44mm. in length by little over 2mm. in breadth, and the same, in spirit, 32mm. by 3mm.
In spirit the body is flattened above, more convex below, of approximately uniform width throughout, with the peripharyngeal aperture in about the middle, and the genital about half-way between it and the posterior extremity.
Eyes numerous, arranged as usual in antero-lateral patches, continued in approximately single series round the anterior tip.
In life the ground-colour of the dorsal surface was pale yellow-brown with six very narrow dark-brown stripes arranged in pairs, two being close together, one on each side of the middle line, and two not quite so close together, just

above each lateral margin and separated from the middle pair by an interval about twice as wide as that between themselves. The outermost stripe on each side may be less distinct than the others.
Ventral surface white, without markings. Anterior tip pinkish-brown.
I have named the species after Mr. H. Suter, who found it at Dunedin and gave me three specimens, one of which I received alive and two in spirit.
