
II.—Zoology.
Art. XII.—Additions to the Marine Mollusca of New Zealand.
[Read before the Wellington Philosophical Society, 2nd November, 1904.]
Plates VII. and VIII.
Some few months ago I received from Mr. A. Hamilton Director of the Colonial Museum, an interesting parcel of small and minute shells, which had been obtained by beach-gatherings at Whangaroa Harbour. This collection, though small in bulk, proves exceedingly rich in species. None of the larger forms are represented, and very few of those of medium size: it must therefore be regarded as little more than a tithe of the molluscan life of this northern locality. Several forms hitherto only recorded from Foveaux Strait and Stewart Island make their appearance, some of them in fair abundance.
In addition to the species recorded in the appended list, there are several which I have been unable to identify. Some of them may prove to be undescribed; others, again, are in too worn and damaged a condition—these include Bittium, Colina(?), and Triphora. Of the latter genus three species are represented, one of which has certainly not been recorded from New Zealand waters; it is little more than a fragment, but characterized by its four smooth spiral keels. Of Cryptodon flexuosum, Mont., a single valve occurs. The species is represented in the Colonial Museum collections by a number of separate valves, but the exact locality does not appear to have been given. Most probably they are from Stewart Island, as Captain Hutton records Cryptodon, sp. ind., from that locality.* The genus does not appear to have been again recognised, and in the “Manual of the New Zealand Mollusca,” page 223, is rejected from the list.
The identification of many of our marine shells, perhaps more especially the smaller forms, is decidedly difficult. There is an absence of figures, and the descriptions, unfortunately, are exceedingly brief.
My best thanks are due to Captain Hutton, Mr. Henry Suter, of Auckland, and Mr. Charles Hedley, of the Australian Museum, for much valuable assistance.
[Footnote] * Cat. Marine Moll. N.Z., p. 75.

Cylichna striata, Hutton. Figs. 1, 2.
Cylichna striata, Hutton, Cat. Marine Moll. N.Z., p. 52 (1873); Cat. Tertiary Moll. and Echin. N.Z., p. 16; Manual N.Z. Moll., p. 120; Trans. N.Z. Inst., vol. xviii., p. 346. C. arachis, Hutton, “Macleay Memorial Volume,” p. 37 (not of Quoy and Gaimard). C. striata, Hutton, “Index Faunæ Novæ-Zealandiæ,” p. 69 (not striata, Pilsbry, Man. Conch. (1), vol. xv., p. 319, pl. 59, figs. 11, 12).
The types of this species are in the Colonial Museum. I have been unable to examine them, but Mr. C. Freyberg kindly did so at my request. They appear to be in good condition, sculptured with longitudinal growth-periods distinctly marked, but a total absence of spiral striæ; in fact, agree perfectly in all characters with the Whangaroa Harbour examples, of which there is a good series.
Pilsbry, when dealing with the specimens he had received from New Zealand, evidently had some doubts as to whether they were the true striata, remarking that the measurements given by Hutton indicated a much smaller form than his specimens. In addition to the larger size of his specimens, the measurements prove that they were of a more narrow, slender form. I have no doubt they were C. arachis, Quoy and Gaimard, with which the description agrees very well indeed. Examples of the latter species from New Zealand (I have only seen two) appear to be rather more minutely spirally striate than those from Port Jackson.
The figures and following description of C. striata is derived from the Whangaroa specimens: Shell short, subcylindrical, thin and white; last half-volution with a lightly concave appearance, more noticeable on approaching the lip. Sculptured with numerous well-marked longitudinal growth-periods, irregularly spaced. The crown concave, with a deep and moderately broad axial perforation. Aperture as long as the shell, narrow above, expanded and effuse below; the inner wall with a very thin callus. Columella thickened, slightly twisted, anteriorly rapidly declining, giving it an almost truncated appearance. In the juvenile stages the columella is usually more gently sloping; the crown with the same crater-like concavity, but the axial perforation shallow; the outer lip almost straight. Dimensions of largest specimen: Length, 3.8 mm.; breadth, 1.87 mm.
It is perhaps nearest to C. pygmœa, A. Adams, from Australia, but this species is adorned with well-marked somewhat distantly spaced spiral striæ.

Clathurella epentroma, n. sp. Figs. 3, 4.
Shell small, narrowly fusiform, the spire slender. Colour light reddish-brown or dull-chestnut. Whorls 5½, rounded or obscurely angled above the periphery, adorned with fine spiral and longitudinal sculpture, the latter strongest. Sutures deep. Protoconch of about one and a half whorls, strongly angled, and with four smooth narrow revolving riblets, the posterior minute and situate near to the suture; the apical half-turn obliquely curved down and somewhat imbedded in the succeeding whorl. The last whorl rather longer than the spire. Adult sculpture: The longitudinals number fifteen to sixteen small riblets on the last whorl, equal to or rather wider than the interspaces, and usually less developed on the anterior end; they are continuous in some, irregular in others. The spirals consist of undulating delicate riblets and threads. On the spire-whorls there are two, and on the last seven or eight slightly stronger; of these the four posterior are more widely spaced, two are above the outer lip and one in line with it; frequently forming beads on crossing the longitudinals. Within these spaces there are, on that adjoining the suture, four or five threadlets, sometime irregular in size; on the succeeding three spaces, usually three threadlets in each, the median one frequently strongest. Anterior to this the interspaces with one to three threads. The old beaks sometimes forming short irregular riblets. Aperture of medium breadth, less than half its length. Outer lip simple, the posterior sinus shallow. Columella lightly curved, the anterior canal short and broad. Length, 5.73 mm.; breadth, 2.11 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
This is a very beautifully sculptured little shell. The spirals on the anterior portion of the last whorl are somewhat variable in number and strength. The lightly angular appearance of the spire whorls in some individuals is due to the more prominent spirals being rather more pronounced; the spaces on these whorls are similarly adorned to the posterior portion of the last.
Clathurella epentroma, Murdoch, var. whangaroaensis, n. var. Fig. 5.
Differs from the typical form in the whorls being strongly angled, spire somewhat turrited, and the principal spiral riblets much more pronounced, but similarly placed. Colour, number of whorls, and protoconch the same as typical. Longitudinal ribs fourteen on the last whorl, equal to or narrower than the interspaces. Spirals—on the spire whorls two, and on the last four or five, prominent and forming beads on crossing the longitudinals.

In addition there are on the anterior end a few more crowded and rather smaller. The interspaces with spiral strlæ, four or more, usually minute, sometimes almost microscopic. Outer lip with usually five well-marked sinuations corresponding to the principal spirals, posterior sinus shallow. Length, 6.83 mm.; breadth, 2.5 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
This form appears to somewhat approach Mangilia connectans, Sowb.,* from South Australia.
Mitromorpha suteri, n. sp. Fig. 6.
Shell small, fusiform, somewhat thin, with fine spiral and usually somewhat obsolete longitudinal riblets. Colour light reddish - brown, sometimes a pale band around the periphery, occasionally a narrow darker band at the sutures. Whorls 5, lightly rounded, the last longer than the spire. Protoconch of two whorls, somewhat globose, smooth and polished, the apical turn oblique to the succeeding whorl. Sculpture: The penultimate whorl with six to seven and the last with sixteen to twenty spiral riblets, seven or eight of which are in front of the aperture; they are slightly variable in strength, some in breadth equal to the interspaces, others rather narrower; also an occasional small thread here and there arises in the interspaces. Longitudinals irregular, low and rounded, more distinct on the spire, frequently obsolete. The growth striæ irregular, somewhat marked, and frequently cutting up the spirals into minute gemmules. Sutures impressed, usually margined with a wider riblet. Aperture somewhat narrow, rather longer than the spire. Outer lip thin, a little flattened; the posterior sinus broad, well marked. Columella almost straight, concave and lightly callused; the canal short and broad. Length, 4.56 mm.; breadth, 1.70 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
At first glance this species might easily pass for a Columbella. From M. substriata, Suter, it is at once distinguished by its colour and much stronger spiral sculpture. It appears to be nearest to M. subabnormis,† Suter, the latter characterized by the well-developed oblique longitudinal riblets. Suter described this species as Clathurella, but it would appear to be more in harmony with Mitromorpha.
I have much pleasure in associating with this species the name of Mr. Henry Suter, of Auckland.
[Footnote] * Proc. Mal. Soc. London, vol. ii., p. 30, pl. in., fig. 14.
[Footnote] † Trans. N Z. Inst, vol xxxi., p. 74, pl. iii., figs. 5, 5a.

Drillia lyallensis, n. sp. Fig. 7.
Shell small, fusiform, rather solid. Colour light or dark brownish-red, or somewhat purple (slightly beach-worn specimens). Whorls 6–6½, rather flattened, spire with a lightly turrited appearance, apex smooth; last whorl much longer than the spire, with eleven or twelve low, strong, rounded and slightly oblique ribs, rather wider than the interspaces, obsolete on the anterior half and usually on approaching the outer lip. Spiral sculpture consists of minute striæ, erased upon the ribs, a few at the anterior end stronger, and frequently several rough irregular ridges the remains of the old beaks. Sutures impressed, somewhat deep. Aperture narrow. Outer lip slightly thickened, the posterior sinus near to the suture and moderately deep. Columella lightly curved, not heavily callused; the canal short, broad, and slightly twisted. Length, 12 mm.; breadth, 4.56 mm.
Type in the Colonial Museum.
Hab. Lyall Bay (Mr. C. Freyberg).
This species is not uncommon in shell sand, but in the majority of specimens the finer sculpture is totally erased. Most examples show prominent scars, flexures repaired during the life of the animal. Of the five specimens before me, three are pierced by a carnivorous mollusc, and, strange as it may appear, each in exactly the same spot—a little above and to the left of the outer lip. In sculpture the species most resembles Clathurella sinclairi, Smith.
Cerithiopsis sarissa, n. sp. Figs. 8, 9.
Shell small, narrow, and tapering to a slender point, adorned with spiral and slightly weaker longitudinal riblets which form gemmules on the lines of intersection, base smooth. Colour light or dark brown, somewhat shining; the sutures of the anterior whorls, fourth spiral on the last, and base dark-purplish. Whorls 11–12, sutures deep. Apical whorls without sculpture; the succeeding with three spiral and numerous slightly oblique longitudinal riblets forming rows of gemmules. On the anterior whorls the sutures are margined with a minute beaded riblet; this gradually strengthens, and on the last whorl forms a fourth spiral, with the beading less marked than on the rows immediately above; beneath this a shallow groove, thence gently curved to the columella, and curving obliquely around the latter is a minute ridge which terminates at the canal. Of the three spiral rows of gemmules the two anterior are the largest, and about equal to the interspaces. The gemmules, of which on the last whorl there are seventeen to twenty per row, are somewhat oval, shining and variously coloured, light-brown, pale-chestnut,

and purple. Aperture ovate, with a well-marked chestnut band on the base, and the colour of the sculpture feebly produced. Outer lip sharp, sinuous. Columella short, nearly straight, and the beak slightly twisted to the left. Length, 6.25 mm.; breadth. 1.8 mm.
Type in the Colonial Museum.
Hab. Kawhia Harbour, immediately within the entrance, on rocks at low tide (R. M.); Whangaroa Harbour and Plimmerton, Cook Strait (Mr. Hamilton).
This beautiful little shell appears to be quite distinct from other New Zealand species. The few specimens obtained are, except that from Kawhia, dead and somewhat bleached; nevertheless the only example with the apex uninjured is from Whangaroa. The protoconch consist of about 3½ smooth rounded whorls, the apex globose, with the first half-turn oblique. The brephic period is not very distinctly marked off—it appears to consist of one volution; upon it a spiral riblet commences, which on the succeeding whorls forms the anterior spiral; thence follows somewhat strong longitudinal striæ, marking as it were short periods of growth, and from amidst these arise two posterior spirals, abruptly followed by the adult sculpture.
Leptothyra fluctuata, Hutton. Fig. 10.
Cyclostrema fluctuata, Hutton, Trans. N.Z. Inst., vol. xvi., p. 215 (1883, issued May, 1884). Leptothyra fluctuata, Hutton, Pilsbry, Man. Conch. (1), vol. x., p. 259, pl. 64, figs. 47, 48. L. fluctuata, Hutton, “Index Faunæ Novæ-Zealandiæ,” p. 81.
This species occurs in fair numbers. The irregular waved longitudinal bands of brown are well marked in some individuals; they extend across the base, but do not reach the umbilicus; the latter usually whitish, at times with broad white radiations. The spiral cingula vary from about twenty-five to thirty-five on the last whorl; the umbilicial area frequently cut up with strong irregular growth-periods. The outer lip descending, rather sharply in some, a thick callus uniting it to the columella. Height, 2.42 mm.; breadth, 3.11 mm.
The figure is derived from specimens collected by Mr. A. Hamilton at Whangaroa Harbour.
Captain Hutton kindly compared these northern forms with the types in the Canterbury Museum.
Hab. Foveaux Strait.
I have also to record the species from the Pliocene formation, Shakespeare Cliff, Wanganui—a single example, smaller than the recent forms.

Leptothyra crassicostata, n. sp. Fig. 11.
Shell small, solid, turbinate, with strong variable spiral sculpture. Colour whitish or light brown, occasionally with irregular markings of brown, most distinct on and below the periphery. Whorls 4, spire short, apex minute and without sculpture. The penultimate whorl with three to five, and the last with twelve to twenty, spiral ribs, four to nine of which are in front of the aperture. The ribs are very variable in size; there are five to eight strong riblets between the periphery and suture on the outer lip; in front of the aperture four or five. about equal to the breadth of the interspaces. On and immediately below the periphery frequently small and crowded, similar on the base, or there may be two or three more prominent intercalated with the smaller spirals, or the basal riblets generally stronger than those on the periphery. The growth-lines strong and irregular, producing here and there a lightly costate appearance, frequently pronounced in the umbilical area. Sutures impressed. Aperture subrotund. Outer lip sharp, descending, rather sharply in some. Columella lightly curved, somewhat produced and expanded anteriorly, a thick posterior callus spreading across to the outer lip. Umbilicus small and deep. Operculum circular, somewhat calcareous, of six or seven narrow folds, the nucleus central. Height, 2.49 mm.; breadth, 3.21 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
Distinguished from L. fluctuata, Hutton, by its much stronger sculpture.
Columbella huttoni, Suter, nom. mut. Fig. 12.
Lachesis sulcata, Hutton, Cat. Marine Moll. N.Z., p. 12 (1873); Cat. Tertiary Moll. and Echin. N.Z., p. 5; Manual N.Z. Moll., p. 45. Columbella huttoni, Suter, for sulcata, Hutton, preoccupied, “Index Faunæ Novæ-Zealandiæ,” p. 72 (1904).
The specimen figured is from the material collected by Mr. A. Hamilton at Whangaroa Harbour. The shell is small, robust, with well-marked spiral riblets somewhat wider than the grooves; last whorl longer than the spire, lightly carinate, more distinct in front of the aperture. Colour ash-grey, brown, or reddish-brown, sometimes variegated with white. Whorls rather flattened, 6 in number, including a 2- or 2½-whorled smooth reddish-brown protoconch. Sculpture: On the penultimate whorl five and on the last twelve to fourteen spiral riblets, five of which are in front of the aperture; a few of the riblets on the anterior end are minute, sometimes absent. These are crossed by irregular growth-lines, which give to the shell a slightly roughish appearance. Sutures impressed. Aperture oval, outer lip simple;

columella curved, lightly callused, and frequently a small denticle at the anterior end. Length, 6 mm., breadth, 2.52 mm.
The type locality is Stewart Island, the types preserved in the Canterbury Museum. They would appear to be somewhat larger than the northern forms.
Columbella transitans, n. sp. Fig. 13.
Shell small, fusiform, somewhat shining, with well-marked spiral grooves usually less than half the breadth of the interspaces. Colour light or dark brown, olive, or reddish-brown; some examples with a lighter band immediately below the sutures and produced around the periphery, in others this band is darker than the ground-colour. Whorls 6, rather flattened, the last longer than the spire. Protoconch of two and a half whorls, smooth and shining, light or dark olive, the apex usually lightest. Sutures impressed. Sculpture: The penultimate whorl with five or six and the last with seventeen to twenty-one spiral grooves, six or seven of which are in front of the aperture; occasionally a wider space surrounds the periphery, lightly cleft with a minute groove; longitudinally striate with irregular growth-periods. Aperture oval. Outer lip simple, uniformly curved. Columella lightly callused, the canal short and broad. Length, 5.11 mm.; breadth, 2.14 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
From C. huttoni, Suter, it may be distinguished by its smaller size, more numerous and much narrower spiral grooves. Its sculpture suggests a position intermediate with that species and the next form described.
Columbella paxillus, n. sp. Fig. 14.
Shell small, slender, smooth or with minute spiral grooves; the last whorl lightly angled, only noticeable in front of the aperture. Colour light - brown or reddish - brown or almost black, the sutures occasionally a little lighter. Whorls 5½–6, lightly convex, the last longer than the spire. Protoconch of about two whorls, not well defined, apex polished. Sutures impressed. Sculpture: A few small spirals on the anterior end of the last whorl; above the periphery and on the spire minute, irregular, or absent; occasionally a few inconspicuous longitudinal riblets on the upper whorls of the spire. Aperture short, oval. Outer lip simple. Columella curved and lightly callused, the anterior canal short and broad. Length, 5.04 mm.; breadth, 2.04 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton); Whangarei

Heads; also Takapuna, Manukau, Kawhia, and New Plymouth.
This little shell occurs in fair numbers. It is nearest to the preceding species; distinguished by the minute irregular spiral grooves, or their frequent absence except on the anterior end.
I may mention that Pyrene flexuosa, Hutton (dark form), is quite different in contour from the above: it has the form of C. choava, Reeve, and is probably only a colour variety of that extremely variable species.
Columbella saxatilis, n. sp. Fig. 15.
Shell small, ovate-elongate, sculptured with fine spiral and stronger longitudinal riblets. Colour light or dark brown, usually a pale band around the periphery. Whorls 6, rather flattened. Protoconch of two whorls, smooth and polished. Last whorl comparatively large. Sutures impressed. Sculpture: Longitudinal ribs sixteen to eighteen on a whorl, low and rounded, equal to or a little less than the breadth of the interspaces, becoming obsolete on the anterior end and usually on approaching the outer lip. Spirals undulating, small but well marked, wider than the grooves; on the penultimate six or seven, and on the last sixteen to seventeen, about seven of which are in front of the aperture; on crossing the longitudinals they have occasionally a somewhat granular appearance. Aperture somewhat narrow, longer than the spire. Outer lip rather flattened, a little thickened, and occasionally a few obscure denticles thereon. Columella frequently with two or three small folds or tubercles, the beak lightly twisted to the left; canal short and broad. Length, 5.86 mm.; breadth, 2.73 mm.
Type in the Colonial Museum.
Hab. Takapuna, from sand in rock-pockets (R. M.); Plimmerton, Cook Strait.
This species is perhaps nearest to C. pisaniopsis,* Hutton, a Pliocene form.
Crossea glabella, n. sp. Figs. 16, 17.
Shell minute, turbinate, smooth and rather solid. Colour white, somewhat shining. Whorls 3, without sculpture. Spire minute. Sutures distinct, not deep, with a narrow slightly concave depression below and adjoining. Base with broad depression, bounded by a prominent ridge; a second ridge arises from the narrow deep umbilicus; both sweep round to the anterior lip, where they form small well-marked expansions. Aperture subcircular. Inner lip reflected and callused, the latter
[Footnote] * Trans. N.Z. Inst., vol. xvii., p. 314; Macleay Mem. Vol., p. 45, pl. vi., fig. 17.

spreading across to the outer lip, which is sharp and simple. Height, 1.97 mm.; breadth, 1.94 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton). Found also at Stewart Island (A. H.).
This species, in its smoothness, solidity, and scanty umbilicus, is perhaps nearest to C. carinata,* Hedley, from New South Wales (75–100 fathoms). It appears to be the first record of the genus from New Zealand.
Odostomia impolita, Hutton. Fig. 18.
Rissoa impolita, Hutton, Cat. Marine Moll. N.Z., p. 29 (1873); Proc. Lin. Soc. N.S.W., vol. ix., p. 941. Barleeia impolita, Hutton, Manual N.Z. Moll., p. 81. B. impolita, Hutton, Suter, Proc. Mal. Soc. London, vol. iii., p. 7. Odontostoma impolita, Hutton “Index Faunæ Novæ-Zealandiæ,” p. 74.
The figure offered is derived from specimens in the collection of Mr. Henry Suter (hab. Foveaux Strait). These specimens have 5 whorls (one more than given in the description of the species), including the minute polished apex, the heterostrophe character of which is not very clear, due apparently to its being much buried in the succeeding whorl. The whorls are convex, the last much longer than the spire. Spiral striæ minute, rather more pronounced on the anterior half of the last whorl. Columella plait small, the lip anteriorly slightly effuse.
The types are in the Colonial Museum.
Hab. Stewart Island.
Odostomia proxima, n. sp. Fig. 19.
Shell small, elongated, rather fragile, smooth. White and slightly shining. Whorls 6, somewhat rounded, the last a little longer than the spire; the apical whorl minute, polished and heterostrophe. Sutures deeply impressed, not channelled. Sculptured with longitudinal irregular minute growth-periods, in places slightly pronounced, usually more marked on the spire. These are crossed by minute spiral striæ and scratches, very irregularly spaced, in some almost absent except for a few on the anterior half of the last whorl. Aperture small, ovate, oblique; columella slightly thickened and reflected, the plait not prominent; the lip anteriorly rather effuse. Base narrowly perforate. Length, 3.9 mm.; breadth, 1.8 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
Distinguished from O. impolita, Hutton, by its more rounded
[Footnote] * “Memoirs Australian Museum,” vol. iv., pt. 6 (Thetis), 1903, p. 345, fig. 71.

whorls and greater number, deeper sutures, irregular sculpture, and comparatively longer spire. It appears to be equally distinct from O. fasciata,* Hutton, a Pliocene form.
Odostomia vestalis, n. sp. Fig. 20.
Shell small, slender, fragile, smooth, and having a somewhat loosely coiled appearance. White and shining. Whorls 6, lightly convex; the last longer than the spire; apical whorl heterostrophe, slightly obliquely tilted. Sutures impressed, shallow and lightly submargined. Sculptured with minute irregular growth-lines, crossed by irregular microscopic striæ, the latter only indicated here and there. Aperture pyriform; columella lightly reflected, the plait small, rather deep within the aperture; anterior lip lightly effuse. Length, 4.31 mm.; breadth, 1.63 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
This form, of which there is but a single example, is nearest to the preceding species; distinguished by its more slender form, more shallow sutures, less rounded whorls, and more rapidly lengthening or loosely coiled appearance.
Odostomia (Pyrgulina) rugata, Hutton.
Odostomia (Parthenia) plicata, Hutton (not of Montford), Trans. N.Z. Inst., vol. xvii., p. 319, pl. xviii., fig. 17 (1884, issued May 1885). Odostomia rugata, Hutton (nom. mut.), Trans. N.Z. Inst., vol. xviii., p. 353 (1885, issued May 1886); “Macleay Memorial Volume,” p. 58, pl. vii., fig. 51.
This species has hitherto been recorded only as a Tertiary fossil. Pliocene: Wanganui and Petane. Found also in the Pareora system. It appears to be rare, both fossil and recent. Of the latter I have seen but three examples, and not in very good condition. I may add—the shell is white, somewhat shining and rather solid; the longitudinal riblets strong, slightly oblique advancing, seventeen to eighteen on the last whorl, feeble or obsolete on the anterior end; spirally adorned with minute dense striæ. Sutures impressed and lightly margined. Columella plait strong. Measurements (approx.): Length, 2.76 mm.; breadth, 1.28 mm. The fig. 51 in the “Macleay Memorial Volume” shows the riblets on the spire-whorls sloping forward, or advancing in a most marked manner. In the examples before me the slope is certainly less than half that shown in this figure.
Hab. Whangaroa Harbour, and Plimmerton, Cook Strait (Mr. A. Hamilton); Takapuna (Rev. W. H. Webster).
[Footnote] * Trans. N.Z. Inst., vol. xvii., p. 320; Macleay Mem Vol, p. 58, pl. vii.,

Vulpecula (Pusia) hedleyi, n. sp. Fig. 21.
Shell small, shortly fusiform, sculpture minute. Colour greyish-white, occasionally very faintly mottled with brown. Whorls 4½–5, rather flattened; spire short, conical, apex blunt; the last proportionally very large. Sutures lightly impressed. Sculptured with minute spiral grooves and with irregular longitudinal plications, the latter slightly more prominent at the sutures. Aperture oblique, narrow and much longer than the spire. Lip uniformly curved and slightly thickened. Columella white, lightly callused and armed with four plaits, the anterior two more oblique; opposite the latter the columella is lightly swollen. Anterior canal short and broad. Length, 5.42 mm.; breadth, 2.63 mm.
Type in the Colonial Museum.
Hab. Whangarei Heads, dredged in shallow water (Mr. C. Cooper, of Auckland).
Distinguished from other New Zealand species by the feeble longitudinal sculpture and short spire. I have much pleasure in naming this species after Mr. Charles Hedley, of the Australian Museum—a token of appreciation of his invariable kindly assistance.
Trophon (Kalydon) curta, n. sp. Fig. 22.
Shell small, ovate, rather solid, spirally and longitudinally ribbed, the latter strongest and forming prominent nodules on the lines of intersection. Colour whitish, occasionally a brown band on the base, rarely a few ill-defined scattered spots on the periphery. Whorls 6, lightly shouldered. Protoconch of two whorls, smooth except the last half-turn, upon which two small spirals arise. The sculpture consists of ten or eleven, rarely twelve, longitudinal ribs, narrower than the interspaces except when the latter number occurs, then equal or rather wider. Of the spirals there are two on the spire-whorls and six or seven on the last—occasionally three on the penultimate and eight on the last; the anterior spiral not infrequently prominent and the nodules sometimes obsolete. Aperture ovate. Outer lip slightly expanded, the margin occasionally feebly dentate; anterior canal short and somewhat curved. Length, 5.7 mm.; breadth, 2.59 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
Occurs in fair numbers, and appears to be the smallest of the genus recorded from New Zealand.
Rissoa leptalea, n. sp. Figs. 23, 24.
Shell minute, subrimate, slender, smooth, and rather fragile. Colour white, shining and semi-transparent. Whorls 5, having a somewhat loosely coiled, and those of the spire swollen, ap-

pearance. Apex minute; the last whorl longer than the spire. Without sculpture except the microscopic growth-lines. Sutures impressed and lightly margined. Aperture ovato-subcircular, peristome continuous; the columella and lip broadly expanded, the latter shortly descending. Length, 1.94 mm.; breadth, 66 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
Of this minute form there are only two examples. It is perhaps nearest to R. lubrica,* Suter, from Stewart Island.
Rissoa microstriata, n. sp. Fig. 25.
Shell minute, ovate-elongate, smooth and white. Whorls 5, somewhat swollen at the shoulders, thence gently flattened to the sutures; the latter impressed and lightly margined. Apex minute; last whorl longer than the spire. The sculpture consists of microscopic dense spiral striæ, broken up into variable lengths by the irregular lines of growth. Aperture obliquely broadly ovate; lip slightly thickened, a callus spreading across the body to the columella; the latter thickened and lightly reflected. Length, 2.11 mm.; breadth, 1.08 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
This little shell is perhaps near to Barleeia neozelanica,† Suter, while the sculpture, though microscopic, may be compared with Rissoa emarginata, Hutton, a Pliocene form.
Rissoa insculpta, n. sp. Fig. 28.
Shell small, elongate, slightly rimate, rather fragile. White. Whorls 5, lightly rounded; the last whorl much longer than the spire; the apex minute. Sutures impressed. Sculptured with minute spiral threadlets, absent on the apical whorls, about sixteen on the penultimate, and on the last about twenty-five, of which seventeen to eighteen are in front of the aperture. These are crossed by irregular growth-lines, here and there rather pronounced. Aperture broadly ovate, slightly oblique, and the peristome continuous. Columella regularly curved, lightly thickened and expanded, anteriorly a little produced; outer lip slightly thickened and projecting from the whorl above. Length, 2.56 mm.; breadth, 1.35 mm.
Type in the Colonial Museum.
Hab. Whangaroa Harbour (Mr. A. Hamilton).
This form is near to R. foveauxiana,‡ Suter. It differs in
[Footnote] * Proc. Mal. Soc. London, vol. iii., p. 5, fig. iii. (in text).
[Footnote] † Proc. Mal. Soc. London, vol. iii., p. 8, fig. v. (in text, on p. 5).
[Footnote] ‡ Proc. Mal. Soc. London, vol. iii., p. 5, fig. ii. (in text).

being more slender, more feebly rimate, sculpture much finer, sutures less deep, and the whorls less rounded. Perhaps the preceding species, R. microstriata, is next in the order of kinship. For comparison I herewith offer a figure of R. foveauxiana, derived from specimens identified by Mr. H. Suter; also a figure of R. fumata, Suter, derived from a typical specimen (hab. Lyall Bay).
List of Species from Whangaroa Harbour.
Class Gasteropoda.
Marinula filholi, Hutton.
Leuconopsis obsoleta, Hutton.
Cylichna striata, Hutton.
Drillia lævis, Hutton.
Surcula zealandica, E. A. Smith = cheesemani, Hutton (not novæzealandiæ, Reeve).
Mitromorpha substriata, Suter.
" subabnormis, Suter.
" suteri, Murdoch.
Mangilia flexicostata, Suter.
Clathurella sinclairi, E. A. Smith.
" epentroma, Murdoch.
" " var. whangaroaensis, Murdoch.
Daphnella lacunosa, Hutton.
Cancellaria trailli, Hutton.
Trophon duodecimus, Gray.
" plebejus, Hutton.
" curta, Murdoch.
" pumila, Suter = ambiguus Philippi var. pumila, Suter = T. bonneti, Cossmann.
Columbella choava, Reeve.
" huttoni, Suter.
" transitans, Murdoch.
" paxillus, Murdoch.
Taron dubius, Hutton.
Vulpecula rubiginosa, Hutton.
Leiostraca murdochi, Hedley.
Odostomia angasi, Tryon.
" proxima, Murdoch.
" vestalis, Murdoch.
Pyrgulina rugata, Hutton.
Turbonilla zealandica, Hutton.
Cæcum digitulum, Hedley.
Turritella carlottæ, Watson.
" kanieriensis, Harris.
Cerithiopsis terebelloides, von Martens.
" sarissa, Murdoch.

Potamopyrgus antipodum, Gray.
Rissoa huttoni, Suter.
" hamiltoni, Suter.
" subfusca, Hutton.
" " var. micronema, Suter.
" foveauxiana, Suter.
" cheilostoma, Ten.-Woods.
" suteri, Hedley.
" rosea, Hutton.
" neozelanica, Suter (Barleeia).
" leptalea, Murdoch.
" microstriata, Murdoch.
" insculpta, Murdoch.
Rissoina rugulosa, Hutton.
Eatoniella olivacea, Hutton.
Littorina mauritiana, Lamarck.
Risellopsis varia, Hutton.
" var. carinata, Kesteven.
Calyptræa maculata, Quoy and Gaimard.
" scutum, Lesson.
Trichotropis inornata, Hutton.
Crossea glabella, Murdoch.
Natica australis, Hutton.
Scala zelebori, Frauenfeld.
Janthina exigua, Lamarck.
Leptothyra fluctuata, Hutton.
" crassicostata, Murdoch.
Phasianella limbata, Hutton.
Trochus tiaratus, Quoy and Gaimard.
Cantharidus dilatatus, Sowerby.
" sanguineus, Gray.
Monilea egena, Gould.
Calliostoma punctulatum, Martyn.
Ethalia zelandica, Hombron and Jacquinot.
Incisura lytteltonensis, E. A. Smith (Scissurella).
Megatebennus moniliferus, Hutton.
Emarginula striatula, Quoy and Gaimard.
Acmæa pileopsis, Quoy and Gaimard.
Class Pelecypoda.
Barnea similis, Gray.
Saxicava arctica, Lin.
Cardium pulchellum, Gray.
Chione crassa, Quoy and Gaimard.
Tapes fabagella, Deshayes.
Cyamiomactra problematica, Bernard.
Kellya sanguineum, Hutton.

Kellya parva, Deshayes.
Lasea miliaris, Philippi.
Diplodonta zelandica, Gray.
Divaricella cumingi, Adams and Angas.
Cryptodon flexuosum, Montague.
Cuna delta, Tate and May.
Verticipronus mytilus, Hedley.
Cardita aviculina, Lamarck.
Venericardia zealandica, Potiez and Michaud.
" corbis, Philippi.
Lima bullata, Born.
Pecten zelandiœ, Gray.
Philobrya meleagrina, Bernard.
" costata, Bernard.
Barbatia decussata, Sowerby.
Glycymeris striatularis, Lamarck.
Nucula nitidula, A. Adams.
Leda concinna, A. Adams.
Malletia australis, Quoy and Gaimard.
Class Brachiopoda.
Terebratella rubicunda, Sowerby.
Rhynchonella nigricans, Sowerby.
Explanation of Plates VII. and VIII.
Plate VII.
| Figs. 1, 2. |
Cylichna striata, Hutton. |
| Figs. 3, 4. |
Clathurella epentroma, n. sp. |
| Fig. 5. |
Clathurella epentroma, var. whangaroaensis, n. var. |
| Fig. 6. |
Mitromorpha suteri, n. sp. |
| Fig. 7. |
Drillia lyallensis, n. sp. |
| Figs. 8, 9. |
Cerithiopsis sarissa, n. sp. |
| Fig. 10. |
Leptothyra fluctuata, Hutton. |
| Fig. 11. |
Leptothyra crassicostata, n. sp. |
| Fig. 12. |
Columbella huttoni, Suter. |
| Fig. 13. |
Columbella transitans, n. sp. |
| Fig. 14. |
Columbella paxillus, n. sp. |
Plate VIII.
| Fig. 15. |
Columbella saxatilis, n. sp. |
| Figs. 16, 17. |
Crossea glabella, n. sp. |
| Fig. 18. |
Odostomia impolita, Hutton. |
| Fig. 19. |
Odostomia proxima, n. sp. |
| Fig. 20. |
Odostomia vestalis, n. sp. |
| Fig. 21. |
Vulpecula hedleyi, n. sp. |
| Fig. 22. |
Trophon curta, n. sp. |
| Figs. 23, 24. |
Rissoa leptalea, n sp. |
| Fig. 25. |
Rissoa microstiata, n. sp. |
| Fig. 26. |
Rissoa fumata, Suter. |
| Fig. 27. |
Rissoa foveauxiana, Suter. |
| Fig. 28. |
Rissoa insculpta, n. sp. |

Art. XIII.—Report on the Mollusca collected by Messrs. Keith Lucas and G. L. Hodgkin in Six Lakes of New Zealand.
[Read before the Wellington Philosophical Society, 7th September, 1904.]
During the year 1902 Messrs. Keith Lucas and G. L. Hodgkin, of Cambridge, made a collection of the fauna of a typical series of lakes in New Zealand, most of the Mollusca being obtained by dredging. Before reaching New Zealand Mr. Lucas had promised Mr. Charles Hedley, conchologist of the Australian Museum, Sydney, to hand him over the Mollusca which would be obtained from our lakes for study, but later on, with Mr. Hedley's consent, the whole of the material was handed over to me. My very best thanks are due to Messrs. Lucas and Hodgkin for allowing me to study the interesting collection, and to my friend Mr. Hedley for his most readily given consent.
Mr. Lucas has published in the Geographical Journal for May and June, 1904, a highly interesting and beautifully illustrated report on the “Bathymetrical Survey of the Lakes of New Zealand,” and Professor W. B. Benham, of Dunedin, has published* the result of his study of the aquatic Oligochæta collected by Messrs. Keith Lucas and Hodgkin.
The lakes from which Mollusca were handed over to me are the following: North Island—Waikare, Rotoiti, Taupo, Waikaremoana; South Island—Wakatipu, Manapouri.
With regard to the deep-water Mollusca of our lakes nothing at all was known, all the collecting hitherto done having been confined to shallow water, and mostly, of course, near the shore. To show how scanty our knowledge of the molluscan fauna of the above-named lakes has been, it may be of interest to mention that the following species have been recorded: Lake Rotoiti—Sphœrium novæ-zelandiœ, Desh. (as lenticula, Dk.), Potamopyrgus badia, Gould (as fischeri, Dk.), both collected by Hochstetter, and Melanopsis trifasciata, Gray. Lake Taupo—Diplodon menziesi, Gray (Dieffenbach), and its subsp. hochstetteri, Dk. (Hochstetter). Lake Waikaremoana—Diplodon menziesi, Gray (as waikarense, Colenso). Lake Wakatipu—Planorbis corinna, Gray, and Isidora antipodea, Sow., both collected by Captain F. W. Hutton.
I propose to describe first, in systematical order, all the molluscs brought together, then to give a synopsis of the molluscan fauna of each lake, and finally tabulate the bathymetrical distribution of the species.
[Footnote] * P.Z.S., 1903, vol. ii., pp. 202–232, pls. xxiv.-xxvi.

Fam. UnionidÆ.
Genus Diplodon, Spix (1827).
Shell elliptical, rounded, elongated or trapezoidal, with rather low beaks which are more or less distinctly radially sculptured, the ridges usually curved and approaching below, with a low or scarcely developed posterior ridge; surface slightly concentrically sculptured, sometimes broken into fine nodules or corrugations; epidermis dull, rayless; hinge with two compressed pseudocardinals in the right valve, and one slender lateral, and two compressed pseudocardinals in the left valve, one in front of the other, and two laterals; nacre bluish to white, dull, often blotched; beak-cavities shallow; dorsal scars numerous, forming a row in the beak-cavity parallel with the hinge line.
Animal with the marsupium occupying nearly the whole length of the inner branchiæ, a few ovules sometimes being found in the outer gills; branchiæ rather large, angular at base, inner much the larger, united their whole length to the abdominal sac; palpi scarcely projecting posteriorly; mantle very thin, thickened on the edges; branchial opening papillose, separated from the smooth anal opening by a strong bridge; supra-anal opening not closed below. (Simpson.)
Type: D. ellipticum, Spix.
Subgenus Hyridella, Swainson (1840).
Beaks rather low, sculpture consisting of curved generally nodulous ridges, which approach below, but usually have a smooth area of shell between them; surface sulcate or sometimes corrugated and nodulous; epidermis rayless; teeth rather delicate, compressed, often somewhat rudimentary.
Animal having the embryos occupying the inner gills for the most part, which are united for their entire length to the abdominal sac; outer gills pointed below in the middle; palpi triangular; branchial opening papillose; anal opening smooth, not separated from the supra-anal opening. (Simpson.)
Type: Unio australis, Lamarck.
Diplodon menziesi, Gray (1843).
Unio menziesi, Gray, in Dieffenbach's “New Zealand,” vol. ii., 1843, p. 257. Unio aucklandica, Gray, l.c., p. 257. Unio waikarense, Colenso, Tasman. Journ. Nat. Sci., vol. ii., 1845, p. 250.
I am following C. Simpson in considering Unio aucklandica as a synonym only. On examining a rather large series of specimens from over twenty localities I tried to uphold it at least as

a subspecies, but I had to give it up, as I had numerous specimens before me which could be either assigned to menziesi or to aucklandica. There is no doubt that when the extreme forms only are compared one would feel inclined to take them for distinct species, but so it is with many other species, as for instance with Helcioniscus tramosericus. However, it is convenient to refer to aucklandica as a form of menziesi which is but little winged, and having the dorsal and ventral margins subparallel.
Unio waikarense will be dealt with further on when describing the mussels from Lake Waikaremoana.
(1.) Lake Taupo (Stat. 9—From dredgings up to 100 ft.).—There are eight specimens, representing quite young to half-grown forms, only one being highly winged. All are distinctly radiately striate, and some of the youngest specimens show the typical beak-sculpture beautifully. It is represented by the accompanying diagram (fig. 1). One of the larger specimens is very distinctly sculptured with elongate nodules on the lower half down to the ventral margin in the region below the beaks. The interior and hinge are the same as in specimens of subspecies hochstetteri, to be described further on. The largest specimen measures—Length, 42 mm.; height, 28 mm.; diam., 13 mm.
(2.) Lake Taupo (Stat. 9F—From dredgings up to 100 ft.).—The eight specimens have the same appearance as those of the last station; all of them have the outline of aucklandica, are finely radiately striate, and one clean olive-coloured specimen also shows nodulous ornamentation. Five quite young specimens have the beaks already so much eroded that no trace of the beak-sculpture is left. A few specimens have a light ferrugineous coating. The largest specimen shows—Length, 43 mm.; height, 28 mm.; diam., 13 mm.
(3.) Lake Waikaremoana (Stat. 14—Dredged in 50 ft.).—Compared with the type of Unio waikarense, Colenso, said to have been obtained in this lake, the four specimens collected at this station are much smaller, very little winged posteriorly, the dorsal margin subparallel to the ventral, darker in colour, and more solid; they are not concentrically sulcated, but only striated, and the marks of rest are much less distinct. All of them are finely radiately striated, a character always to be found in menziesi. The pseudo cardinals are typical, the upper lateral tooth in the left valve is much lower than the other, and crenate

in all specimens. Inside bluish-white, pearly, blotched with yellowish, especially in the umbonal cavity.
(4.) Lake Waikaremoana (Stat. 37—Dredged in 15 ft.).—Four specimens of different size, showing all the characters of those from Stat. 14.
In some specimens in my collection, kindly sent to me by Mr. Elsdon Best, and collected in Lake Waikaremoana, the radiate sculpture is distinctly nodulous, sometimes V-shaped, thus approaching D. websteri, Simpson. Typical specimens of aucklandica from creeks near Auckland show the same sculpture to a most marked degree, and I consider D. websteri as a D. menziesi in which the nodulous sculpture is developed to the highest degree.
I have seen, thanks to Mr. E. Best's great kindness, a large number of Diplodon from Lake Waikaremoana, but not one approaching Colenso's type of waikarense, which is a large, thin, yellowish-olive-coloured shell, having more the appearance of an Anodonta. The Waikaremoana specimens are all much smaller, thicker, darker in colour, and less winged. Mr. A. Hamilton, Director of the Colonial Museum, to whom I spoke about it, and who has visited the locality, suggests that Colenso did not get his specimens from Lake Waikaremoana itself, but from some small lake or lagoon in the vicinity. This seems to be correct, as I obtained, again through the unremitting kindness of Mr. E. Best, a number of specimens from a lagoon near Ruatoki, Tuhoeland, and these are typical waikarense, Colenso; one of them showing the same outline and the same dimensions as the cotype in the Canterbury Museum.
An error in Colenso's diagnosis of Unio waikarense (Trans. N.Z. Inst., xiv., 169) wants correcting. He says, “Posterior slope keeled.” This, however, is not correct: the shell is not keeled. How misleading such an incorrect statement is may be guessed from Simpson's remark on the species in his “Synopsis of the Naiades,” p. 890 (footnote): “Suter thinks this is a variety of menziesi, but Colenso states that the posterior slope is keeled. If this is so it must be quite different from the species.” There is also an error in the quotation in Simpson's work, as he gives the year 1841 as the date of the publication of the Tasman. Journ. Nat. Sci. vol. ii., whereas it is 1845. Colenso says that he discovered the shell in 1841, but his description was published four years later.
I have compared typical specimens of waikarense with many specimens of menziesi from about twenty different localities, and I am unable to separate the two. In all essential characters the two agree, and there remains nothing but to make Colenso's species a synonym of D. menziesi, Gray.

Diplodon menziesi, Gray, subsp. hochstetteri, Dunker (1862).
Unio hochstetteri, Dunker, Malak. Blätter, vol. viii., 1862, p. 153.
The type was collected by Hochstetter in the Waikato River.
(1.) Lake Taupo (Stat. 46—From dredgings in 10–30 ft.).—Six specimens, blackish-brown, much corroded round the beaks, most of them more produced and rounded posteriorly than the type, but some have the posterior end distinctly truncated and more or less biangular. In correspondence with Mr. C. T. Simpson I expressed the opinion that hochstetteri is a pathological subspecies,* having seen the same form amongst specimens of rugatus, Hutton, from the Kopuaranga River,† and also amongst menziesi from the River Avon. The young shells are invariably typical D. menziesi, but on growing larger the deformity constituting hochstetteri becomes more and more apparent. Some specimens are more affected, others less, thus producing a strongly truncated biangular posterior margin, or it remains only flatly rounded. At the posterior end the periostracum is produced in thick, foliated layers, and the inner margin is considerably thickened by pearly substance, forming large rugosities, and very often pearls adhering to the shell are met with. Loose small pearls of irregular form are only exceptionally found. In my opinion the cause of this is most likely some parasitic creature, as is the case in most of the pearl-producing bivalves. The outer exposed layers round the beaks are smooth, light-brown, waxy. The concentric striation is rather coarse, the marks of rest distinct and elevated. The inside is but little iridescent, except along the ventral margin, outside the mantle impression, grey to light-brown, sometimes blotched with brown; there are more or less considerable rugosities beyond those at the posterior margin. The dorsal scars in the umbonal cavity are small and deep, and to the number of two to four. The anterior adductor impressions are irregular in shape, and much deeper than the posterior ones, which are oval and shallow. Right valve with two pseudocardinals, the upper anterior forming a small lamella, the posterior being strong, compressed, high, triangular, and slightly corrugated. The single lateral tooth is regularly slightly curved and somewhat crenate at its posterior end, which is abruptly descending. Left valve with two pseudocardinals arising from a common base, both blunt, the anterior tooth larger and but slightly crenate. The two laterals are also curved, the lower of them is strongly lamellated and denticulated at its posterior end, which slopes down very gradually.
[Footnote] * C. T. Simpson, “Synopsis of the Naiades,” p. 889 (footnote)
[Footnote] † Trans. N.Z. Inst., xxiv., 275.

(2.) Lake Rotoiti (Stat. 8—Obtained by dredging in 12 ft.).—Two specimens of medium size, strongly concentrically ridged, umbones much eroded, posterior end subtruncated, not yet distinctly biangular. They represent the intermediate stage between the young menziesi and the full-grown hochstetteri. Interior olive-bluish, pearly, a row of small dorsal scars in the umbonal cavity, parallel to the hinge line. Muscular scars strongly impressed. Right valve with two very unequal pseudo-cardinals, the anterior small, lamellar, the posterior compressed, with a broad posterior base, crenulated, trifid; the lateral tooth slightly curved, narrow, high, truncate and corrugate posteriorly. Left valve with the two pseudocardinals coalescent, separated only by a groove, the anterior strong, triangular, rugose. The two laterals with crenate edges, obliquely truncated behind. The dimensions of the two specimens are—Length, 50 mm.; height, 34 mm.; diam., 16 mm.; and, length, 41 mm.; height, 29 mm.; diam., 12.5 mm.
Diplodon menziesi, Gray, subsp. rugata, Hutton (1883).
Unio rugatus, Hutton, N.Z. Journ. Sci., vol. i., 1883, p. 478.
Lake Waikare (Stat. 41—From dredgings in all parts of the lake; most common on sand and on stony shore).—Six specimens, two of which are quite young. The latter are winged posteriorly, the beaks already eroded, and there are three to six nodulous ridges descending in front of the umbo, nearly parallel to the anterior margin; they are concentrically finely thread-striated; colour olive-brown; the inside is bluish-pearly, yellow under the beaks. The larger specimens are subventricose and thin, less winged, the dorsal margin nearly parallel to the length-axis; the beaks are corroded, the anterior margin rounded, sometimes slightly truncate, the posterior margin produced and the ventral margin broadly convex. The concentric striae are irregular, rugose posteriorly, fine and more regular at the anterior end; the marks of rest are fairly distinct. Most of the specimens are partly covered with a dark-brown ferrugineous coating. Interior bluish, yellowish, or purple pearly, sometimes strongly blotched with olive. Muscular scars shallow. Right valve with two compressed triangular pseudocardinals close together, the anterior tooth smaller, lamellar and smooth, the posterior stronger, broader, and somewhat rugose. The lateral tooth is slender, curved, rugose on the edge of the posterior part. Left valve with two elongated, rounded, rugose pseudocardinals which sometimes coalesce, when only a slight notch indicates the original two teeth. The two laterals are long, slender, sinuate, and distinctly pectinate at the posterior edges. There are always a few small rather deep dorsal scars in the umbonal cavity.

Diplodon menziesi, Gray, n. subsp. lucasi.
Lake Manapouri (Stat. 35—Dredging in 60 ft.).—Three specimens; one of them may be taken as adult. Shell (figs. 2 and 3) oblong-ovate, very much compressed, thin and fragile, inequilateral, beaks low, eroded; surface with close strongly pronounced rest-marks and between them a few lines of growth, all close together and foliated at the anterior end. In the adult specimen the middle part has distinct radiate nodulous sculpture, partly V-shaped, but no such ornamentation is to be found on the young specimens. The straight dorsal margin is subparallel to the ventral margin, which is slightly sinuate; the anterior margin is angularly rounded, the posterior obliquely truncated and slightly produced. Nearly the whole of the shell is covered with a thin ferrugineous coating; the epidemis is olive-green, waxy. The ligament is small, not much raised. In the right valve the two pseudocardinals are compressed, small; the upper anterior tooth is a small, smooth lamella, the lower tooth is more elevated, conoidal, and strongly crenate; the lateral tooth is almost straight, thin and rugose at its posterior portion. In the left valve there is a rather long compressed lower anterior rugose pseudocardinal, the upper tooth is quite rudimentary; the upper lateral tooth is a little higher and more rugose posteriorly than the other. Interior bluish-white, pearly, a little blotched with olive in the umbonal cavity, where there are rather large and deep dorsal scars. The adductor-muscle scars are shallow. The young specimens are slightly winged.
This subspecies is nearest to the typical aucklandica, but is distinguished from it by its exceptionally compressed form, the thinness of the shell, the strongly marked and close concentric lines, the more tapering posterior margin, and the feebly developed pseudocardinals. The radiate nodulous sculpture is found in many specimens of menziesi and its subspecies. Adult specimen—Length, 45 mm.; height, 24 mm.; diam., 8 mm.
Type in my collection.
I have much pleasure in naming the subspecies in honour of its discoverer, Mr. Keith Lucas.

Diplodon lessoni, Küster (1856).
Unio lessoni, Küster, Conch. Cat., 1856, p. 135, pl. xxxvi., fig. 4. Type from New South Wales.
Lake Wakatipu (Stat. 37—Dredged in 10 ft. to 30 ft.).—Nine specimens were obtained. Shell (fig. 4) oblong, obliquely truncated behind, compressed, inequilateral, beaks strongly eroded in all specimens, no trace of sculpture left; surface with distant flatly elevated rest-marks, which, together with the intervening space, are covered with very fine thread-like concentric lines; towards the base and posterior margin the growth-lines are more distinct and slightly foliated. There is no trace of radial sculpture. The epidermis is olive to dark-brown, dull. Interior bluish nacre, with yellowish patches under the beaks, where there are several deep small dorsal scars. Right valve: The pseudocardinals are compressed, the anterior upper tooth is small, grooved, the posterior tooth much larger and crenate; the lateral tooth is slightly arched and rugose posteriorly. Left valve: Anterior pseudo-cardinal compressed, tongue-shaped, slightly rugose, the posterior subtriangular, deeply denticulate; the laterals are of nearly equal height and crenate posteriorly.
The largest specimen was selected for the diagram and description, and its dimensions are—Length, 51 mm.; height, 30 mm.; diam., 14½ mm.
I submitted specimens to Dr. W. H. Dall, Hon. Curator of Mollusks, U.S. Nat. Museum, for examination, and he very kindly informed me that according to the material in the museum they are D. lessoni, Küster. I have not seen any Australian specimens, nor the description and figure published by Küster, but I do not hesitate to accept Dr. Dall's view, although some may think it hazardous to refer our shell to a New South Wales species. I have not seen this species from anywhere else in New Zealand.
Dr. Von Jhering once suggested that a number of measurements should be taken to ascertain the range of variability in species of the family Unionidœ. I have measured a great number of specimens from New Zealand some years ago, and may publish the results later on. I have done the same for a number of the Unionidœ collected by Messrs. Lucas and Hodgkin, and

the result is contained in the following table. [The ciphers i. to xi. in the table indicate—i. Length of shell. ii. Greatest height. iii. Index of height (length = 100, height in % of it). iv. Distance of greatest height from anterior end. v. Index of position of greatest height (length = 100, distance of greatest height in % of it). vi. Diameter. vii. Index of diameter (length = 100, diameter in % of it). viii. Length of hinge line. ix. Distance of beak from anterior end of hinge line. x. Umbonal index (length of hinge line = 100, distance of beak from anterior end of hinge line in % of it). xi. Cardinal index (length of shell = 100, length of hinge line in % of it).]
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
| — | i. | ii. | iii. | iv. | v. | vi. | vii. | viii. | ix. | x. | xi. |
|---|---|---|---|---|---|---|---|---|---|---|---|
| D. menziesi. | |||||||||||
| Taupo, 9 | 32 | 19 | 59 | 9 | 28 | ||||||
| " 9F | 37 | 23 | 62 | 10 | 27 | ||||||
| Waikaremoana, 14 | 44 | 28 | 64 | 23 | 52 | 13 | 29 | 32 | 9 | 28 | 72 |
| " 14 | 41 | 23 | 56 | 25 | 61 | 10 | 24 | 25 | 6 | 24 | 61 |
| " 14 | 39 | 22 | 56 | 21 | 53 | 10 | 27 | 24 | 6 | 25 | 61 |
| Mean of the three above | 41 | 24 | 58 | 23 | 55 | 11 | 27 | 27 | 7 | 25 | 64 |
| Waikaremoana, 37 | 55 | 32 | 58 | 30 | 54 | 18 | 32 | 38 | 10 | 26 | 69 |
| " 37 | 60 | 33 | 55 | 28 | 47 | 17 | 28 | 40 | 10 | 25 | 67 |
| " 37 | 61 | 36 | 59 | 31 | 50 | 19 | 31 | 40 | 10 | 25 | 66 |
| " 37 | 62 | 33 | 53 | 31 | 50 | 19 | 30 | 40 | 10 | 25 | 64 |
| Mean of the four above | 59 | 33 | 56 | 30 | 50 | 18 | 30 | 39 | 10 | 25 | 64 |
| Cotype of waikarense | 79 | 48 | 61 | 40 | 51 | 25 | 31 | 47 | 10 | 21 | 59 |
| D. hochstetteri. | |||||||||||
| Taupo, 46 | 48 | 31 | 65 | 24 | 50 | 16 | 33 | 36 | 8½ | 24 | 75 |
| " 46 | 52 | 36 | 70 | 30 | 58 | 18 | 35 | 35 | 8 | 23 | 67 |
| " 46 | 54 | 36 | 67 | 30 | 56 | 19 | 35 | 40 | 10 | 25 | 74 |
| " 46 | 64 | 40 | 63 | 33 | 52 | 22 | 34 | 48 | 11 | 23 | 75 |
| Mean of the four above | 55 | 38 | 66 | 29 | 54 | 19 | 34 | 40 | 9½ | 24 | 72 |
| From Lake Rotorua, typical | 61 | 44 | 72 | 32 | 52 | 24 | 39 | 42 | 10 | 24 | 69 |
| D. rugata. | |||||||||||
| Waikare, 41 | 47 | 28 | 59 | 22 | 47 | 14 | 29 | 31 | 8 | 26 | 66 |
| " 41 | 46 | 29 | 63 | 26 | 56 | 15 | 32 | 30 | 8 | 26 | 65 |
| " 41 | 44 | 26 | 59 | 20 | 45 | 13 | 29 | 25 | 6½ | 26 | 57 |
| " 41 | 41 | 25 | 61 | 21 | 51 | 13 | 31 | 29 | 8 | 27 | 70 |
| Mean of the four above | 44 | 27 | 60 | 22 | 50 | 14 | 30 | 29 | 8 | 26 | 64 |
| From Lake Pearson, cotype | 52 | 33 | 63 | 26 | 50 | 17 | 32 | 35 | 9 | 25 | 67 |
| D. lucasi. | |||||||||||
| Manapouri, 35 | 45 | 24 | 53 | 19 | 42 | 8 | 18 | 28 | 8 | 28 | 62 |
| " 35 | 34 | 21 | 61 | 16 | 47 | 8 | 23 | 22 | 7 | 31 | 64 |
| " 35 | 28 | 17½ | 62 | 15 | 53 | 6½ | 23 | 18 | 5 | 28 | 64 |
| D. lessoni. | |||||||||||
| Wakatipu, 37 | 51 | 30 | 58 | 23 | 45 | 14½ | 28 | 35 | 9 | 26 | 68 |
| " 37 | 44 | 24 | 58 | 22 | 53 | 11½ | 28 | 26 | 7½ | 28 | 63 |

Fam. Sphæriidæ.
Cardinal teeth not exceeding two in each valve, and exhibiting a cessation of development at an early stage. (Dall.)
The New Zealand species of Sphærium and Corneocyclas are not easily separated with regard to their generic position, as both are about to the same extent inequilateral, though in Corneocyclas the anterior part is mostly more attenuated and the nepionic shell sometimes more conspicuous than in Sphærium. The cardinal teeth are very variable, and it is a tedious job to ascertain the genus from these alone, but they assist in the general diagnosis. To be absolutely certain living specimens should be examined, Sphærium having two siphonal tubes, Corneocyclas only one; but with the specimens to be described that was out of the question. Fortunately I have specimens of both genera from New Zealand in my collection, which I examined when alive, and these were of great help to me for the study of the various specimens collected by Messrs. Lucas and Hodgkin.
Much useful information was obtained from the chapter on the family Sphæriidæ in the classical work of Dr. W. H. Dall, “Tertiary Fauna of Florida.”
The indices I calculated for the Sphæriidæ are part of those used for the Unionidœ: iii. = index of height; vii. = index of diameter; x. = umbonal index.
Genus Sphærium, Scopoli (1777).
Subgenus Sphærium, Scopoli, s.s.
Type: S. corneum, Linné.
The nepionic shell passing into the adult without any distinct demarcation; the anterior end shorter; the ligament subinternal; the two right cardinals widely divergent and coalescent at their adjacent or upper ends, thus apparently forming but one tooth, but which if it had continued in development would have separated into two; the widening of the ventral angle causes the A-shape to disappear; the nepionic shell (and consequently the beaks) is finely concentrically striate or even nearly smooth and rather convex. (Dall.)
Sphærium novæ-zelandiœ, Deshayes (1853).
Cat. Conchif. Brit. Museum, p. 272; P.Z.S., 1854, p. 342.
(1.) Lake Rotoiti (Stat. 15—From muddy bottom and weeds in 6 ft.).—Three specimens of nearly equal size, suborbicular, subequilateral; colour grey, near the ventral margin yellowish; beaks covered with a ferrugineous incrustation. Nepionic shell

hardly to be distinguished from the later growth, obtuse; anterior side of shell slightly produced, rounded, posterior side broadly convex; the whole of the valves concentrically finely striate. Ligament inset. Left valve with a straight lamellar upper cardinal, and below and slightly in front of it a curved stouter tooth. In the right valve the two cardinals are united, forming an oblong squarish tooth with three denticulations on the lower margin. Laterals smooth. Fig. 5 illustrates the cardinal teeth of the two valves. Length, 5 mm.; height, 4¼ mm.; diam., 2¾ mm. Umbo 3 mm. from anterior end. The indices are—iii. = 85; vii. = 55; x. = 60.
This is no doubt the same as S. lenticula, Dunker, as Hochstetter obtained the specimens from the lakes Rotoiti and Taupo, and I consider it as a synonym of S. novæ-ze-landiæ.
(2.) Lake Taupo (Stat. 4—Dredged with weeds in 20 ft., and from bottom of coarse pumice in 100 ft.).—Four specimens of straw colour, the nepionic shell rather distinct, but otherwise not differing from the typical form. Right valve with two minute cardinals, the anterior oblong, the posterior short, triangularly oval, elevated. Left valve with two minute teeth, broadly rounded behind, pointed at the anterior end, one in front of the other. Fig. 6 shows the form and position of the cardinals. Length, 4½ mm.; height, 4 mm.; diam., 2½ mm. Umbo 2¾ mm. from anterior end. Indices—iii. = 84–89; vii. = 56; x. = 61.
(3.) Lake Taupo (Stat. 34—With weeds from 20 ft. to 80 ft.).—Six small specimens, a little more oval and compressed, but otherwise the same as the examples from Stat. 4. Length, 4 mm.; height, 3 mm.; diam., 1¾ mm. These are the dimensions of Dunker's S. lenticula. Indices—iii. = 75; vii.—44.
(4.) Lake Waikaremoana (Stat. 33—Dredged in 10 ft. to 20 ft.).—There are twenty-three small to minute specimens. The larger examples are yellowish-grey, the small ones straw-colour. In the right valve there are two lamellar cardinal teeth meeting at an obtuse angle; in the left valve there is one crescent-shaped

rounded cardinal, and a strong bifid crenulated cardinal below and slightly in front of it. Fig. 7 illustrates the cardinals of the valves. The lateral teeth are smooth; the ligament inset. Dimensions of two specimens—Length, 4¼ mm.; height, 3½ mm.; diam., 2 mm.; beak 2½ mm. from anterior margin: length, 4 mm.; height, 3½ mm.; diam., 2 mm.; beak 2¼ mm. from anterior margin. The mean of the indices is—iii. = 85; vii. = 49; x. = 58.
Genus Corneocyclas, Férussac (1818).
(= Pisidium, Pfeiffer, &c.)
Subgenus Corneocyclas.
Nepionic shell convex, concentrically striated; hinge with two separate cardinals in the left, and a single compound, usually arcuate, cardinal in the right valve. (Dall.)
Sec. Corneocyclas, s.s.
Type: Tellina pusilla, Gmelin.
Nepionic valves passing into the mature disc without any strong demarcation; the anterior cardinal and lateral adjacent and retaining traces of their original connection; ligament internal. (Dall.)
Corneocyclas novozeelandica, Prime (1862).
P.Z.S., 1862, p. 3.
(1.) Lake Waikare (Stat. 29—From weeds in 1½ ft. to 2 ft.).—Nine specimens of different size; colour greyish-yellow, shining, inequilateral, with fine concentric lines, which are crossed by close microscopic radiate striæ. Nepionic shell convex, concentrically striated, well delimited, but passing without any change into the disc of the adult valve. Ligament inset. Right valve with the two cardinals remaining united, crescent-shaped; four laterals. Left valve with two cardinals, the posterior bifid, the anterior triangular; two laterals. The accompanying fig. 8 shows the characters of the hinge.

Four specimens were measured—Length, 2¾–5 mm.; height, 2½–4 mm.; diam., 1¼–2½ mm.: and the mean indices are—iii. = 83; vii. = 48; x. = 59.
These specimens are very much like those I collected in the River Avon, Christchurch.
(2.) Lake Waikare (Stat. 29F—From weeds in 1½ ft. to 2 ft.).—Four specimens, dead when collected. They do not differ much from those of Stat. 29, except being slightly more globular. The dimensions of three specimens measured are—Length, 5 mm.; height, 4½ mm.; diam., 3¼ mm.; umbo 3 mm. from anterior margin: length, 4¼ mm.; height, 3½ mm.; diam., 2¼ mm. umbo 2½ mm. from anterior margin: length, 3 mm.; height, 2½ mm.; diam., 1½ mm.; umbo 1¾ mm. from anterior margin. The mean indices are—iii. = 85; vii. = 56; x. = 58.
(3.) Lake Wakatipu (Stat. 17—Dredged with weeds in 20 ft. to 100 ft.).—One minute specimen, rounded, yellowish-white, subequilateral. Length, 2¼ mm.; height, 2 mm.
This small specimen has the aspect of a Sphærium, but the nepionic shell is so distinct that I prefer placing it in Corneocyclas for the present. I tried to separate the valves, but could not do it without running the risk of breaking the valves. Without ample material, and especially examining the living animal, it is impossible to be quite certain about its generic position.
(4.) Lake Wakatipu (Stat. 34—Dredged in 10 ft. to 30 ft.).—One adult and four young specimens. The adult is light-brown, the nepionic shell distinctly limited; the young examples are grey to yellowish, more inequilateral and tapering in front. The dimensions of two specimens are—Length, 5¼ mm.; height, 4½ mm.; diam., 2½ mm.; umbo 2¾ mm. from anterior end: length, 3½ mm.; height, 1¾ mm.; diam., 1½ mm.; umbo 2 mm. from anterior end: and the mean indices are—iii. = 83; vii. = 46; x. = 55.
There is one specimen, covered with a black coating, which is suborbicular, much compressed, inequilateral, with inconspicuous beaks, the anterior side broadly rounded. The dimensions are—Length, 4 mm.; height, 3¼ mm.; diam., 1½ mm.; umbo 2½ mm. from anterior end: and the indices are—iii. = 81; vii. = 38; x. = 62. I have seen similar abnormities amongst specimens I collected in the River Avon.
(5.) Lake Manapouri (Stat. 13—Dredged in shallow water near the shore).—Three young specimens, which are at once recognised as being Corneocyclas, being more inequilateral, compressed and attenuated anteriorly than adult specimens. Had I not seen the same forms from other localities, collected together with adult examples, I might have been inclined to consider

them a distinct species. With few exceptions the young shells of C. novozeelandica show the characters just indicated. The colour is light-grey, and the nepionic shell is distinct. The dimensions of two specimens are—Length, 3¼ mm.; height, 2¾ mm.; diam., 1½ mm.; umbo 2 mm. from anterior end: length, 2¾ mm.; height, 2 mm.; diam., 1 mm.; umbo, 2 mm. from anterior end. The mean indices are—iii. = 78; vii. = 41; x. = 66.
Corneocyclas hodgkini, n. sp. Fig. 9.
(1.) Lake Waikaremoana (Stat. 18—Dredged in 800 ft.).—Two specimens. They are very small, oval, much compressed, the anterior side produced and attenuated; posterior and ventral margins regularly rounded; colour yellowish-white; beaks obtuse, unconspicuous; nepionic shell distinct, very finely concentrically striated, passing without change into the adult valves, which are irregularly finely concentrically striate. Posterior part with a ferrugineous coating. The dimensions are—Length, 2¾ mm.; height, 2 mm.; diam., 1 mm.; and the indices—iii. = 72; vii. = 36: length, 2 mm.; height, 1¾ mm.; diam., ¾ mm.; and the indices—iii. = 87; vii. = 50.
Named in honour of Mr. G. L. Hodgkin, who so ably assisted Mr. Lucas in his arduous work.
Type in my collection.
(2.) Lake Taupo (Stat. 10—Dredged in 280 ft. and 320 ft., muddy bottom).—Six small specimens, thickly coated all over with ferrugineous earth, so that the form of the shell is unrecognisable. I succeeded in cleaning one specimen, and found it to agree with the specimens just described from Lake Waikaremoana. Dimensions of the cleaned specimen—Length, 3 mm.; height, 2¼ mm.; diam., 1 mm.
Fam. HydrobiidÆ.
Genus Potamopyrgus, Stimpson (1865).
Americ. Journ. Conch., vol. i., 1865, p. 53.
Potamopyrgus corolla, Gould (1847).
Melania corolla, Gould, Proc. Bost. Soc. Nat. Hist., vol. ii., 1847, p. 223.
(1.) Lake Waikare (Stat. 19—From reeds).—One specimen, with black coating and six rounded whorls. Length, 5½ mm.; breadth, 3¾ mm.
(2.) Lake Taupo (Stat. 16 and 16F—Dredged in 280 ft. and 320 ft., muddy bottom).—Four specimens, all of which are

shouldered and with traces of stout setæ; covered with a dark-brown coating; there are seven whorls, the peritreme is continuous. The dimensions range from 7 × 4½ mm. to 6½ × 4½ mm.
Potamopyrgus corolla, Gould, subsp. salleana, Fischer (1860).
Paludestrina salleana, P. Fischer, Journ. de Conch., vol. viii., 1860, p. 208, pl. iv., fig. 6.
(1.) Lake Waikare (Stat. 14 and 14F—Among reeds).—From Stat. 14 are thirteen adult and a number of young shells; from Stat. 14F twelve specimens, exactly the same as the former. Most of them are of horn-colour, with a white calcareous coating; three only are coated with black. All, with the exception of one, are spinous, the spines being distant, rather short, bent upward. In some specimens a distinct angle below the periphery is present. All the adult shells are smaller than the type, with six whorls. The dimensions range from 4½ × 2½ mm. to 5¼ × 3¼ mm.
(2.) Lake Rotoiti (Stat. 18—Dredged with weeds in 6 ft.).—Seventeen specimens of rather uniform size and shape, covered with a greenish-black coating. There are five rounded smooth whorls; only one specimen shows traces of bristles on the upper whorls. The peritreme is sometimes, not always, black. A few specimens are approaching corolla, being more ventricose than the subspecies. The dimensions vary from 5 × 3 mm. to 6 × 4 mm.
(3.) Lake Rotoiti (Stat. 18F—Dredged with weeds in 6 ft.).—Sixteen specimens, all “dead shells.” Very much the same as the preceding. Dimensions range from 5 × 3 mm. to 5½ × 3¼ mm.
(4.) Lake Taupo (Stat. 6—Dredged in 25 ft. to 420 ft.).—Many specimens, all light-horn colour, about two-thirds with smooth rounded whorls, the others with a carina above the periphery on which very short spines are situated, having a broad base, from which two to three separate spines arise. This is a feature met with in specimens of P. corolla from Lake Kanieri, South Island. The variability of the arrangement of the spines is just as great as in P. badia. The pullus is mostly brown and shining. The first two and a half whorls are always convex, never shouldered. The peritreme is continuous and brown. Young specimens have sometimes the body-whorl angled below the periphery, but no chordate carina is present. The size is very variable, ranging from 5 × 3 mm. with six whorls to 8½ × 5 mm. with seven whorls.
(5.) Lake Taupo (Stat. 6F—Dredged in 25 ft. to 420 ft.).—About two dozen specimens, showing the same characters as those of Stat. 6. The dimensions vary from 5½ × 3 mm. to 8½ × 5 mm., with six and seven whorls respectively.
(6.) Lake Taupo (Stat. 16F—Dredged in 280 ft. and 320 ft.,

muddy bottom).—All the seven specimens are “dead shells,” therefore very fragile. They are shouldered, with traces of spines, covered with a dark-brown coating; no carina below the periphery. Dimensions range from 6½ × 4 mm. to 8½ × 5 mm.
(7.) Lake Taupo (Stat. 39—With weeds, from 20 ft. to 80 ft.).—There are seventeen specimens, of light-horn colour, and of an astonishing variability. Some call to mind the graceful, slender P. egenus, while others are more ventricose and short. Especially the elongated specimens show a very distinct angle on the body-whorl, arising from the junction of the outer lip with the whorl. Only a few are shouldered, but devoid of spines. The peritreme is dark-brown. The dimensions of four specimens are—4 × 2¾ mm.; 5½ × 2½ mm.; 6 × 3 mm.; 6 × 3½ mm. All adult specimens have six whorls.
(8.) Lake Taupo (Stat. 39F—With weeds, from 20 ft. to 80 ft.).—Four small specimens, covered with a grey coating, with six smooth convex whorls and a brown peritreme. They come very near P. antipodum, subsp. zelandiœ, but the whorls are more convex and the suture deeper. This is the most extreme form of salleana I have seen. The dimensions are—4 × 2¼ mm.; 4¼ × 2¼ mm.; 4½ × 2¼ mm.; 5 × 2½ mm.: the ratio of breadth to length varying from 1:1.6 to 1:2.
(9.) Lake Taupo (Stat. 40—With weeds, from 20 ft. to 80 ft.).—A large number of young specimens, most of which are distinctly angled below the periphery. Only two adult specimens, one smooth, one with spines. Colour, &c., the same as in examples from Stat. 39. The dimensions are 5 × 2½ mm. and 5 × 3 mm., with six whorls.
(10.) Lake Taupo (Stat. 40F—With weeds, from 20 ft. to 80 ft.).—Numerous young specimens, none adult, showing the same characters as those from Stat. 40, but they are covered by a green coating.
(11.) Lake Taupo (Stat. 47—Dredged in 400 ft.).—Four adult specimens, with a very thick black coating, very markedly shouldered, seven whorls, no spines, but there is an exceptionally strong carina; mouth snow-white. Dimensions, 7 × 4¼ mm.
(12.) Lake Waikaremoana (Stat. 3—Dredging in 50 ft. to 100 ft.).—Fifteen adult specimens, of a cinereous colour, with six whorls which are mostly shouldered and have rudimentary bristles; a few only have rounded smooth whorls. Peritreme light-brown. Dimensions range from 4½ × 2½ mm. to 6 × 3½ mm. These may be considered as typical forms.
(13.) Lake Waikaremoana (Stat. 17—Dredging in 800 ft.).—One specimen, a “dead shell,” with the last whorl broken off. It is thickly coated with calcareous substance, stained orange by oxide of iron. There are rudimentary spines visible on one

whorl. Currents very likely brought this specimen to this considerable depth.
(14.) Lake Waikaremoana (Stat. 34—Dredging in 10 ft. to 20 ft.).—There are thirty-four adult shells, cinereous to black according to the amount of coating covering the shell. Most of them have six rounded smooth whorls, a few only are shouldered, none have spines. Peritreme brown. The dimensions vary from 5 × 2¾ mm. to 6½ × 3¾ mm. These also are typical examples.
(15.) Lake Manapouri (Stat. 15—From shallow water near the shore).—Two large adult specimens, both of which are of horn-colour, broadly shouldered, with distant brown spines on the carina, two or three arising from a common broad base. The larger example shows distinctly the slightly chordate carina below the row of spines on the last whorl, a character mentioned by P. Fischer. The other species, however, shows no trace of it; and it is, as I have pointed out elsewhere, not a constant but an extremely rare feature of salleana. The dimensions of the two shells are—8 × 5 mm. and 7 × 4 mm.
I have similar specimens from Lake Kanieri, kindly collected for me by Dr. Macandrew, of Hokitika, but they are more ventricose, and I assign them to P. corolla.
Potamopyrgus badia, Gould (1848).
Amnicola badia, Gould, Proc. Bost. Soc. Nat. Hist., vol. iii., 1848, p. 75.
(1.) Lake Wakatipu (Stat. 10—Dredged with weeds in 200 ft. to 300 ft.).—Seven shells of very light horn-colour, rather variable in size and shape. Only one, the largest, has spines; the others have the whorls convex, a few showing indications of a keel. They are very thin and fragile, the peritreme continuous in all, and light-brown. The largest shell measures 7 × 3½ mm., and it is very similar to the large specimens found in Lake Te Anau. The smallest shell, with six whorls, is 5 × 2½ mm.
(2.) Lake Wakatipu (Stat. 18—Dredged with weeds in 20 ft. to 100 ft.).—Ten shells, three of which are not adult, very variable in size, of light-horn colour, and very thin. Four have smooth, rounded whorls, two are slightly shouldered, and four are spinous. On the lower whorls the setæ are far apart, sometimes two to four bristles arising from a common base. All have six whorls, and the dimensions range from 5½ × 2½ mm., 6 × 3¼ mm., to 6½ × 3½ mm.
(3.) Lake Wakatipu (Stat. 6—From weeds fringing shore).—Seven shells of horn-colour, covered with a thin white coating, six convex whorls, suture impressed. Some examples are slightly shouldered on the upper whorls, with minute close-set short bristles. All are of about the same size—4¼ × 2¼ mm.

Fam. Limnæidæ.
Sub-fam. Ancylinæ.
Genus Gundlachia, Pfeiffer (1849).
Zeitschrift f. Malak., vol. vi., 1849, p. 98.
Gundlachia lucasi, n. sp. Figs. 10, 11.
Lake Waikare (Stat. 30 and 30F—Netting in weeds).—Three specimens; two of them were collected alive. Shell obliquely conical, thin, semitransparent, horn-colour, covered by a blackish coating; apex inclined to the right, situated at the posterior third of the length; convex anteriorly, slightly concave on the posterior slope; a few concentric lines of growth. Aperture oval; peritreme sharp, extremely fragile. No septum, the shells being in the Ancylus stage of development only. Dimensions of two specimens—Length, 3 mm.; breadth, 2 mm.; height, 1 mm.: length, 4 mm.; breadth, 2¾ mm.; height, 1½ mm. The dentition is very similar to that of the Gundlachia sp. from the River Avon,* which, settles the generic position.
This species stands nearest to G. tasmanica, T.-Woods. It is more rounded and elevated than the species from the River Avon. There is also one specimen of G. lucasi from Inglewood in my collection.
The occurrence of two species of Gundlachia in three different localities leaves no doubt that the genus is endemic, and accidental introduction out of the question.
Type in my collection.
I have great pleasure in uniting with the species the name of Mr. K. Lucas, who so ably and successfully collected the fauna of New Zealand lakes.
Sub-fam. Latiinæ.
Genus Latia, Gray (1850).
Latia neritoides, Gray (1850).
P.Z.S., 1849, p. 168 (1850).
Lake Waikare (Stat. 35—Dredged in 4 ft.; stony shore).—One small typical specimen, 5 mm. long.
Sub-fam. Limnæinæ.
Genus Amphipeplea, Nilsson (1822).
Nilsson, Hist. Moll. Suec., p. 58.
Amphipeplea arguta, Hutton (1885).
Trans. N.Z. Inst., vol. xvii., p. 54, pl. xii., fig. 1.
(1.) Lake Waikare (Stat. 19 and 19F—From reeds).—Fourteen
[Footnote] * Trans. N.Z. Inst., xxvi., pl. xiv., fig. 5.

mostly young shells. Colour horny, columellar lip broadly reflexed. Dimensions—Shell: length, 7 mm.; breadth, 4 mm. Aperture: height, 5 mm.; breadth, 4 mm. Shell: length, 5½ mm.; breadth, 4 mm. Aperture: height, 4½ mm.; breadth, 3 mm.
(2.) Lake Taupo (Stat. 42—With weeds from 20 ft. to 80 ft.).—One specimen collected alive; yellowish-white, rather slender. The aperture more elongated than in the type. Shell: length, 5½ mm.; breadth, 3½ mm. Aperture: height, 4 mm.; breadth, 2½ mm.
(3.) Lake Wakatipu (Stat. 10—Dredged with weeds in 200 ft. to 300 ft.).—One specimen collected alive; nearly colourless, transparent, very thin and fragile, with broad columellar reflection; regular distinct incremental lines. The aperture is longer and narrower than in the type. Shell: length, 5¼ mm.; breadth, 3¼ mm. Aperture: height, 4½ mm.; breadth, 2½ mm.
(4.) Lake Wakatipu (Stat. 33—Dredged in 30 ft. to 60 ft.).—Two specimens collected alive. A slender form with elongated aperture, of horn-colour, with regular lines of growth, and the spire a little higher than typical. Shell: length, 5 mm.; breadth, 3 mm. Aperture: height, 3½ mm.; breadth, 2¼ mm. Shell: length, 4 mm.; breadth, 2½ mm. Aperture: height, 3½ mm.; breadth, 1¾ mm.
The Dentition.—Figs. 11–14 represent the most characteristic teeth of the radula of specimens from the four localities—fig. 11 from Lake Waikare, fig. 12 from Lake Taupo, fig. 13 from Stat. 10, and fig. 14 from Stat. 33, Lake Wakatipu. Compared with Hutton's description and figure of his species* a considerable variability, especially in the transitional teeth, is at once apparent. The central tooth shows mostly a second small denticle on the left side; the lateral teeth have all three cutting-points, but the entocone and mesocone may coalesce, forming only one cutting-point, as was evidently the case in the example
[Footnote] * Trans. N.Z. Inst., xvii., p. 54, pl. xii., fig. 10.

figured by Hutton. The number of laterals and marginals is very variable, and the figures sufficiently show the different arrangements of the cutting-points on the transitional teeth.
Sub-fam. Planorbinæ
Genus Planorbis (Guett.), Geoffroy (1767).
Planorbis (Gyraulus) corinna, Gray (1850).
P.Z.S., 1849, p. 167 (1850).
Lake Waikare (Stat. 33—Obtained by netting in weeds).—One specimen only was found, which has three whorls and a diameter of 2½ mm.
Genus Isidora, Ehrenberg (1831).
Isidora tabulata, Gould, subsp. moesta, H. Adams (1861). P.Z.S., 1861, p. 144.
(1.) Lake Waikare (Stat. 19—From reeds).—One specimen only was obtained. It is of light-horn colour, very thin, covered with a greenish coating; there are four whorls, the last two distinctly shouldered and keeled. Columella excavated in the middle, fold distinct, reflection of columellar lip small; outer lip sharp, regularly arched. It is a little more slender than the type. Shell: length, 7 mm.; breadth, 4 mm. Aperture: height, 5 mm.; breadth, 2½ mm. Ratios: i. = 1: 1.75; ii. = 1: 2; iii. = 1: 1.4. (The ratios are the same as in the revision of Isidora.)
(2.) Lake Waikare (Stat. 28—From weeds in water from 1½ ft. to 2 ft.).—Two specimens of dark-brown colour, solid and large, keeled, the keel becoming obsolete on approaching the aperture; columella twisted. They differ but little from the type. Shell: length, 17½ mm.; breadth, 11 mm. Aperture: height, 11 mm.; breadth, 6 mm. Shell: length, 14 mm.; breadth, 9½ mm. Aperture: height, 10 mm.; breadth, 5 mm. Ratios: i. = 1: 1.6; ii. = 1: 1.8; iii. = 1: 1.6. i. = 1: 1.5; ii. = 1: 2; iii. = 1: 1.4. The accompanying fig. 15 shows some teeth of the radula, which need no explanation. The dentition of our forms of Isidora is so variable that I doubt whether it can be used as a help to separate the species. However, a considerable number of animals of each species has to be examined before this point can be settled definitively.
(3.) Lake Waikare (Stat. 28F—From weeds in water from 1½ ft. to 2 ft.).—Two “dead shells,” one young, the other nearly adult. They resemble those from the last station, but are a little more ventricose, and have the spire a little shorter. Shell:

length, 15 mm.; breadth, 10 mm. Aperture: height, 10 mm.; breadth, 5.5 mm. Shell: length, 9 mm.; breadth, 6½ mm. Aperture: height, 6 mm.; breadth, 3½ mm. Ratios: i. = 1: 1.5; ii. = 1: 1.8; iii. = 1: 1.5. i. = 1: 1.5; ii. = 1: 1.7; iii. = 1: 1.5.
Isidora lirata, Tenison-Woods (1879).
P.L.S. N.S.W., vol. iii., 1879, p. 138, pl. xiii., fig. 6.
(1.) Lake Rotoiti (Stat. 12—From weeds in 6 ft.).—Six specimens, one only adult. The colour is light-horny, some specimens having a dark coating; the whorls are rounded, but slightly flattened below the suture, the spiral striæ are present but not very distinct. Aperture produced at the base, columella twisted. They are slightly more ventricose than the type. Shell: length, 12 mm.; breadth, 7 mm. Aperture: height, 8½ mm.; breadth, 4 mm. Shell: length, 10 mm.; breadth, 6 mm. Aperture: height, 7 mm.; breadth, 3½ mm. Shell: length, 8½ mm.; breadth, 5½ mm. Aperture: height, 5 mm.; breadth, 3 mm. Ratios: i. = 1: 1.7; ii. = 1: 2.1; iii. = 1: 1.4. i. = 1: 1.7; ii. = 1: 2; iii. = 1: 1.4. i. = 1: 1.5; ii. = 1: 1.7; iii. = 1: 1.7.
(2.) Lake Taupo (Stat. 7—Dredged with weeds in 75 ft.).—One specimen of cream colour, fragile; four whorls, shouldered down to the middle of the last whorl, carina with short bristles; shell indistinctly spirally striated; columella twisted, aperture much produced anteriorly. More ventricose than the type. Shell: length, 8 mm.; breadth, 5 mm. Aperture: height, 6 mm.; breadth, 3 mm. Ratios: i. = 1: 1.6; ii. = 1: 2; iii. = 1: 1.3.
(3.) Lake Taupo (Stat. 38—With weeds, from 80 ft.).—Many, mostly young specimens. They are of a dirty-white colour with a thin whitish coating. Whorls 4, rounded, sometimes flattened below the suture, beautifully regularly spirally striate. Columella twisted, aperture produced anteriorly. These I consider to be typical forms. Shell: length, 9½ mm.; breadth, 6 mm. Aperture: height, 7 mm.; breadth, 3 mm. Shell: length, 8½ mm.; breadth, 4¾ mm. Aperture: height, 6 mm.; breadth, 3 mm. Shell: length, 7½ mm.; breadth, 4½ mm. Aperture: height, 5 mm.; breadth, 2½ mm. Ratios: i. = 1: 1.6; ii. = 1: 2.3; iii. = 1: 1.4. i. = 1: 1.8; ii. = 1: 2; iii. = 1: 1.4. i. = 1: 1.7; ii. = 1: 2; iii. = 1: 1.6. Fig. 16 shows some teeth of the radula. A remarkable feature are the additional denticles on the outer upper side of the marginal teeth.
(4.) Lake Taupo (Stat. 38F—With weeds, from 80 ft.).—Many, mostly young shells. Very much like the specimens from

Stat. 38, but the spiral striation is not so distinct, and the coating is thicker, of a greenish hue.
(5.) Lake Waikaremoana (Stat. 4—Dredging in 50 ft.).—Two “dead shells,” one adult with the apex broken off. They are rather large, of cream colour, with four rounded but slightly shouldered whorls; distinctly spirally lirate. Columella twisted, and aperture produced at base. Typical forms, though one much larger. Shell: length, 15½ mm.; breadth, 9 mm. Aperture: height, 10 mm.; breadth, 5 mm. Shell: length, 10½ mm.; breadth, 6 mm. Aperture: height, 7 mm.; breadth, 3 mm. Ratios: i. = 1: 1.7; ii. = 1: 2; iii. = 1: 1.6. i. = 1: 1.8; ii. = 1: 2.3; iii. = 1: 1.5.
(6.) Lake Waikaremoana (Stat. 5—Dredging in 50 ft.).—Six specimens, two of them quite young. The colour is light-horny, some shells with a ferrugineous coating. The whorls in some are rounded, in others shouldered and the keel ornamented with short bristles. In young specimens the spiral striation is quite distinct. Columella twisted, and aperture produced anteriorly. These again are typical forms. Dimensions of largest specimen: Shell: length, 13 mm.; breadth, 6¾ mm. Aperture: height, 8 mm.; breadth, 4 mm. Ratios: i. = 1: 1.9; ii. = 1: 2; iii. = 1: 1.6.
(7.) Lake Waikaremoana (Stat. 32—Dredging in 10 ft. to 20 ft.).—Four nearly adult and four young specimens. They are dirty-white, the longer examples with a light-brown coating. The four whorls are convex, very little flattened below the suture; spiral striation indistinct. These shells are very nearly typical. Dimensions of largest specimen:—Shell: length, 12 mm.; breadth, 6½ mm. Aperture: height, 8.5 mm.; breadth, 4 mm. Ratios: i. = 1: 1.8; ii. = 1: 2.1; iii. = 1: 1.4.
Synopsis of the Molluscan Fauna of the Six Lakes.
Lake Waikare.
Diplodon menziesi, Gray, subsp. rugata, Hutton.
Corneocyclas novozeelandica, Prime.
Potamopyrgus corolla, Gould.
" " " subsp. salleana, Fischer.
Gundlachia lucasi, Suter.
Latia neritoides, Gray.
Amphipeplea arguta, Hutton.
Planorbis corinna, Gray.
Isidora tabulata, Gould, subsp. moesta, H. Adams.
Lake Rotoiti.
Diplodon menziesi, Gray, subsp. hochstetteri, Dunker.
Sphærium novæ-zelandiœ, Deshayes.

Potamopyrgus corolla, Gould, subsp. salleana, Fischer.
Isidora lirata, Tenison-Woods.
Lake Taupo.
Diplodon menziesi, Gray.
Sphærium novæ-zelandiœ, Deshayes.
Corneocyclas hodgkini, Suter.
Potamopyrgus corolla, Gould.
" " " subsp. salleana, Fischer.
Amphipeplea arguta, Hutton.
Isidora lirata, Tenison-Woods.
Lake Waikaremoana.
Diplodon menziesi, Gray.
Sphærium novæ-zelandiœ, Deshayes.
Corneocyclas hodgkini, Suter.
Potamopyrgus corolla, Gould, subsp. salleana, Fischer.
Isidora lirata, Tenison-Woods.
Lake Wakatipu.
Diplodon lessoni, Kuster.
Corneocyclas novozeelandica, Prime.
Potamopyrgus badia, Gould.
Amphipeplea arguta, Hutton.
Lake Manapouri.
Diplodon menziesi, Gray, subsp. lucasi, Suter.
Corneocyclas novozeelandica, Prime.
Potamopyrgus corolla, Gould, subsp. salleana, Fischer.
" badia, Gould.
Bathymetrical Distribution of the Mollusca.
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
| —— | Lake. | Depth in Feet. |
|---|---|---|
| Diplodon menziesi, Gray | Taupo | 100 |
| " " | Waikaremoana | 10–50 |
| " " subsp. hochstetteri, Dunker | Rotoiti | 12 |
| " " subsp. hochstetteri, Dunker | Taupo | 10–30 |
| " " subsp. rugata, Hutton | Waikare | 2 |
| " " subsp. lucasi, Suter | Manapouri | 60 |
| Diplodon lessoni, Kuster | Wakatipu | 10–30 |
| Sphærium novæ-zelandiæ, Deshayes | Rotoiti | 6 |
| " " | Taupo | 20–100 |
| " " | Waikaremoana | 10–20 |
| Corneocyclas novo-zeelandica, Prime | Waikare | 2 |
| " " | Wakatipu | 10–100 |
| " " | Manapouri | Shallow |
| Corneocyclas hodgkim, Suter | Taupo | 280–320 |
| " " | Waikaremoana | 800 |
| Potamopyrgus corolla, Gould | Waikare | Shallow |
| " | Taupo | 280–320 |
| " " subsp. salleana, Fischer | Waikare | Shallow |
| " " subsp. salleana, Fischer | Rotoiti | 6 |
| " " subsp. salleana, Fischer | Taupo | 20–420 |
| " " subsp. salleana, Fischer | Waikaremoana | 10–100 |
| " " subsp. salleana, Fischer (dead) | " | 800 |
| " " subsp. salleana, Fischer | Manpouri | Shallow |
| Potamopyrgus badia, Gould | Wakatipu | 20–300 |
| " " | Manapouri | Shallow |
| Gundlachia lucasi, Suter | Waikare | " |
| Latia neritoides, Gray | " | 4 |
| Amphipeplea arguta, Hutton | " | Shallow |
| " " | Taupo | 20–80 |
| " " | Wakatipu | 20–300 |
| Planorbis corinna, Gray | Waikare | Shallow |
| Isidora tabulata, Gould, subsp. moesta, H. Adams | " | 2 |
| Isidora lirata, Tenison-Woods | Rotoiti | 6 |
| " " | Taupo | 75–80 |
| " " | Waikaremoana | 10–50 |

Summary.
The Mollusca collected by Messrs. Lucas and Hodgkin in the six lakes of New Zealand belong to nine genera, and these are represented by twelve species and five subspecies. As new we find one subspecies of Diplodon menziesi, one species of Corneocyclas, and one species of Gundlachia. Am interesting addition to our fauna is Diplodonta lessoni, Küster.
From Lake Rotoiti no specimens of Potamopyrgus badia, Gould, and Melanopsis trifasciata, Gray, were obtained, but it is almost certain that Potamopyrgus corolla, subsp. salleana, was taken for P. badia, and I never have seen an example of Melanopsis trifasciata from Lake Rotoiti, though I tried hard to get some. In the Lake of Tiberiade, in Palestine, Melanopsis has been found at a depth of about 150 ft.

From Lake Wakatipu Planorbis corinna and Isidora antipodea were not represented.
All the Mollusca living in deeper water have much lighter colours, and the test is much thinner and more fragile. With one exception (Corneocyclas hodgkini) there is no real deep-lake fauna in these lakes, such as it is known from the subalpine lakes in Switzerland, &c., and this may partly be explained by the poverty of our lakes in molluscan life. Limnæa is absent, its place being taken by Amphipeplea, and it is interesting to find this genus represented by one species only.
The littoral fauna, according to Forel's investigations,* extends to 50–75 ft., and it is therefore evident that many of our known species can live in deeper water without undergoing any great structural change. Generally Unionidœ are not found in very deep water. However, Anodonta ponderosa, Pfr., was obtained in 33 ft. in Lake Tschaldyr, in Armenia, by Dr. Brandt, and five species of Unio were dredged in 150–300 ft. in Lake Tiberiade, in Palestine, by Lortet.
Sphœrium and Isidora have, as far as I know, not been known to live in deeper water, and of Potamopyrgus it is for the first time we get any accurate knowledge of the depths of water in which it may be found living.
The deep-lake molluscan fauna of New Zealand, as far as Messrs. Lucas and Hodgkin's investigations go, may be considered to be composed of the following species:—
(1.) Diplodon menziesi, Gray.
(2.) " " subsp. lucasi, Suter.
(3.) Sphærium novæ-zelandiœ, Deshayes.
(4.) Corneocyclas novozeelandica, Prime.
(5.) " hodgkini, Suter.
(6.) Potamopyrgus corolla, Gould.
(7.) " " subsp. salleana, Fischer.
(8.) Potamopyrgus badia, Gould.
(9.) Amphipeplea arguta, Hutton.
(10.) Isidora lirata, Tenison-Woods.
[Footnote] * Dr. F. A. Forel, “La Faune profonde des Lacs Suisses,” 1885.

Art. XIV.—The First-discovered New Zealand Gundlachia
[Read before the Wellington Philosophical Society, 7th September, 1904.]
Gundlachia neozelanica, n. sp.
Ancylus, sp., Suter, Journ. de Conch., vol. xl., pp. 248–250 (1892). Ancylus tasmanicus, Suter, P.L.S. N.S.W. (2), vol. vii., p. 624 (1893), (not of T.-Woods). Gundlachia, sp., Suter, Trans. N.Z. Inst., vol. xxvi., p. 122, pl. xiv., figs. 1–5 (1894); Gundlachia, sp., Suter, P.L.S. N.S.W. (2), vol. viii., p. 486 (1894); Gundlachia, sp., Hedley, l.c., pp. 505, 507, 511, pl. xxiv., figs. 12–15; Gundlachia, sp., Suter, Journ. de Conch., vol. xli., p. 229 (1894).
Shell depressed-conoidal, oval-oblong, thin, subtransparent, horn-colour, with a blackish-green coating; apex a little inclined to the right, situated at the posterior sixth of the length, flatly convex anteriorly; concentric lines of growth at regular intervals. Interior light-brown, shining; aperture elongated-oval, slightly broadened anteriorly. Length, 3 mm.; breadth, 2 mm.; height, ¾ mm.
Hab. River Avon, near Christchurch.
Type in my collection.
With regard to the dentition I have to rectify a mistake in the figure: the rhachidian tooth is bicuspid, not tricuspid.
The formation of a septum has been described and figured by Hedley (l.c.) from specimens I sent him.
I always hesitated giving this species a name, as I hoped fully developed specimens might turn up. This, however, has not been the case, and, as a second species has been discovered, it is incumbent to name the first-discovered form.
Art. XV.—Revision of the New Zealand Species of the Genus Potamopyrgus, with Description of a New Species.
[Read before the Wellington Philosophical Society, 7th September, 1904.]
A Revision of the New Zealand Hydrobiinæ was published by Captain F. W. Hutton in 1882,* reducing the already described
[Footnote] * Trans. N.Z. Inst., vol. xiv., pp. 143–146, pl. i., figs. A-H.

species to three, and adding a new species (P. pupoides). Hutton says: “The absence of books prevents me feeling certain that all the synonyms I have given are correct.” With regard to books we are not much better off than we were twenty-two years ago, and, besides this, there is the very great inconvenience for us that the types of all the species, Hutton's species excluded, are in foreign museums. It was many years back, when material in my collection was fast accumulating, that I found Hutton's restriction to a total of only four species unsatisfactory. I am fully aware of the great variability of fresh-water molluscs, also of the fact that many species of Potamopyrgus are polymorphic, and therefore one and the same species may have been described under different names. There is a spinous angulate form, then an angulate espinous form, and thirdly an acuminate ecarinate one. Of the New Zealand species only two are polymorphic.
I tried to get as much information as I possibly could about those species of which I did not possess sufficient knowledge, and I have to thank especially Dr. W. H. Dall, Hon. Curator of Mollusks, U.S. National Museum, Washington; also Dr. H. Fischer, of Paris; Dr. R. Sturany and Dr. Oberwimmer, K.K. Hofmuseum, Vienna, for the great readiness with which they acceded to my request. The revision now undertaken is to a large extent based on the information thus obtained, and I hope it may prove useful to students of conchology.
The species of Potamopyrgus described from New Zealand up to now number eleven (omitting crossei, Frfld., ciliata, Gould, and gracilis, Gould, for reasons shown later on), and they were formerly classed under five genera: Melania, Amnicola, Paludestrina, Hydrobia, and Bythinella. These eleven species I now reduce to five, with three subspecies.
Genus Potamopyrgus, Stimpson (1865); Stimpson, Amer. Journ. of Conchology, vol. i. (1865), p. 53; Smithon, Miscell. Coll., No. 201 (1865), pp. 49, 50.
Type: Melania corolla, Gould, Proc. Bort. Soc. Nat. Hist., vol. ii. (1874), p. 223.
Synonym: Pyrgophorus, Ancey, Bull. Soc. Mal. France, vol. v. (1888), pp. 188, 192.
Stimpson's diagnosis of the genus is reproduced by Hutton in Trans. N.Z. Inst., vol. xiv., p. 143, with some additional remarks on the dentition.
Pilsbry says, “Potamopyrgus is a genus of great antiquity, extending at least as far back as the early Eocene. It now comprises all of the fresh-water rissoids of New Zealand, a majority of those of Australia, with species in West Africa and

tropical America.”* Tasmania has forms very nearly allied to ours. The genus is, as far as I am aware, not known in the fossil state from New Zealand.
Potamopyrgus corolla, Gould, sp.
Melania corolla, Gould, Proc. Bost. Soc. Nat. Hist., vol. ii. (1847), p. 223. Melania corolla, Reeve, Conch. Sc. (Melania), fig. 366. Amnicola corolla, Gould, U.S. Expl. Exp., vol. xii. (1852), p. 129, pl. ix., figs. 149 a-c. Paludestrina cumingiana, P. Fischer, Journ. de Conch., vol. viii. (1860), p. 208, pl. iv., fig. 7. Potamopyrgus corolla, Stimpson, Smithon. Miscell. Coll. No. 201 (1865), pp. 49, 50. Hydrobia crossei, Frauenfeld, Verh. Zool. Bot. Gesell. Wien, vol. xiv. (1865), p. 595. (?) Hydrobia ciliata, Frauenfeld, l.c., p. 1025. Bythinella corolla, Tenison-Woods, P.L.S. N.S.W., vol. iii. (1879), p. 135, pl. xiii., figs. 2, 3, 5. Hydrobia corolla, v. Martens, Crit. List N.Z. Moll. (1873), p. 14. Hydrobia corolla, Hutton, Manual N.Z. Moll. (1880), p. 83. Potamopyrgus cumingiana, Hutton, Trans. N.Z. Inst., vol. xiv. (1882), p. 144, pl. i., figs. A, F. Potamopyrgus cumingiana, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii. (1893), p. 619. Potamopyrgus cumingiana, Suter, Journ. de Conch., vol. xli. (1893), p. 222.
Dr. Sturany, of Vienna, kindly informed me (in lit.) that Frauenfeld proposed the specific name crossei solely in case the name of Fischer, Paludestrina cumingiana, should be accepted, as there exists a Paludestrina cumingi, d'Orb., of earlier date.
Regarding the species Hydrobia ciliata, Gould, I am indebted to Dr. W. H. Dall, of Washington, for the following (in lit.): “Described as from Liberia with other shells from the missionary Dr. Perkins, but appears very similar to Hutton's figure in Trans. N.Z. Inst., vol. xiv., pl. i., fig. B 1. It seems as if some mistake had occurred, and this shell should really have come from New Zealand, but it is difficult to see how, as Gould was a very careful man.” Dr. P. Fischer no doubt admitted this species to be of African origin, for he says, “Nous connaissons deux autres espèces de la Nouvelle-Zélande, une des Antilles, une de l'Afrique; toutes sont caracterisée par la série d'épines qui ornent le dernier tour de spire.”† Pilsbry also, as already stated, gives West Africa as the habitat of Potamopyrgus, and is most likely alluding to the above species.
Dr. W. H. Dall (in lit. 20, xi., 97) also kindly sent me the following information respecting the type of the genus: “Melania
[Footnote] * Proc. Acad. N. Sci., Philadelphia, 1891, p. 327.
[Footnote] † Journ. de Conch., vol. viii. (1860), p. 209.

corolla, Gould (1847), is not corolla of Hutton, but the shell figured by Reeve as corolla. It is the largest, with spare long spines. The locality may have been erroneous. The whorls are rounded, cf. Hutton, Trans. N.Z. Inst., vol. xiv., pl. i., fig. A 1.” Considering this statement, we must conclude that the synonyms given by Hutton in his Manual, p. 83, are mostly correct, but that in his revision he took the wrong species for corolla—viz., Amnicola badia, Gould. He says (l.c., p. 143) that P. cumingiana, Fischer, which is held by v. Martens to be the same species as P. corolla, Gould, does not occur on Banks Peninsula, and that its dentition does not correspond with the description given by Dr. Stimpson. Now, Dr. Dall admits that the locality given by Gould may have been erroneous, and this appears now to be quite certain. I examined the dentition in typical specimens of P. corolla from the Ruamahanga River, Wairarapa, and I cannot say that it does not agree with Stimpson's description. The rhachidian and the lateral teeth correspond with his diagnosis, but the marginal teeth have a number of larger denticles, their number agreeing with Stimpson's statement, but there are a number of minute denticles besides which I am unable to count with a magnifying-power of 720. The number of denticles on the marginal teeth is variable, and can hardly be used for specific distinction.
The diagnosis of the species is as follows: Shell horn-colour, sometimes covered with a black coating, ventricose, thin, sub-pellucid; 6½ rounded whorls, the last three spinous, the others smooth and sometimes of a darker colour, the last swollen, subcarinated at the encircling series of spines. Spines long, rather distant, curved, directed upwards. Aperture ovate, peristome continous, thickened. There may be only rudimentary spines, or they may be absent altogether, leaving only a carina, or, finally, all the whorls may be smooth and convex, without a trace of a carina or spines. Length, 6.5 mm.; breadth, 4.5 mm. Ratio of B: L = 1: 1.44.
This species is distinguished by its globosely ventricose form, the absence of spines on the first 3½ whorls, the great number of spines on the last whorl (17 to 20), their length, curvature, and direction towards the apex.
My collection contains specimens from fourteen localities:—
(1.) Typical form with long spines is represented from Lakes Takapuna and Kanieri. Those from the latter locality have shorter spines. Most of the specimens are of horn-colour, sometimes with the first few whorls darker, and a few have a dark-brown coating.
(2.) Specimens with rudimentary spines occur in the creeks at Henderson and Swanson; in the creek at Hastwell's and

Ruamahanga River in the Forty-mile Bush; at Petone, near Wellington; Parua Bay, near Whangarei; and on the Chatham Islands. The majority of these specimens have a thick, black coating, and the continuous peristome much thickened.
(3.) With only a thread-like carina on the whorls there are specimens from Hastwell's and the Chatham Islands.
(4.) Shells with the whorls smooth, rounded, mostly coated with black and callous peristome, were found in creeks at North-cote, Hastwell's, Parua Bay, Kawau Island, the Waikato, and Chatham Islands.
(5.) Dwarf forms with spines, horn-colour, and six whorls, were collected in Lake Takapuna (3½ × 4½ mm.), and near Greymouth (4 × 6 mm.).
Type in the U.S. Nat. Mus., Washington.
Potamopyrgus corolla, Gould, subsp. salleana, P. Fischer, subsp.
Paludestrina salleana, P. Fischer, Journ. de Conch, vol. viii. (1860), pp. 208, 209, pl. iv., fig. 6.
Shell horn-colour, conical, thin; whorls 7 to 7½, flatly convex, the last four spinous, the others smooth, the last whorl with a chordate carina below the row of spines. The setæ are short, black, directed upwards. Aperture ovate, margins continuous, but little callous. Length, 6 mm.; breadth, 3½ mm. Ratio of B: L = 1: 1.7.
This subspecies differs from corolla in being more slender, less ventricose, in having spines on the last four whorls and a chordate carina below the row of spines; the last whorl is also less voluminous in proportion.
Although I have examined many hundreds of specimens I have not yet seen one with a chordate carina below the row of setæ, but some specimens from the Great Barrier Island and from Nelson are distinctly angled below the periphery, and I take this quite peculiar character mentioned by Fischer as of quite exceptional occurrence.
The distribution of this subspecies extends over a large area. In my collection there are specimens from Lake Takapuna, Western Springs (Auckland), Lake St. John, Onehunga Springs, Great Barrier Island, Maketu (Hunua Range), Waipoua River (near Masterton), Petone (near Wellington), Pelorus River, Nelson, and Collingwood.
All specimens are yellowish-brown, thinner than corolla, mostly without spines, and some are larger than the type. Length, 8 mm.; breadth, 4½ mm. The Nelson specimens are from brackish water.
Type in the collection of the Journal de Conchyliologie, Paris.

Potamopyrgus antipodum, Gray, sp. (em.).
Amnicola antipodanum, Gray, in Dieffenbach's “New Zealand,” vol. ii. (1843), p. 241. Hydrobia antipodum, v. Martens, Crit. List N.Z. Moll. (1873), p. 14. Hydrobia antipodum, E. A. Smith, Voy. “Erebus” and “Terror,” Zool., vol. ii. (1875), p. 3, pl. i., fig. 19, above. Bythinella antipoda, Hutton, Manual N.Z. Moll. (1880), p. 81. Potamopyrgus antipodum, Hutton, Trans. N.Z. Inst., vol. xiv. (1882), p. 145, pl. i., figs. C, G (except upper right figure). Potamopyrgus antipodum, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii. (1893), p. 619. Potamopyrgus antipodum, Suter, Journ. de Conch., vol. xli. (1893), p. 221.
The statement made by v. Martens (l.c., p. 141) that “some specimens are bristly” is not correct. This species has never been found with spines or a carina.
The diagnosis of the species may be found in Hutton's Manual, and in Trans. N.Z. Inst., vol. xiv.
P. antipodum is just as variable as most species of the genus, especially in size; but the conical form, flatly convex whorls, and little-impressed suture are fairly constant characters.
I have specimens from twelve localities, ranging from Auckland to Southland. Specimens from Owaka (Clutha) are very small and ventricose, with five whorls only, and measuring 4½ × 2¾ mm. Another dwarf form was collected by Mr. R. Murdoch, of Wanganui, on the Waimate Plains.
The great majority of the specimens are covered with a black coating. It is remarkable that P. antipodum is very often met with in brackish water, and almost without an exception such specimens have the tip of the shell eroded. In some places I saw this species living on Ulva plants. I have specimens from brackish water from seven different localities.
Type in the British Museum.
Potamopyrgus antipodum, Gray, subsp. zelandiœ, Gray, subsp. Amnicola (?) zelandiœ, Gray, in Dieffenbach's “New Zealand,” vol. ii. (1843), p. 241. Hydrobia zelandiœ, v. Martens, Crit. List. N.Z. Moll. (1873), p. 15. Hydrobia zelandiœ, E. A. Smith, Voy. “Erebus” and “Terror,” Zool., vol. ii. (1875), p. 3, pl. i., fig. 19, below. Bythinella zelandiœ, Hutton, Manual N.Z. Moll. (1880), p. 81.
This subspecies is fairly well represented by the upper right figure on pl. i., fig. C, in Trans. N.Z. Inst., vol. xiv. It is distinguished from the species by its somewhat smaller size and more tapering form. Dimensions of type are: Length, 5mm.; breadth, 2½ mm. It is much rarer than antipodum, but also shows a good amount of variation.

My collection contains specimens from Nelson, three localities near Wellington, Wanganui, Hastwell's, and Riverhead (near Auckland), the latter being from brackish water, and small.
Type in the British Museum.
Potamopyrgus badia, Gould, sp.
Amnicola badia, Gould, Proc. Bost. Soc. Nat. Hist., vol. iii. (1848), p. 75; U.S. Expl. Exp., vol. xii. (1852), p. 126, fig. 150. Hydrobia fischeri, Dunker, Mal. Blätter, vol. viii. (1862), p. 152. Hydrobia reevei, Frauenfeld, Verh. Zool. Bot. Gesell. Wien, vol. xiii., p. 1024. Hydrobia fischeri, badia, reevei, v. Martens, Crit. List. N.Z. Moll., pp. 14, 15. Bythinella fischeri, badia, reevei, Hutton, Manual N.Z. Moll. (1880), p. 82. Potamopyrgus corolla, Hutton, Trans. N.Z. Inst., vol. xiv. (1882), p. 145, pl. i., figs. B, F (not of Gould). Potamopyrgus corolla, Hedley and Suter, P.L.S. N.S.W. (2), vol. viii. (1893), p. 619 (not of Gould). Potamopyrgus corolla, Suter, Journ de Conch., vol. xli. (1893), p. 619 (not of Gould).
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
The type of this species is not spiny, the whorls moderately convex, shouldered above, and the dimensions given are: Length, ⅕ in. = 5 mm.; breadth, 1/12 in. = 2 mm.; ratio of B: L = 1:2.5.
This is the species that. since 1882 was erroneously called corolla by New Zealand conchologists. I have specimens from the River Avon which perfectly agree with the description and measurement given by Gould. Some are spinous, some shouldered and without spines, and others have smooth, flatishly rounded whorls. These typical specimens are, according to my experience, very rare, and it is curious that this form, instead of the very common one, should have been collected. The common form has the following dimensions—length, 5 mm.; breadth, 2½–3 mm.—having thus a somewhat greater angle of the spine. Both forms were found living together in the River Avon.
I know this species from the South Island only, and the finest specimens I found in the Rivers Avon and Heathcote, near Christchurch. Specimens from Akaroa have smooth, rounded whorls; a globose form, also smooth, and measuring 4 ½ × 3 mm., comes from Kowai Bush. Similar smooth forms are in my collection from Mount Somers, Bealey, Birch Hill (Tasman Valley), and Opawa, near Albury, the latter agreeing with H. fischeri. From the Leith, Dunedin, smooth and spinous forms are mixed, but the former are more abundant. I mentioned the occurrence of a large form of P. cumingiana from Lake Te Anau in these Transactions (vol. xxvi., p. 121), measuring 8 × 4 mm. Careful

examination and comparison have now convinced me that it is really a very large form of P. badia.
Considering the great variability of Potamopyrgus I refrain from establishing any new species or subspecies unless for very good reasons, and merely mention the localities where forms differing considerably from the type have been found. Temperature, chemical composition, movement and size of the water-area, and food available have a great influence on the growth of fresh-water shells, and it is difficult to find the same form of a species in more than two or three localities. Taking into consideration the polymorphism of some species, and the great variability, we can congratulate ourselves on having not more synonyms to record.
Type in the U.S. Nat. Museum, Washington.
Potamopyrgus egenus, Gould, sp.
Amnicola egena, Gould, Proc. Bost. Soc. Nat. Hist., vol. iii. (1848), p. 75. Amnicola gracilis, Gould, U.S. Expl. Exp., vol. xii. (1852), p. 127, figs. 151 a, b. Hydrobia egena, v. Martens, Crit. List. N.Z. Moll. (1873), p. 15. Bythinella egena, Hutton, Manual N.Z. Moll. (1880), p. 82.
I am indebted to Dr. W. H. Dall for the following information (in lit.): “The shell first described by Gould as Amnicola egena is imperfect. It was inadvertently called gracilis in the final report. It appears to be identical with specimens called gracilis in the collection, except that the type is brown and the others greenish.” Gould's diagnosis is copied in Hutton's Manual, and the type was found on Banks Peninsula.
This is undoubtedly a good species, but it seems to be rather rare. I have it in my collection from three localities only: Two specimens from Kaiwarra River, near Wellington; one from Nelson; and fourteen from Little River, Banks Peninsula. The graceful elongated shape, the convex whorls, and the last whorl amounting to half the length of the shell, distinguish it at once from the other species. No carinated or spinous forms are known. The dimensions given by Gould are: Length, 5 mm.; breadth, 2 ½ mm.; with five whorls. The dimensions of the Little River specimens, with five whorls, are 4 × 2 mm.; those of the Kaiwarra specimens, 4 × 1 ¾ mm. with six whorls, and 5 × 2 ¼ mm. with seven whorls; while the Nelson specimen measures 5 ½ × 2 ½ mm., and has six whorls. Thus my specimens must have slightly shorter whorls and a somewhat narrower spire than the type. In all specimens the last whorl is a trifle longer than half the axis of the shell.
Type in the U.S. Nat. Museum, Washington.

Potamopyrgus spelæus, Frauenfeld, sp.
Hydrobia spelæa, Frauenfeld, Verh. Zool. Bot. Gesell. Wien, vol. xiii. (1862), p. 1022; vol. xv., p. 526, pl. viii. Hydrobia spelæa, v. Martens, Crit. List. N.Z. Moll. (1873), p. 15. Bythinella spelæa, Hutton, Manual N.Z. Moll. (1880), p. 82.
This species was found together with P. reevei in moa-bone caves. A translation of the diagnosis is given by Hutton, but the dimensions are not quite those of Frauenfeld; the type is 3 mm. long by 1.6 mm. in breadth. This minute species is nearly allied to pupoides, Hutton, which, however, is mostly, but not always, smaller, and constantly of pupoid form.
The specimens in my collection I take to be almost typical were collected by Mr. A. Hamilton, now Director of the Colonial Museum, in the salt springs at Te Mahia, Hawke's Bay, and they show the same dimensions as the type specimen. A rather large form comes from the tidal part of the Wanganui River, and its dimensions vary from 3 × 1 ¾ mm. to 3½ × 2 mm. Very small specimens occur at Nelson, the largest measuring 2½ × 1¼ mm., the smallest 2 × 1 mm. The latter is the usual size of P. pupoides, but the Nelson examples are distinguished by much more convex whorls, deeper suture in consequence, and a more conoidal spire. Forms with ventricose body-whorl were collected by Mr. Chadwick on the upper Wanganui River, and in a cold mineral pool at Rotorua by Lady Frances Brown, a most enthusiastic collector. What I consider to be the same species are examples collected by Mr. Charles Cooper, of Auckland, in hot-spring water at Te Aroha. These specimens are very variable in shape, some approaching the type and measuring 3 × 1½ mm., others with a more inflated body-whorl show¼ mm. greater breadth; they all have only four whorls. This is the only instance known to me of a mollusc living in hot water in New Zealand, while it is well known that Neritina and Bithynia have been found in France living in water from 68° to 122° Fahr.
Type in the K.K. Hofmuseum, Vienna.
Potamopyrgus spelæus, Frfid., subsp. pupoides, Hutton.
Potamopyrgus pupoides, Hutton, Trans. N.Z. Inst., vol. xiv. (1882), p. 146, pl. i., figs. D-H. Potamopyrgus pupoides, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii. (1893), p. 620. Potamopyrgus pupoides, Suter, Journ. de Conch., vol. xli. (1893), p. 222.
I have typical specimens from brackish water in the Heathcote Estuary, near Christchurch, and they are so nearly allied to Frauenfeld's spelœus that I find it advisable to give P. pupoides only subspecific rank. It is distinguished from the species

by its cylindrical form, the much flatter whorls, and the less impressed suture. I have also specimens from Parua Bay, near Whangarei, and from the Onehunga Springs. Those from the latter locality are variable, some corresponding with the type, while others have more convex whorls, and approach the very small form of spelœus from Nelson.
Type in the Canterbury Museum, Christchurch.
Potamopyrgus subterraneus, n. sp.
Shell minute, subcylindrical, fragile, opaque-white, smooth. Spire pupoid; apex blunt; whorls 5, rather convex, the body-whorl more than half the axis; suture well impressed; mouth oval, oblique, peristome continuous; outer lip membranaceous, the specimen being apparently not quite full-grown; inner lip slightly callous, subvertical. Operculum not seen. Length, 2¾ mm.; breadth, 1¼ mm.
Hab. The only specimen was obtained by Mr. W. W. Smith, of Ashburton, by pumping water from a well 48 ft. deep.
It was alive when caught, but upon reaching me the animal was already decomposed. Mr. Smith very kindly presented the specimen to me in February, 1892. I did not describe it then because I hoped to get some specimens with the animal in sufficiently good condition for study. No other specimens, however, turned up. The species is exceedingly fragile, and distinguished, like pupoides, by its subcylindrical form. The nearest allies are egenus and spelœus, but it differs considerably from both. It most likely has been derived from the latter species.
Type in my collection.
Art. XVI.—Revision of the New Zealand Species of the Genus Isidora, with Description of a New Subspecies.
[Read before the Wellington Philosophical Society, September 7th, 1904.]
The first short list of species we find in von Marten's “Critical List” (1873), p. 15, consisting of three species: Physa variabilis, Gray; P. tabulata, Gould; and Limnæa (?) wilsoni, Tryon (a sinistral shell). The next list, in Hutton's Manual, is more extensive, comprising ten species: Physa wilsoni, Tryon; P. antipodea, Sow.; P. gibbosa, Gould; P. guyonensis,

T.-Woods; novæ-zelandiœ, Sow.; P. tabulata, Gould; P. variabilis, Gray; P. moesta, Ad.; P. lirata, T.-Woods; and P. cumingii, Ad. In vol. vii., P.L.S. N.S.W., Captain Hutton published a list of the fresh-water shells of New Zealand, in which the species of the genus were reduced to four—Aplexa antipoda, Sow.; A. tabulata, Gould; A. variabilis, Gray (= gibbosa, Sow. not Gould, guyonensis, T.-Woods, hochstetteri, Dkr.); and A. moesta, Ad. In 1885 Captain Hutton gave a list of the Limnæidæ in these Transactions, vol. xvii., enumerating Bulinus antipodeus, Sow.; B. variabihs, Gray (= gibbosus, Hutt. non Gould, novæ-zealandiæ, Sow., guyonensis, T.-Woods); B. tabulatus, Gould; and B. moesta, Adams (= lirata, T.-Woods), giving a description and figure of the dentition of the latter. The following species were omitted as not really inhabiting New Zealand: Limnæa wilsoni, Tryon, like Physa pyramidata, Sow., from Australia; Physa gibbosa, Gould, inhabiting New South Wales; and Physa cumingi, Ad., inhabiting Queensland.
In Fischer's Manual, p. 257, we find only two species recorded: Physa guyonensis and P. moesta. With my friend Mr. Charles Hedley, of Sydney, I published in 1893 a “Reference List of the New Zealand Land and Fresh-water Shells,” in which, for the species of Isidora, Hutton's latest classification was chiefly adopted, reducing, however, the species to three: Bulinus antipodeus, Sow.; B. variabilis, Gray (= guyonensis, T.-Woods, novœ-zelandiœ, Sow.); and B. tabulatus, Gould (= moesta, Adams, lirata, T.-Woods). Mr. Hedley added the following species, which for want of literature had escaped the notice of New Zealand conchologists: B. novœ-seelandiœ, Clessin; tenisoni, Clessin; coromandelicus, Dkr.; and hochstetteri, Dkr.
The following year I published, at the request of Mr. H. Crosse, a somewhat more extensive list in the Journ. de Conch., vol. xli., adhering still to three species of Isidora, and reducing Clessin's and Dunker's species to synonyms, as follows: Bullinus variabilis, Gray (= guyonensis, T.-Woods, novœ-zelandiœ, Sow., gibbosus, Hutt. non Gould, novœ-seelandiœ, Clessin); B. tabulatus, Gould (= moesta, H. Ad., lirata, T.-Woods, coromandelicus, Dkr., hochstetteri, Dkr.); and B. antipodeus, Sow.
I have studied now the description and copy of figure, kindly supplied to me by Mr. Hedley, of Physa novœseelandiœ, Clessin, and find it to be identical with Physa lessoni, E. A. Smith, an Australian shell, and it has therefore to be omitted from the list of New Zealand shells. Physa tenisoni, Clessin, is according to the figure a Limnœa, being dextral. Physa hochstetteri, Dunker, was mentioned by Hutton as a synonym of Aplexa variabilis, Gray (antea), but was never mentioned again in his later publications.

When working out the Mollusca collected by Mr. K. Lucas in New Zealand lakes, I had the same experience with Isidora as with Potamopyrgus: the lumping of species had been carried on too far. I do not wish to exonerate myself from blame, and I freely confess that never before have I made a careful study of our species of Isidora, as their great variability makes it extremely difficult to decide the limit of species and subspecies. A good collection of specimens from various localities, besides plenty of time and patience, is necessary for the successful study of these fresh-water molluscs. Specimens from over twenty localities were used to write the present revision, and I hope that it will form a sound basis to work upon.
Physa variabilis was the first species described by Gray (in “Dieffenbach's Travels,” vol. ii. (1843), p. 248). The very short diagnosis, unaccompanied by a figure, has in my opinion been the curse of New Zealand conchologists. From the many species judged to be synonyms of this unfortunate variabilis it can be gathered that no one ever knew what Gray's species is—perhaps not even Gray himself, for his diagnosis fits nearly all our species. So it has become a regular olla podrida: all the forms that did not fall under a recognised species were simply labelled “variabilis, Gray.” I have come to the conclusion that as long as we retain this species there is no possibility of classifying our various forms of Isidora correctly, and I reject it as insufficiently described, unfigured, and embracing perhaps several distinct species.
Genus Isidora, Ehrenberg (1831).
Synonyms: Diastropha, Gray (1840); Ameria, H. Adams (1861); Glyptophysa, Crosse (1872); Pyrgophysa, Crosse (1879); Physastra, Tapparone Canefri (1883).
Animal without the produced and reflected mantle-lobes of Physa; radula Limnæidian, approaching Planorbis rather than Limnœa; central tooth bicuspid, cusps rather blunt, base square; laterals tricuspid; marginals serrate. Laterals about 6–10, marginals about 25–33. Number of rows varying between 140 and 220.
Shell sinistral, resembling that of Physa, acuminated or gibbous, smooth or keeled; texture somewhat thick, covered with a deciduous epidermis; columella strong, often reflected, umbilicus sometimes very wide and deep.
Distribution: Australia, Tasmania, New Zealand, New Guinea, New Caledonia, Fiji, Tonga, Africa (north, north-east, west, and south), southern France, Spain, and all countries bordering the Mediterranean.
Ameria was proposed for Physœ with keeled whorls. The

distinction is untenable. Every gradation of keeling is observable in the Australian Isidorœ (Rev. A. H. Cooke).
Isidora tabulata, Gould (1848), sp.
Physa tabulata, Gould, Proc. Boston Soc. Nat. Hist., vol. ii. (1848), p. 214; U.S. Expl. Exp., vol. xii., p. 116, figs. 136 a, b. Physa tabulata, v. Martens, Crit. List N.Z. Moll. (1873), p. 15. Physa tabulata, Hutton, Man. N.Z. Moll. (1880), p. 30. Aplexa tabulata, Hutton, P.L.S. N.S.W., vol. vii., p. 67. Bulinus tabulatus, Hutton, Trans. N.Z. Inst., vol. xvii. (1885), p. 57. Physa tabulata, A. H. Cooke, P.Z.S. (1889), pp. 139, 140, fig. 4 (radula). Bulinus tabulatus, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii. (1893), p. 627. Bullinus tabulatus, Suter, Journ. de Conch., vol. xli. (1894), p. 233.
The diagnosis is to be found in Hutton's Manual. Fig. 1 represents the species after a copy from Reeve (Conch. Icon., vol. xix., Physa, fig. 17b), kindly supplied with several others by Mr. Hedley. Taken from the figure the dimensions are—Shell: length, 22 mm.; breadth, 16 mm. Aperture: length, 12 mm.; breadth, 8 mm. I think it to be a most useful thing to establish the following rationis to help in separating the various forms: i. Ratio between breadth and length of shell = 1: 1.4; ii. ratio between breadth and length of aperture = 1: 1.5; iii. ratio between length of aperture and length of shell = 1: 1.8. Hereafter I shall designate these rationis simply by i., ii., iii.
Hob. A mountain-stream, Bay of Islands (Drayton).
Type in the U.S. Nat. Museum, Washington.
I have not seen this species.
Isidora tabulata, Gould, subsp. moesta, H. Adams (1861), subsp. Physa moesta, H. Adams, P.Z.S. (1861), p. 144. Physa coromandelica, Dunker, Malac. Blätter (1862), p. 150. Physa moesta, Hutton, Man. N.Z. Moll. (1880), p. 31. Aplexa moesta, Hutton, P.L.S. N.S.W., vol. vii., p. 67. Bulinus moesta, Hutton, Trans. N.Z. Inst., vol. xvii., p. 57, pl. xii., fig. 12 (dentition). Bulinus moesta et coromandelica, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii., pp. 627, 628. Bullinus moesta et coromandelica, Suter, Journ. de Conch., vol. xli., p. 233.
The very short diagnosis is fortunately supplemented by a figure in Reeve's Conch. Icon., fig. 32, the outlines of which I here reproduce (fig. 2). The dimensions, taken from the figure, are—Shell: length, 17 mm.; breadth, 11 mm. Aperture: length, 10 mm.; breadth, 5 mm. The proportions are: i. = 1: 1.5; ii. = 1: 2; iii. = 1: 1.7.

I have examined and measured specimens in my collection from the following localities:—
(1.) Ditch near Lake Takapuna, Auckland. The specimens are fuscous, with a ferrugineous coating, shouldered, larger and more slender than the type. Two specimens measured gave length of shell 15 mm. and 17 mm., breadth 9 mm.; and the mean proportions were found to be—i. = 1: 1.8; ii. = 1: 2; iii. = 1: 1.7.
(2.) From Lake Takapuna, Auckland. Two specimens, dark-brown, strongly carinated, larger and a little less ventricose than the type. Shell: length, 12–15½ mm.; breadth, 7–9 mm. Mean rationis—i. = 1: 1.7; ii. = 1: 2; iii. = 1: 1.5.
I first tried to uphold I. coromandelica, Dunker, as a separate subspecies of I. tabulata, as the specimens of the above three localities approach the ratio between breadth and length of shell of I. coromandelica, which is 1: 2, but I soon found out that all intermediate forms, from the elongated coromandelica to the more ventricose moesta, are met with, and, as there seems to be no other character available to distingusih the two, I thought it advisable to make the former a synonym of the latter.
(3.) Wanganui (no exact locality). One adult and two not quite full-grown specimens were measured. The adult had—length, 17 mm.; breadth, 10 mm. All specimens are shouldered. The mean proportions are—i. = 1: 1.6; ii. = 1: 2.1; iii. = 1: 1.4.
(4.) Fresh-water stream, Parua Bay, Whangarei. All specimens are blackish-brown, two are distinctly shouldered, the other has smooth whorls. The columellar fold is very distinct. Three shells showed the length to vary from 9–15 mm., the breadth from 5½–9½ mm. The mean proportions are—1 = 1: 1.7; ii. = 1: 1.9; iii. = 1: 1.6.
(5.) Waikato River, near Huntly. Shells of good size, chestnut-colour, some with a ferrugineous coating; young specimens are strongly carinated, in adult specimens the carina is reduced to a slight angulation on the last whorl. Columella strongly twisted. Four shells were measured, the length being 12–15 mm.; breadth, 8–11 mm. The mean proportions were—i. = 1: 1.4; ii. = 1: 1.9; iii. = 1: 1.5.
(6.) Chatham Islands. One specimen only, of yellowish-brown colour, with four shouldered whorls. This is a slender form. Length, 8 mm.; breadth, 4½ mm. Rationis—1 = 1: 1.8; ii. = 1: 2; iii. = 1: 1.6.
The result of measuring twenty-eight specimens is the following: Ratio i.—variability, 1: 1.4 to 1: 1.8; mean, 1: 1.65. Ratio ii.—variability, 1: 1.9 to 1: 2.3; mean, 1: 2. Ratio iii.—variability, 1: 1.4 to 1: 1.7; mean, 1: 1.5. It shows that the shells of this subspecies are a little more slender and the spire

is somewhat higher in proportion to the length of the aperture than in the type specimen figured.
Type in the British Museum (?).
Isidora hochstetteri, Dunker (1862), sp.
Physa hochstetteri, Dunker, Malac. Blatter, vol. ix. (1862), p. 150. Physa guyonensis, T.-Woods, P.L.S. N.S.W., vol. iii. (1879), p. 138, pl. xiii., fig. 4. Physa guyonensis, Hutton, Man. N.Z. Moll., p. 30. Physa guyonensis et hochstetteri, Hutton, P.L.S. N.S.W., vol. vii., p. 67. Bulinus guyonensis et hochstetteri, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii., pp. 627, 628. Bullinus guyonensis et hochstetteri, Suter, Journ. de Conch., vol. xli., pp. 232, 233.
The diagnosis of guyonensis is in Hutton's Manual. The accompanying fig. 3 is drawn from a tracing of the figure given by Tenison-Woods. Captain Hutton mentions (Manual) that Dr. Dohrn determined specimens from the same locality (Lake Guyon) as P. hochstetteri, Dkr. The diagnosis agrees in the main points with that of T.-Woods, and Dunker particularly mentions the deep suture and the amplitude of the body-whorl. I do not hesitate to accept the determination of the distinguished conchologist, Dr. Dohrn. The dimensions of P. hochstetteri are—Height, 17 mm.; breadth, 9 mm. For P. guyonensis the dimensions given by T.-Woods are—Height of shell, 15 mm.; breadth, 7½ mm.: aperture—height, 9 mm.; breadth, 5 mm. I have not seen specimens from Lake Guyon, but I have shells from Lake Nga-tu, Kaitaia, which I consider to belong to this species, although they are somewhat shorter and more ventricose than the type; otherwise they agree with the diagnosis. Especially the deep suture is characteristic, no other of our species showing this character in such a marked degree. All my examples are “dead shells” of a light-brown colour.
For I. guyonensis the rationis are—i. = 1: 2; ii. = 1: 1.8; iii. = 1: 1.7. For hochstetteri—i. = 1: 1.9. I measured four of my specimens. The length of the shell varies from 13½–14½ mm., the breadth from 8–9 mm. Aperture—length, 8–9 mm.; breadth, 4–5 mm. The mean proportions are—i. = 1: 1.6; ii. = 1: 1.9; iii. = 1: 1.6.
Hab. The only localities known to me are Lake Guyon, Nelson, and Lake Nga-tu, in the high north.
Type in the K.K. Hofmuseum, Vienna (?).

Isidora novæ-zelandiœ, Sowerby (1873), sp.
Physa novœ-zelandiœ, Sowerby, Reeve, Conch. Icon., vol. xix., Physa, sp. 29 (1873). Physa novœ-zelandiœ, Hutton, Man. N.Z. Moll., p. 30. Bulinus novœ-zelandiœ, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii., p. 627. Bullinus novœ-zelandiœ, Suter, Journ. de Conch., vol. xli., p. 232.
The diagnosis of this species is also contained in Hutton's Manual, and I reproduce here a figure (4) from a tracing of fig. 29b in Conch. Icon. Taken from the figure the dimensions are—Shell: length, 20½ mm.; breadth, 12¼ mm. Aperture: length, 15 mm.; breadth, 6 mm. Proportions—i. = 1: 1.7; ii. = 1: 2.5; iii. = 1: 1.4.
I have only two specimens from the North Island, exact locality unknown. One adult specimen has smooth whorls, the other, not full-grown, is shouldered. The dimensions are—Shell: length, 17 mm.; breadth, 12 mm. Aperture: length, 14 mm.; breadth, 6½ mm. Shell: length, 14 mm.; breadth, 9½ mm. Aperture: length, 10 mm.; breadth, 4½ mm. The mean proportions—i. = 1: 1.5; ii. = 1: 2.2; iii. = 1: 1.3. The adult specimen is of chestnut-colour, the young horny-olive. The spire is shorter and the last whorl not quite so broad posteriorly, nor so flat at the periphery, as the figure of the type indicates.
Type in Mr. Sowerby's cabinet (?).
Isidora antipodea, Sowerby (1873), sp.
Physa antipodea, Sowerby, Reeve, Conch. Icon., vol. xix., Physa, sp. 37 (1873). Physa antipodea, Hutton, Man. N.Z. Moll., p. 30. Aplexa antipoda, Hutton, P.L.S. N.S.W., vol. vii., p. 67. Bulinus antipodeus, Hutton, Trans. N.Z. Inst., vol. xvii., p. 56. Bulinus antipodeus, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii., p. 627. Bullinus antipodeus, Suter, Journ. de Conch., vol. xli., p. 233.
Diagnosis in Hutton's Manual. I reproduce here (fig. 5) the outlines of the species after Reeve. This seems, as far as our scanty knowledge goes, to be the only one of our species that has constantly smooth whorls. The dimensions taken from the figure are—Shell: length, 22 mm.; breadth, 12 mm. Aperture: length, 13 mm.; breadth, 6½ mm. Rationis—i. = 1: 1.8; ii. = 1: 2; iii. = 1: 1.7.
This species is represented in my collection by two specimens from Lake Wakatipu, and the dimensions are—Shell: length, 17 mm.; breadth, 10½ mm. Aperture: length, 11 mm.; breadth, 5½ mm. Shell: length, 18½ mm.; breadth, 11 mm. Aperture: length, 12 mm.; breadth,

6 mm. Mean rationis—i. = 1: 1.7; ii. = 1: 2; iii. = 1: 1.5. These specimens are slightly more ventricose and have the aperture in proportion shorter than the type.
Hab. Lake Hayes, Otago; near Napier; Lake Wakatipu.
Type in Mr. Sowerby's cabinet (?).
Isidora lirata, Tenison-Woods (1879), sp.
Physa lirata, T.-Woods, P.L.S. N.S.W., vol. iii. (1879), p. 138, pl. xiii., fig. 6. Physa lirata, Hutton, Man. N.Z. Moll., p. 31; Trans. N.Z. Inst., vol. xvii., p. 57. Bulinus liratus, Hedley and Suter, P.L.S. N.S.W. (2), vol. vii., p. 627. Bullinus liratus, Suter, Journ. de Conch., vol. xli., p. 233.
The diagnosis is also in Hutton's Manual, and the accompanying fig. 6 is reproduced from T.-Wood's figure. Fig. 6a shows a few teeth of the radula: the rhachidian tooth is bicuspid, the laterals tricuspid, the following transition teeth with four and more denticles, and the marginals are elongate and serrate.
This is one of the best-characterized species; the fine spiral lirae and the anteriorly produced lip distinguish it at once from all the other New Zealand species. The dimensions given by T.-Woods are—Shell: length, 10 mm.; breadth, 5 mm. Aperture: length, 5 mm.; breadth, 3 mm. The proportions are—i. = 1: 2; ii. = 1: 1.7; iii. = 1: 2: but taken from the figure we get—i. = 1: 1.9; ii. = 1: 2.2; iii. = 1: 1.5.
In my collection the species is represented from twelve localities: Rivers Heathcote and Avon, creek in St. Albans, pond in Fendalton, all near Christchurch; Wellington; Greymouth; Pelorus River; pond near Lake St. John; Toko; Lakes Virginia and Westmere, near Wanganui; and North Island, exact locality unknown. The specimens vary a great deal in size and proportions, but the main features mentioned above are always present. The colour is usually light-horny, the whorls are sometimes smooth but mostly strongly carinated, and the carina usually adorned with short light-brown bristles, a character only met with in this species.
I measured altogether thirty-one specimens and found the rationis to vary—i., from 1: 1.6 to 1: 1.9; ii., from 1: 1.6 to 1: 2.4; iii., from 1: 1.3 to 1: 1.7. The means are as follows—i. = 1: 1.7; ii. = 1: 1.9; iii. = 1: 1.5.
Type where ?

Isidora lirata, T. Woods, subsp. conferta, n. subsp.
Shell globosely ovate, sinistral, corneous-translucent, thin, almost imperforate, the columellar reflexion leaving only a narrow chink. The distant lines of growth are rather regular, especially on the upper whorls, and under the lens fine regular and close spiral liræ are visible. The colour varies from very light horny to light olive-brown. The spire is short, about one-third the length of the shell. The pullus is acuminate, of darker colour, consisting of two whorls with fine incremental striation. There are four whorls, either strongly shouldered or having only a posterior angle, which usually gets lost on the body-whorl; the latter is forming the greater part of the shell. The base is convex, the suture impressed. Aperture vertical, elongately oval, acuminate above and produced anteriorly. The outer lip is regularly rounded, sharp, thin; the inner lip is twisted, forming a distinct fold near the axis of the shell; not much reflexed. Fig. 7 represents a distinctly shouldered form, and fig. 8 with only a slight angulation on the whorls.
This subspecies is distinguished from the species by its ventricose form and the much broader aperture, approaching T. tabulata, Gould. I have specimens from two localities:—
(1.) From swamps near Otorohanga, King-country. These are strongly keeled (fig. 7). Three specimens were measured—Shell: length, 11 mm.; breadth, 7 mm. Aperture: length, 7 mm.; breadth, 4 mm. Shell: length, 9½ mm.; breadth, 6½ mm. Aperture: length, 6½ mm.; breadth, 3½ mm. Shell: length, 9½ mm.; breadth, 6 mm. Aperture: length, 6 mm.; breadth, 3¼ mm. The mean rationis are—i. = 1: 1.6; ii. = 1: 1.7; iii. = 1: 1.6.
(2.) From Wairau River, south of Birch Hill Station, Nelson. These specimens are much lighter in colour, the upper whorls slightly angled (fig. 8). Three specimens were measured—Shell: length, 11–12 mm.; breadth, 7–7½ mm. Aperture: length, 8–8½ mm.; breadth, 4½ mm. The mean proportions are—i. = 1: 1.6; ii. = 1: 1.9; iii. = 1: 1.4. The means for the two localities are—i. = 1: 1.6; ii. = 1: 1.8; iii. = 1: 1.5.
Type in my collection.
To repeat, the following species are omitted:—
(1.) Physa variabilis, Gray, insufficiently described and unfigured.
(2.) Physa gibbosa, Gould, inhabits Australia.

(3.) Physa cumingi, H. Ad., inhabits Australia.
(4.) Physa wilsoni, Tryon, inhabits Australia; perhaps pyramidata, Sow.
(5.) Physa novœseelandiœ, Clessin = P. lessoni, E. A. Smith, Australia.
Art. XVII.—Some, New Species of New Zealand Marine Shells, together with Remarks on some Non-marine Species, and some Additions to the “Index Faunœ.”
[Read before the Auckland Institute, 27th February, 1905.]
Plates IX and X.
New Species of New Zealand Marine Shells.
Turbo (Lunella) radina (= delicate), n. sp. Fig. 1, a, b.
Shell flat-topped, whorls 3½, of which the slightly depressed, white, smooth protoconch claims 1½, the next half-whorl, which is slightly rounded, is pale-green, the last is pale-brown. In the neanic stage the sharply angled periphery shows small white projections increasing in size until each assumes the form of a blunt hollow spine, having a dark-grey patch in front of it. The upper surface of the body-whorl slopes sinuously to the keeled periphery, below which are two similar but less developed keels; a single but obscure keel surrounds the deep umbilicus. The entire shell is irregularly but closely radiately striate, the striæ on the upper surface sloping backward. Suture well marked, aperture not continuous, circular, interior iridescent with a white margin inside the aperture. Columella vertical, curved, white, margin reflexed, outer lip thin. Height, 1½ mm.; breadth, 2½ mm. The operculum is characteristic.
Hob. Takapuna.
Type in my collection.
The colour and size of this shell agree with that of half a dozen specimens in my collection, also with a living specimen found at Whangarei by Mr. C. Cooper.
Astralium pyramidale, n. sp. Fig. 2, a.
Shell a pyramid with straight sides, brown above, marbled with green on the base; protoconch obscured by coralline growth, whorls 4, flat in outline but heavily ribbed, the body-whorl having twelve ribs which slope forwards from the suture towards the periphery and are crossed almost at right angles by numerous

squamose growth-lines; each rib ends with a flat hollow tooth projecting beyond the periphery; suture only marked by the serrated periphery, as in A. heliotropium. A rounded thread runs round the entire shell about one-third above the periphery. The squamose base is slightly rounded, and has six rounded threads between the outside edge and the columella; the second from the outside is very prominent, the fifth is somewhat so, and both persist into the aperture beneath the nacre. Columella white, vertical, the inside rounded and coated with a layer of dull nacre; there is a very slight umbilical depression. Animal and operculum unknown. Height, 10 mm.; breadth, 13 mm.
Hab. Takapuna.
Type in my collection.
N.B.—This shell does not resemble the young of any of our Astralium.
Rissoia vulgaris (= common), n. sp. Fig. 3.
Shell pale-horny, imperforate, of 4½ well-rounded deeply sutured whorls, the sutures channelled, protoconch small, shining. The entire shell is longitudinally, finely, diagonally striate. Aperture ovate anteriorly, lip thin, columella vertical, slightly reflexed and continued on the body-whorl by a slight callous line. Height, 2 mm.; breadth, 1¼ mm. Animal and operculum unknown.
Hab. Waipipi.
Type in may collection.
Rissoia micans (= shining), n. sp. Fig. 4.
Shell minute, imperforate, dark rich golden-brown, highly polished; whorls 4, rounded, divided by deep sutures, no sculpture. Columella vertical, arched, dark-brown, slightly reflexed; aperture circular, not continuous, lip simple, very slightly expanded, and semitransparent milk-white in colour. Height, 1¼ mm.; breadth, 1 mm. Animal and operculum unknown.
Hab. Takapuna.
Type in my collection.
Rissoia zosterophila, n. sp. Fig. 5, a, b.
Shell thin, imperforate, with brightly contrasting colours, the upper half of each whorl milk-white, the lower half dark redbrown; the base is horny, thus leaving the red-brown colour as a sub-peripheral band on the body-whorl; this band forks lengthwise about half a turn from the aperture, on reaching the inner edge of which it stops abruptly in most full-grown specimens. Whorls 6½, scarcely rounded, the smooth elevated protoconch taking up two of these. Periphery bluntly angled, a similar projection

being noticed in the outline near the columella. Aperture angled behind, rounded in front, continuous, thin-edged, the edge slightly expanded. Columella nearly vertical, interior transparent. Operculum quite colourless and presenting a malleated appearance on the internal surface. Since deciding upon describing these shells I have had no opportunity of obtaining the animal alive. Height, 2½ mm.; breadth, 1 mm.
Hab. Devonport.
This shell is labelled R. annulata in the Auckland Museum, but is not that species as named by Professor Hutton.
Rissoia carnosa (= flesh-coloured), n. sp. Fig. 6.
Shell thin, imperforate, flesh-brown, with a cream-coloured band above the periphery, some specimens being a darker brown without the band, others having the last whorl entirely creamcoloured. Whorls 5½ (of which two form the smooth domeshaped dark protoconch), slightly rounded, the body-whorl more so, and having about twenty straight longitudinal ribs crossed by about twenty-five spiral liræ, which diminish in strength as they ascend to the apex, while the ribs are stronger on the upper whorls and in some specimens die away on the body-whorl. The shell is so thin that the sculpture is plainly visible when looked at through the aperture. Suture well marked; aperture auriform, not thickened anywhere, not continuous. Animal and operculum unknown. Height, 2¼ mm.; breadth,¾ mm.
Hab. Takapuna.
Type in my collection.
Rissoia candidissima (= very white), n. sp. Fig. 7.
Shell thin, imperforate, semitransparent-white when fresh. Whorls 4½, of which two form the smooth elevated protoconch, very round, with about twelve distant longitudinal ribs on the body-whorl crossed by twelve spiral lirae. Suture deep, aperture rounded but not continuous, slightly thickened. Animal and operculum unknown. Height, 2 mm.; breadth, 1 mm.
Hab. Takapuna.
Type in my collection.
Rissoina (Eatoniella) limbata, Hutt. Fig. 8, a.
I give a drawing of the radula × 600 and of the operculum of this species, which was originally described by Professor Hutton as a Cingula (the animal being unknown), and then found its way into Phasianella (see P. Mal. Soc., iii., p. 8). I also examined the dentition and operculum of transparent-pink and pellucid-white specimens, finding them the same as the type.

Rissoina (Eatoniella) olivacea, Hutt. Fig. 9, a.
I examined the radulæ of many typical dark-green and also brown specimens of this shell, further also of the purple-black shells with white sutural band called R. annulata, Hutt. I find radula and operculum identical in all. I consider therefore that R. annulata intergrades with R. olivacea. With Professor Hutton's concurrence I propose for R. annulata varietal rank, thus: Rissoina (Eatoniella) olivacea, Hutt. (1882); Rissoina (Eatoniella) var. annulata, Hutt. (1884).
Rissoina agrestis (= clumsy), n. sp. Fig. 10, a, b,
Shell solid, imperforate, dull, of a purple-brown shading to cream-colour on the body-whorl. Whorls 5½, including a 2½-whorled domed striato-punctate protoconch with a well-defined convex channel at the suture. The three lower whorls are longitudinally, slightly diagonally, rudely ribbed, the channel being ribbed independently. Aperture somewhat angled behind and rounded in front, very thick. Columella vertical, marked by a white shining stripe. Animal unknown. Length, 1½ mm.; breadth, ⅔ mm.
Hab. Takapuna.
Type in my collection.
Very like the illustration of Rissoa impressa, Hutt.
Eulimella coena (= ordinary), n. sp. Fig. 11, a.
Shell elongated, slender, white, smooth, imperforate, whorls 7, flat with a blunt heterostrophe apex. Sculpture none, suture slight, aperture ovate anteriorly, columella vertical and slightly twisted. Height, 2½ mm.; breadth, 1 mm. Animal and operculum unknown.
Hab. Takapuna.
Type in my collection.
Corbula pura (= clean), n. sp. Fig. 12, a.
Shell white, thin, tinged with pink before and behind the umbones, which are one-third from the posterior rounded end. The fold is sharp, its outline concave, its anterior end being far in advance of the anterior dorsal margin. Prodissoconch quadrate, plain, slightly eroded, the entire shell lightly but clearly and regularly both concentrically and radiately striate, left valve very slightly the larger. Hinge teeth, sinus, and scars, &c., normal. Height, 5 mm.; breadth, 10 mm.; depth from valve to valve, 3 mm.
Hab. Dredged in Rangitoto Channel.
Type in my collection.

Standella aequalis (= uniform), n. sp. Fig. 13, a.
Shell a true oval, umbones two-fifths from the anterior end; the entire shell, prodissoconch included, lightly and irregularly concentrically striate. Hinge teeth, sinus, scars, &c., normal. Height, 17 mm.; breadth, 27 mm.; depth of single right valve, 6 mm.
Hab. Kauri Point.
Type in my collection.
Remarks on some Non-marine Shells.
Thaumatodon varicosa, Pfeiffer.
All the shells collected near Waiuku have two teeth, one on the body-whorl and another at the base of the columella margin. This is the case with both adult and juvenile specimens.
Ptycodon pseudoleiodon, Suter.
This shell is found in Waiuku in the adult stage normal and also with combinations, 3 . 1 . 7; 3 . 1 . 10; 3 . 1 . 12, &c.
Phrixgnathus phrynia, Hutton.
This and other shells are found in Waiuku with callous patches indicating the development or loss of throat teeth.
Laoma marina, Hutton.
This occurs (adult) with the throat teeth of L. nerissa, Hutton.
Shells to be added to Fauna List.
The following shells in my collection should be added to or reinstated in the list of our fauna:—
Sigaretus undulatus, Hutt. Cape Maria van Diemen.
Ancilla depressa, Sow. Orua Bay.
Terebra venosa, identified by Mr. Suter. Cape Maria van Diemen and Port Waikato.
Pyrgulina rugata, Hutt. Takapuna.
Leuconopsis inermis, Hed. Takapuna.
Pholadidea spathulata, Sow. Narrow Neck.
Venericardia amabilis, Desh. Cape Maria van Diemen.
Venerupis carditoides, Lam. Takapuna.
Philene teres, Hed. Cheltenham Beach, Auckland.
Also the following two shells, which have been omitted from the “Index Faunæ”:—
Venus (Gomphina) maorum, Smith, J., of Mal. 1902, ix., pt. 4.
Unio (Diplodon) websteri, Simpson, “Nautilus,” July, 1902.

Art. XVIII.—Some Earthworms from the North Island of New Zealand.
[Read before the Otago Institute, 13th September, 1904.]
Plate XI.
All the earthworms, with two exceptions,* that have hitherto been ascribed with certainty to New Zealand have, as a matter of fact, been collected in the South Island, and chiefly from Canterbury and Otago; but during the last year or two I have received numerous specimens from various localities in the North Island, and in the early months of the present year I received from Mr. Elsdon Best a collection of worms made at Ruatahuna, the interest in which is enhanced by the fact that several of the species were in former times used as an article of diet by the Maoris, and are referred to under their Maori names in his article “The Food-products of Tuhoeland” in these Transactions.† I gladly take this opportunity of thanking Mr. Best for his kindness in taking the trouble, at my request, to collect, preserve, and forward to me these extremely interesting species; for not only is this the first occasion in which earthworms have been recorded as being eaten by man of any race, but, from a zoological point of view, they introduce us to a family of earthworms hitherto scarcely represented in New Zealand. To Mr. Charles Cooper, of Auckland, my thanks are also due, for it was at his suggestion that I put myself into communication with Mr. Best, whose article I had not at that time read, and was therefore ignorant of the fact that the Maori esteemed the earthworm as an article of diet. To several other correspondents I herewith offer thanks for forwarding to me specimens, including Captain Hutton, Professor H. B. Kirk, and Mr. H. Suter.
I have already written a detailed account of these new species, and have sent it to the Zoological Society for publication‡; nevertheless it seems desirable to place on record in these Transactions all new species of animals described from New Zealand, even if a certain amount of duplication of articles
[Footnote] * These two exceptions are Maoridrilus plumbeus and Microscolex monticola. both described by Beddard, from Mount Pirongea, near Auckland.
[Footnote] † Trans. N.Z. Inst., xxxv., p. 45.
[Footnote] ‡ Proc. Zool. Soc., 1904.

results. Here and now it is only necessary to present a list of the new species, but I cannot let the opportunity pass by without emphasizing the fact that the predominant earthworms in the North Island belong to a subfamily quite different from that to which the predominant earthworms of the South Island belong, and heretofore believed to be characteristic of New Zealand as a whole.
The South Island earthworms (extending up to and including Stephen Island in Cook Strait) belong to the subfamily Acanthodrilinœ; on the other hand, the commonest earthworms in the North Island belong to the subfamily Megascolecinœ, which is characteristic of Tasmania and Australia.
The problem of distribution presented to us renders our former ideas on the subject as to the relation of our fauna to that of Australia somewhat confusing: and I do not at present propose to discuss it till I have worked out the line along which the two subfamilies came into contact in New Zealand. I shall be extremely grateful for any earthworms from Nelson and Marlborough, as well as from the southern parts of the North Island, for we are very deficient in knowledge as to the fauna in these parts of the country.
List of New Species.
Fam. Megascolecid æ
Subfam. Acanthodrilin æ.
1. Maoridrilus mauianus, Benham.
Loc. Auckland.
A single individual. Collected by Mr. Suter.
2. Octochætus michaelseni, Banham.
Loc. Wellington.
A single specimen. Collected by Captain Hutton.
Dinodriloides, Benham (1904).
3. Dinodriloides beddardi, Benham.
Loc. Auckland.
A single individual. Collected by Mr. Suter.
4. Rhododrilus edulis, Benham.
Loc. Ruatahuna.
Two individuals. Collected by Mr. Elsdon Best. This worm is known to the natives of Tuhoeland as tarao (= wharu, and perhaps huharu); it appears to be common, and was used as food.*
[Footnote] * I have added diagrams to illustrate the specific characters of this and the following species, for comparison with other species of Rhododrilus described in the present volume.

5. Rhododrilus besti, Benham.
Loc. Ruatahuna.
A single individual. Collected by Mr. Elsdon Best.
This is the tokerangi of the natives, and is not included in Mr. Best's list of edible species. He writes me that it is “found on tracks, &c., in the morning, generally after a wet night.”
Subfam. Megascolecin æ.
Tokea, Benham (1904).
6. Tokea esculenta, Benham.
Loc. Ruatahuna.
Collected by Mr. Elsdon Best, in stony places: it is termed Kurekure by the natives, who eat it.
7. Tokea sapida, Benham.
Loc. Ruatahuna.
Collected by Mr. Best. This is also an edible species, and is not distinguished by the Maori from the preceding species, the same name, kurekure, being used for both. It was one of the most prized kinds, and was reserved as food for the chiefs.
8. Tokea ureweræ, Benham.
Loc. Ruatahuna.
Collected by Mr. Best. It is the pokotea of the Maoris, who included it in their bill-of-fare.
9. Tokea huttoni, Benham.
Loc. Whangarei.
One individual. Collected by Captain Hutton.
10. Tokea suteri, Benham.
Loc. Auckland.
One individual. Collected by Mr. Suter.
11. Tokea kirki, Benham.
Loc. Ohaeawai.
Five specimens. Collected by Professor H. B. Kirk.
12. Tokea maorica, Benham.
Loc. Auckland: Waitakerei Bush and Nikau-palm Bush, near Auckland.
Eight individuals. Collected by Mr. Suter.
It will be seen that out of a collection containing twelve new species, only two belong to the predominant genera of the South Island—viz., Maoridrilus and Octochœtus. The genus Rhododrilus is represented by one species in the South Island, one species in the Chatham, and two species (described in the present volume) in the southern islands.

The new genus Dinodriloides is allied to Dinodrilus of the South Island, of which only one species is at present known. There are five species belonging to the subfamily Acanthodrilinœ: the remaining seven belong to a new genus, allied to the Cryptodriline series of Australia.
The genus Tokea (from the Maori toke, an earthworm) is very widely distributed over the North Island, as will be seen from the varied localities at which it has been collected—from Ohaeawai at the north to Ruatahuna in the south-east portion of the island.* It is probably the commonest earthworm in these parts, as two species occur in and around Auckland, three species at Ruatahuna, and from the majority of the other localities no other genus has been received. It is, so far, unknown in the South Island.
To this list of new species from the North Island we must add Beddard's species, Maoridrilus plumbeus and Microscolex monticola, from Mount Pirongea, described in 1895. But before that date Schmarda described “Hypogœon orthostichon,” from “Mount Wellington, New Zealand.” This worm is now placed, by those who have re-examined the specimen, in the genus Notoscolex, a thoroughly characteristic Australian genus; and doubt has been thrown on the accuracy of Schmarda's statement that it was collected in New Zealand—firstly, because no representative of that genus had hitherto been found here, and secondly, because there is no “Mount Wellington” of conspicuous size in New Zealand. It has been suggested that the well-known mountain of that name near Hobart was the real source whence Schmarda obtained the worm, and that the words “New Zealand” were due to a lapsus calami†; and I went so far as to refer to it, in a note read before the Australasian Association for the Advancement of Science at Hobart in 1902, as a “neglected Tasmanian earthworm.” But these two reasons for doubt may now be laid aside—for, firstly, the genus Tokea is closely allied to Notoscolex, and the differences are such that unless particular attention were paid to the points of difference they might readily be overlooked, and the description is insufficient to decide the question; it is possible, then, that Schmarda's worm may indeed belong to this new genus Tokea. Then, secondly, in reply to a query from me, Professor Kirk informs me that “Mount Wellington is the name of a small volcanic cone just outside Auckland, now under cultivation”; so that the suggestion of a geographical error seems to have
[Footnote] * I have recently received species of this genus from Little Barrier Island.
[Footnote] † Hutton, “Catalogue of the Worms of New Zealand,” Trans. N.Z. Inst., xi., footnote, p. 317.

been unnecessary. It is probable that the point will never be decided, and, as Schmarda's worm no longer forms an exception to the zoo-geographical problem, its decision is now of little importance.
Explanation of Plate XI.
The illustrations of the anatomy of the earthworms described in this article are purely diagrammatic, indicating only the segmental position of the various organs, the worm being supposed to be slit up along the dorsal line and the body-wall pinned aside.
A group of three diagrams refers to each worm herein described. The left-hand diagram in each of the groups referring to a species represents the external features. The location of the various genital pores is represented as round black dots (if on a papilla this is left white), the clitellum is obliquely shaded, the tubercula pubertatis are vertically shaded.
In addition, the arrangement of the chætæ—labelled a, b, c, d—is indicated in segments 5 to 23 on one side; they are omitted on the other side for clearness' sake. The true relative spacing of the chætæ is shown.
The position of the nephridiopores is indicated by the small circles on one side of the figure.
The middle figure represents the alimentary canal and so much of the vascular system as is diagnostic. The latter is black. The gizzard is indicated by vertical shading, the œsophageal glands by more or less horizontal lines The intestine is not represented as being constricted, which is, however, the case in most worms.
The right-hand figure shows the reproductive system. The gonads are in black. The sperm-sacs are dotted The sac with penial chætæ when present is indicated, and the muscular duct of the spermiducal gland is transversely striped. The transverse muscles in the 18th segment are shown.
No attempt is made to give the relative sizes of the worms or of the various organs.
Art. XIX.—On the Oligochœta from the Southern Islands of the New Zealand Region:
[Read before the Otago Institute, 13th September, 1904.]
Plates XII. and XIII.
In the winter of last year (July, 1903) Dr. L. Cockayne paid a visit to the southern islands on the Government steamer “Hinemoa,” and while collecting plants was good enough to collect earthworms, which he kindly handed over to me. The following is the list of worms described in the present paper, all but the last being collected during this expedition:—

Fam. Megascolecid æ.
Subfam. Acanthodrilin æ.
(1.) Notiodrilus aucklandicus, Benham.
(2.) Notiodrilus campbellianus, n. sp.
(3.) Rhododrilus cockayni, n. sp.
Fam. Enchytræid Æ.
(5.) Marionina antipodum, n. sp.
(6.) Enchytrœus albidus, Henle.
(7.) Lumbricillus macquariensis, n. sp.
We already know two species of Notiodrilus from these southern islands—viz., N. macquariensis, Beddard, originally described some years ago, and more recently studied and illustrated by myself,* and N. aucklandicus, described by myself in 1902. Hitherto these two worms have been the only Oligochæta recorded from this region, and the other islands have been unexplored for this branch of their fauna. The present contribution thus adds a third species of Notiodrilus,† as well as two new species of Rhododrilus, which is endemic in New Zealand, and, unlike some of our other genera, is represented alike in the North and South Islands, as well as in the Chathams. The presence, too, of Enchytræids in these southern islands is of interest, as species of both Marionina and Lumbricillus have been recorded from South Georgia and Tierra del Fuego, while the same species of Enchytrœus has been met with all over the globe.
The Oligochœtal fauna, so far as are known, may here be summarised:—
Antipodes Island:—
Notiodrilus aucklandicus.
Marionina antipodum.
Lord Auckland Isles:—
Notiodrilus aucklandicus.
Rhododrilus cockayni.
Rhododrilus leptomerus.
Campbell Islands:—
Notiodrilus aucklandicus.
N. campbellianus.
Rhododrilus cockayni.
Enchytrœus albidus.
[Footnote] * Trans. N.Z. Inst, 1900 and 1902.
[Footnote] † It may be that these three forms are local varieties of one and the same species.

Macquarie Islands:—
Notiodrilus macquariensis.
Enchytrœus albidus.
Luwbricillus macquariensis.
In the present paper I confine myself to detailing the characters of the new species, and will postpone a consideration of the bearing that the facts of geographical distribution have in regard to the previous extension of the New Zealand land surface. I have so much new material waiting investigation that it is desirable to gather together these facts in a general paper at a later date.
Dr. Michaelsen has recently* pointed out that Notiodrilus kerguelarum is to be found not only on dry land but also on the sea-shore within reach of the sea-spray, in company with typically littoral Oligochæta such as Enchytrœus albidus—or, to use his term, it is “euryhaline.” It is worthy of note, therefore, that in the collection made by Dr. Cockayne there is further evidence in support of this statement, for in the bottle containing N. campbellianus there are Enchytrœus albidus, a few Polychæta, and a Nemertine. After reading Michaelsen's remarks I wrote to Dr. Cockayne for further details as to the “stations” at which these worms had been obtained, and he writes me that “all the Campbell Island worms were collected near one another on the stony shore of Perseverance Harbour, just above high-water mark. I collected no other worms on Campbell Island. It is possible that sea-spray may reach any place at which the Adam Island worms were collected, but the soil cannot be termed brackish in any degree. As for Ewing Island worms, these were collected on or near the shore. On the other hand, the ‘peat bog’ in which the worms from Antipodes Island were collected is quite out of the influence of the sea-spray.”
Notiodrilus aucklandicus.
The single specimen from the Antipodes Island is, curiously enough, truncated anteriorly, and has lost four anterior segments; this injury, however, is entirely healed up, but no new prostomium has yet been formed, nor have the anterior segments been regenerated, for the porophores, instead of being on segments 17 and 19, occur on the 13th and 15th. Granting this loss of four segments, the organs occupy the normal position.
The colour differs somewhat from the type in being yellowish instead of grey posterior to the clitellum. This is not due to the preservative, for formaline was used in both cases, but is due to colour of intestinal contents.
[Footnote] * Die Geograph. Verbreit. d. Oligochœten, 1903.

The specimen is smaller than the type, being only 55 mm. × 2 mm.
It was collected in “bog land, in peat of Pleurophyllum.” This “Pleurophyllum meadow” is described in detail by Dr. Cockayne in his interesting article “A Botanical Excursion during Midwinter to the Southern Islands of New Zealand”*: “Then the leaves of the past year lie rotting upon the surface of the soil, while their bases form great decayed masses many centimeters in thickness round the leaf-bases of the young leaves of Pleurophyllum criniferum or P. hookeri. In these sheaths of decaying leaves considerable numbers of earthworms were found, and they appear also to be fairly numerous in the peat itself.”
Those from the Campbell Islands were obtained from the “roots of plants” in some number, mature and immature. Their colour is pinkish-brown anteriorly, instead of sienna-brown as in the type. The only noticeable difference anatomically from the type is the absence of definite œsophageal glands, though the gut-wall is thicker and dilated in segments 13, 14, and 15. Also the diverticula of the spermathecæ arise from the duct free from the body-wall.
From Adam Island (one of the Lord Auckland Group) I received nine specimens, of which three are immature. These show no sensible variation in colour.
Notiodrilus campbellianus, n. sp.
Three specimens were obtained, of which two are immature; all are ill-preserved.
Colour (in formaline).—Dark chocolate-brown, with paler clitellum.
Length, 50 mm., approaching N. macquariensis.
Prostomium is epilobic, one-third, with a transverse groove.
The chœtœ are isolated, the interspaces nearly equal, though the gap bc (= aa) is greater than ab or cd, and dd is considerably greater than aa. This chætal formula agrees with that of N. macquariensis.
The clitellum is complete, extending over segments 13 to 16 in the only mature specimen in my possession.
Genital Pores, &c.—The porophores are white, in line with b. There are also paired copulatory tubercles, quite pale in colour, near the anterior margins of segments 17, 19, and 20, in line with a; while behind the spermathecal pores in segments 8 and 9 similar tubercles occur in the same line. This, in general, agrees with what I have recorded for N. aucklandicus, for in N. macquariensis no mention has been made of such tubercles.
Internal Anatomy.—The dorsal vessel is single, and the last
[Footnote] * Trans. N.Z. Inst., 1903, p. 267.

heart is in the 12th segment, as in N. aucklandicus. There are no distinct œsophageal glands.
Reproductive System.—The long undulating prostates extend through six and four segments respectively, terminating in segment 22. The duct is thick and muscular, and rather swollen at its entrance to the body-wall.
The penial chæta is rather strongly curved below the tip, as in the allied species, but is more delicate than in either of these. The tip is blunt, as in N. aucklandicus, and the ornamentation is in the form of pectinated ridges, as in that species, but are not so close together as in the type, nor do they extend so far towards the tip.
The spermatheca resembles that of N. macquariensis in its globular form and two divergent diverticula.
Remarks.—This species is evidently very closely allied to N. aucklandicus, yet it differs in several respects from it, and approaches N. macquariensis in these points—viz., in its small size, in the spacing of the chætæ, absence of distinct œsophageal glands, form of the spermatheca.
Although I have given a new specific name to this worm, it may become desirable to consider these three forms as varieties of one species.
Hab. Campbell Island. In the bottle containing these specimens were also some Enchytrœus albidus, Polychæta, and Nemertine. They were all collected near one another close to the sea.
Rhododrilus cockayni, n. sp.
Seven individuals were collected, of which the majority are sexually mature.
Colour.—The general colour is a greyish-purple, darker anteriorly, with brown clitellum. The posterior region is very pale, with a purplish-grey line along the dorsum. The body-wall is thin.
Dimensions.—The specimens have a length of from 100 to 150 mm., with a diameter of 3.5–4 mm. immediately behind the clitellum. The segments number 72 in the shorter and 110 in the larger individuals.
The prostomium is epilobic, about one-quarter, without a transverse furrow.
The chœtœ: a and b are nearer together than are c and d, and the formula here is aa = bc; dd = 2cd = 3ab; or ab < cd < bc < dd; while towards the tail ab widens out so that the four chætæ on each side are nearly equidistant.
The clitellum is saddle-shaped, extending as far downwards as chæta b, and covers segments 13–17.
Genital Pores, &c.—The single pair of porophores on the 17th

segment are in line with ab, but the male pore is in line with b. Copulatory tubercles are variously developed, but in all the mature specimens there are paired oval post-chætal glands on segments 10 and 11 in line with ab. Some individuals have others on segment 9 and even on the 8th as well. In a single case, similarly paired papillæ occur on segments 19 and 20 as well as on the two anterior segments, and all are in the same relative position on the segment, and have the same form—viz., transversely oval, with a slight depression—giving the impression of a sucker.
Spermathecal pores: Three pairs, at the anterior margin of segments 7, 8, and 9, in line with b. Nephridiopores also in line with b.
Internal Anatomy.—The six septa behind segments 8–13 are stout. The dorsal vessel is single; last heart in segment 12. The worm is meganephric.
Alimentary Tract.—A very small gizzard occupies the 5th segment. A large lobulated and highly vascular salivary gland lies on each side of the pharynx. The œsophagus is distinctly dilated in the 13th segment to form a spherical sac, the lining of which has the general villous structure of a gland, though the lateral region is not constricted from the axial œsophagus.
In segments 14–17 the gut is narrow, with a yellowish wall; in the 18th it suddenly dilates to form the thin-walled intestine.
Reproductive System.—Three pairs of sperm-sacs occupy segments 9, 11, and 12. The spermiducal (prostate) glands are long, tubular, and undulating: each extends through the five segments 17 to 21 inclusive. The short muscular duct is confined to segment 17. The sacs of penial chætæ are also long. On the removal of the arcuate muscles from segments 16, 17, and 18, the two sperm-ducts on each side are readily seen on the body-wall, and can be traced backwards outside the prostate duct, where they bend mesially behind it and the penial sac to open to the exterior.
The penial chæta (Plate XIV., fig. 6) is long, delicate, and curved rather abruptly near the tip, which is slightly recurved.
The only “ornamentation” is in the form of a few scattered minute serrated markings some little distance below the tip.
The three pairs of spermathecæ are in segments 7, 8, and 9 (and in one specimen from Adam Island there are three on the left side but four on the right). These increase in size backwards, and the diverticulum becomes relatively longer. Each spermatheca (Plate XIV., fig. 2) consists of a large somewhat ovoid sac with a short wide duct, into which opens a curved cylindrical diverticulum. This, in mature forms, is longer than

the “ampulla,” though in the young form and in the anterior sacs it is shorter.
Hab. Campbell Island; Ewing and Adam Islands of the Lord Auckland Group.
Rhododrilus leptomerus, n. sp.
This peculiar species is represented by two mature individuals collected “on moss (? Auckland Islands).”
Colour.—Pale-yellowish, with a white clitellum.
Dimensions.—Length, 75 mm.; breadth, 3 mm; with 82 segments.
Prostomium epilobic, one-half, with a posterior transverse groove.
Chætœ in couples, the individuals of the ventral couple being rather near, the other spaces wide: aa = bc = cd = 2ab; dd = 1¾aa.
The clitellum is saddle-shaped, and covers segments 13 to 16; it has well-defined margins anteriorly and posteriorly, and appears to be fully developed, as the intersegmental furrows are obliterated except on the ventral surface.
Genital Pores, &c.—A single pair of porophores occurs on the 16th segment in line with cheeta &: it is only feebly prominent, and on it are three pores, as in other species of the genusviz., one common to the sperm and prostate ducts, the other two for the two penial chætæ. There are no copulatory tubercles. There is a single pair of spermathecal pores between segments 7 and 8. Nephridiopores in line with chæta c.
Internal Anatomy.—There are no thickened septa. The only notes on the vascular system are from longitudinal sections, which show two pairs of hearts, in segments 10 and 11 respectively, of which those in the former segment are especially large. But owing to the hard-coagulated blood in the wall of the gut and blood-vessels the sections were somewhat torn, and I was unable to trace the matter further.
The Alimentary Tract.— There is a short thick-walled gizzard in segment 6, concealed by the muscles of the pharynx. The œsophagus is dilated in segments 12 and 13, but no definite glands are formed. The intestine commences in the 15th segment, where the thick wall of the œsophagus is replaced by a thin wall. The typhlosole is a very low ridge.
Reproductive Organs.—There are two pairs of testes in segments 9 and 10. The three pairs of sperm-sacs are in segments 8, 10, and 11. The first is small and attached to the hinder septum; the others larger and attached to the anterior septa of their segments. All are “botryoidal” and contain developing spermatozoa. The spermiducal gland on each side is very long,

undulating, and bent in a U-shaped fashion. It occupies five segments, 16 to 20 inclusive. The free end of the gland lies in the former segment, and the closely undulating or zigzag portion extends into the 20th, against the hind wall of which it bends sharply on itself, and the rather wider recurved limb passes forwards into the 19th, at the anterior end of which the muscular duct commences. This is narrow and long, passing through three segments to open to the exterior at the 16th segment. The penial sac is also long, occupying the four segments 16 to 19 inclusive.
The penial chæta (Plate XIV., fig. 7) is long, curved, with a spoon-shaped pointed tip, and is without ornamentation. The total length of a penial chæta is 4 mm.
The ovaries are plainly visible in the dissected worm in the 12th segment.
The single pair of spermathecæ lie in the 8th segment. Each (Plate XIV., fig. 3) is a subglobular sac with a short thick duct, which receives a relatively large cylindrical diverticulum which is bent upon itself.
Remarks.—The most remarkable feature about this species is the apparent shifting forwards of the genital organs by one segment—i.e., the genital pore, instead of being on the 17th, is on the 16th segment, and each of the internal genital organs-ovaries, testes, and sperm-sacs—similarly occupy the segment preceding the one normal for the genus. As to the spermatheca in the 8th segment, it is uncertain whether this has been affected, for in R. edulis the single spermatheca is in this segment, though in other species with a single pair it is in the 9th.
The position of the male and female gonads is so constant throughout the whole group of earthworms, with the exception of the Moniligastridœ, that this forward movement is very puzzling.
Both individuals present this same dislocation. Naturally, this led me to re-examine very carefully the external and internal segmentation, both with a lens of high power and, after bisection and clearing, with the compound microscope, and, further, I cut one individual into a series of longitudinal sections. It at once occurred to me that the prostomium and peristomium might be invaginated, or that an injury had been received at this end; but the latter suggestion was negatived by the occurrence of two individuals presenting precisely the same phenomenon; and, moreover, there is no sign of injury at this anterior end. The prostomium is perfectly well defined; it has the usual relations to the peristomium—it is, indeed, “dovetailed” into the latter segment.
The 1st segment has no chætæ—i.e., it agrees with the usual

character of the peristomitum. The chætæ commence in the 2nd segment. This and the following segments are perfectly normal: there is no indication of a double set of chætæ in any of them. There is no furrow in the 1st or 2nd or any subsequent segment to suggest a fusion of two segments; in fact, the segments are not annulated, as is sometimes the case. The worms are fairly soft, not contracted at all, so that the limits of the segments are quite distinct and definite. There seems no possibility of error in the enumeration of the segments such as does occur in a strongly annulated worm (e.g., Octochætus). There seems to be no means of explaining the peculiarity from an external examination. Internally, too, the septa, though thin, are quite distinct. There is none of that shifting of septa that occurs in many large worms; and, as the soft condition permitted it to be fully extended, there is no crowding of the septa. Repeated countings, both externally and internally, gave the same result—viz., the ovaries (fortunately large and readily seen) in segment 12, and other organs one segment forward. There is no “spiral segment” such as has been observed by Morgan and others. Longitudinal sections show that the cerebral ganglia occupy the normal position at the hinder part of the 3rd segment; the circumpharyngeal commissure lies in this segment, and the first ventral ganglion occupies the anterior half of the 4th segment, and a single ganglion corresponds to each of the subsequent segments.
The condition of preservation is not sufficiently good to enable me, with certainty, to study the detailed distribution of the ganglion-cells in the first ventral ganglion; but, from a consideration of the facts, the only way to explain the shifting of the organs is to imagine an “excalation” of a segment in front of the testes: if the spermatheca has been unaffected, then the original 9th segment has disappeared; or if the spermatheca has been moved forwards, then one segment lying between the 2nd and 8th has disappeared; but as the gizzard is in the 6th segment the former suggestion is the more probable. If we had had only a single individual this peculiar forward shift would have been remarkable enough, and might have been explained by supposing that the anterior extremity had been cut off, and that an imperfect regeneration of segments had followed—a regeneration in which one segment short of the full number had been formed; but when two individuals exhibit exactly the same phenomenon it does not seem possible to refer it to regeneration. Nevertheless, I do not consider it necessary to form a new genus for the reception of the worm—as we should have done a few years back—for in all essential structural points it agrees so closely with species of Rhododrilus that it must be looked upon

as an extremely abnormal species of that genus: though it is true that in no other species is the spermiducal gland provided with such a long muscular duct.
Marionina antipodum, n. sp.
Four small Enchytræids appear to belong to this new species.
The longest is 11 mm. in length and 0.75 mm. broad, with 35 segments.
The chætæ are very feebly sigmoid, the points being slightly curved: there are nearly constantly four in each bundle throughout the body, only in a few anterior segments are there five in each bundle.
The clitellum is girdle-like, completely covering segments 12 and 13.
The usual head-pore was noted.
The œsophagus passes gradually into the intestine, being, however, rather dilated in segments 8, 9, and 10.
There are four pairs of septal glands, lying in segments 4, 5, 6, and 7.
The dorsal blood-vessel commences at the hinder end of the 13th segment, and immediately in front of its origin it becomes a good deal dilated, but no “cardiac body” is present. The blood was in life apparently colourless, as the blood-vessels, instead of being filled with a red-stained or yellowish coagulum, are empty. In an entire specimen (stained) I was unable to detect the vessels; but in transverse sections the dorsal vessel appears as a small empty tube with a single nucleus on either side; while even anteriorly the ventral vessel appears to be closely adherent to the œsophageal wall.
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
The segments 10 and 11 are filled with spermatozoa, and the septum 13/12 is pushed back to the end of the 13th segment.
The testes are quite small and relatively loose in structure, though there are no definite lobes, the edge being slightly frayed.
The sperm-funnel is a good deal curved in the entire specimen, but appears to be about four times as long as its breadth. The penial apparatus (Plate XIV., fig. 9) is comparatively small, as it scarcely exceeds the thickness of the longitudinal muscles of the body-wall. Opening into it, however, is a conspicuous prostate gland.
The spermatheca (Plate XIV., fig. 10) is a long pyriform organ, in which the muscular duct is not distinctly marked off from the sac, which, moreover, is empty in all the individuals examined. The duct is without glands, but at the pore is a couple of groups of gland-cells—one anterior and one posterior.
It should be noted that the longitudinal muscles, instead of

consisting of a single row of fibres, present several layers, as Ude has noted in Lumbricillus verrucosus.
Loc. Antipodes Island.
Remarks.— I cannot fit this worm to any of the species diagnosed in Michaelsen's monograph.* It does not agree with any of the South American forms described by Ude;† for, amongst other differences, I do not find any “subneural glands” in the present worm, although the specimens sectioned are sexually mature.
Enchytræus albidus, Henle.
Specimens indistinguishable from this widely distributed species were collected on Campbell Island, close to the sea; and I have specimens from Macquarie Island, which were collected some years ago. I may add that a re-examination of the worm collected and named E. simulans by me‡ convinces me that this name must be eliminated: they are E. albidus, as I at first imagined them to be.
Lumbricillus macquariensis, n. sp.
Several specimens of this Enchytræid were collected some years ago by Mr. Hamilton on Macquarie Island, and are entered in the Museum register by the late Professor Parker as “small Oligochæta from brackish pools, with Siphonaria, planarians, &c.” I take this opportunity of giving an account of this species.
Length, 23 mm.; breadth, 1.25 mm.; with 60 segments.
Chætæ very feebly sigmoid and relatively small, in the usual four bundles of 4–7 per bundle. Anteriorly there are in several, segments six dorsal and four ventral, while in the 2nd segment six dorsal and seven ventral. Posteriorly the usual numbers are four or five dorsal and six or seven ventral. There are no ventrals in segment 12, and the dorsals are few in the 12th and 13th segments.
The clitellum covers segment 12 and part of the 13th, ceasing at the level of the chætæ. The male pore, in 12, is in some cases prominent owing to the protrusion of the terminal organ.
The usual head-pore is present between the prostomium and the 1st segment. The œsophagus passes very gradually into the intestine, which commences in the 15th segment. There are three pairs of septal glands, as usual.
The dorsal blood-vessel arises either at the hinder end of the
[Footnote] *Michaelsen, “Das Tierreich: Oligochæten,” 1900.
[Footnote] †, Ude, “Enchytraeiden,” in Hamb. Magalhaen, Sammelr., 1896.
[Footnote] ‡ P.Z.S., 1903, vol. ii., p. 219.

13th or at the commencement of the 14th segment. The blood appears to have been reddish in life.
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
In segments 10 and 11 there are great lobulated. masses attached to the body-wall at the insertion of the septum 10/11, some lobes passing forwards and even reaching into the 9th segment, others backwards. This is the “multiple testis” (of Claparede); but, as Michaelsen has shown, it is only “testis” at its base—towards the extremities of the lobes the sperm mother cells are found dividing up, and various stages in sperm-formation occur.
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
The tub-shaped sperm-funnel (Plate XIV., fig. 13) in the 11th segment has a length equal to about twice its breadth. The narrow sperm-duct coils considerably immediately after passing through the septum 11/12, then takes a straight course to the “penis.” This consists of a spherical mass of gland-cells enclosed in a thin muscular coat, consisting of circular and longitudinal muscles. Some of the gland-cells open directly to the exterior, others into the sperm-duct as it passes through the apparatus. The sperm-duct arrives at the outer side of the “penis,” which it perforates obliquely to open externally at its centre. The area of the body-wall over which the gland-cells open is, in the specimen sectionised, retracted, so that a deep narrow pit results.
The ovary, like the testis, is “multiple”—i.e., lobulated—each lobe being moniliform, consisting of strings of ova, while a few large ova lie free in segment 13.
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
The oviducal pore is visible on the mounted specimen at the junction of segments 12/13.
The spermatheca has the usual position, opening, that is to say, between segments ⅘. It (Plate XIV., figs. 11, 12) is a cylindrical tub-shaped sac, with a sharply constricted, very short duct which is surrounded by a circle of gland-cells. At the opposite end of the sac a narrow tube puts it into communication with the œesophagus.
The nephrida have a small pre-septal portion, and a large post-septal region whence the duct passes from the hinder end to the body-wall, usually bending forward below the rest of the organ.
There are three subneural (? copulatory) glands—two large ones segments 14 and 15, and a smaller one in the 16th segment. In transverse section it is seen that the gland (Plate XIV., fig. 8) rises up the sides of the nerve-cord and leaves the greater part of its upper surface uncovered. Each lobe is of considerable size—at least four or five times the diameter of the nerve-cord—and extends laterally as far as the ventral chætæ. The ducts i.e., necks—of the gland-cells pass through the circular muscular

coat of the body-wall, and spread out fan-wise below the epidermis.
Remarks.—This species appears to be nearly allied to L. maritimus, Ude,* from which, however, it differs in the following points: It is more than twice the length, and contains nearly twice as many segments; the excess of chætæ in the dorsal bundle of some of the anterior segments, and the fewer chætæ in the bundle of the hinder segments; the less extent of the clitellum; the relatively smaller size of the sperm-funnel, which in L. maritimus is three or four times longer than broad (the form of the spermatheca appears to differ, for Ude makes no mention of the narrow œsophageal duct, saying that the “beutel-formige” sac communicates by its narrow end with the gut); the relatively great size of the subneural glands in the present species; and the presence of a third small one in the 16th segment.
Explanation of Plate XIV.
| Fig. 1. |
A spermatheca of Rhododrilus kermadecensis. |
| Fig. 2. |
A spermatheca of Rh. cockayni. |
| Fig. 3. |
A spermatheca of Rh. leptomerus. |
| Fig. 4. |
Penial chætæ of Rh. kermadecensis, seen from above (x 350). |
| Fig. 5. |
The same, side view, showing spoon-like tip (X 350)., |
| Fig. 6. |
Penial chæta of Rh. cockaym (x 350). |
| Fig. 7. |
Penial chseta of Rh. leptomerus (x 350). |
| Fig. 8. |
Lumbricillus macquariensis. A transverse section of a subneural gland (camera × 175). a, nerve-cord; c, circular muscular coat; e, epidermis; l, longitudinal muscular coat |
| Fig. 9. |
Marionina antipodum. The aperture of the sperm-duct, from a transverse section (× 175, camera). d, sperm-duct; gl, spermiducal gland, opening into terminal apparatus; p, pore; l, longitudinal coat of muscles consisting of several fibres in depth. |
| Fig. 10. |
Marionina antipodum. The spermatheca. gl, gland at its exit; a, œsophageal opening; p, external pore. |
| Fig. 11. |
Lumbricillus macquriensis. A spermatheca, with rosette of glands at the pore. o, œsophageal opening. |
| Fig. 12. |
The same. Base of spermatheca (from a transverse section), showing very short muscular duct, and the glands around the pore. |
| Fig. 13. |
The same. The male apparatus for an entire individual. d, duct; f, funnel; p, pore; s, septum. |
[Footnote] *Ude, “Enchytraeiden,” Hamb. Magalhaen, Sammelr., 1896, p. 6.

Art. XX.—Earthworms from the Kermadecs.
[Read before the Otago Institute, 8th November, 1904.]
Plates XIII. and XIV.
Owing to the thoughtfulness of Mr. R. Shakespear, of Little Barrier Island, I am indebted to Captain Bollons, of the Government steamer “Hinemoa,” for the first samples of the Oligochæta that have been collected in the Kermadecs. Captain Bollons was good enough to take a considerable amount of trouble to obtain these worms. The soil forms, he tells me, a very thin covering of decayed vegetation upon pumice rock, and he was unsuccessful in his earlier efforts to find any earthworms, but finally, at Coral Bay, his persistency was rewarded. My best thanks are due to these gentlemen for their kindness.
I received the specimens alive, packed with soil and moss in a tin box: they consisted of three small earthworms and six Enchytræids. Of the earthworms, two were immature individuals of some species of the genus Allolobophora belonging to the family Lumbricidæ—the members of which are readily transported, and are, as a matter of fact, widely distributed by the agency of man. Though I am unable to identify the species to which these individuals belong, there can be little doubt but that they have been “introduced” by some means, though Captain Bollons states that this part of the island has never been inhabited; nevertheless, some parts have been occupied, and there can be no difficulty in explaining the presence of these worms here. The third worm is mature, and belongs to the genus Rhododrilus,* which, according to Michaelsen's most recent writings, has its headquarters in New Zealand, and is represented on both the main islands as well as on the outlying islands. The present species, however, differs from any hitherto described, and I propose to name it
Rhododrilus kermadecensis, n. sp.
Colour.—Pinkish, with transparent body-wall.
Dimensions.—Length, 65 mm.; diameter, 1.75 mm.; with 108 segments.
Chœtœ.—The eight bristles are wide apart, and approximately their arrangement in the midbody is ab =½bc; bc = cd = 2ab; aa = bc; dd = 2cd. But at about the 20th segment the lines
[Footnote] * It appears from a recent work by Michaelsen that he would fuse the species of Rhododrilus with those of Microscolex, the older genus. I have given reasons in a paper forwarded to the Zoological Society for distinguishing the two genera.

of c and d descend slightly, so that the gap bc is reduced, and dd increased, in the forepart of the body.
Prostomium epilobic, about one-half, without a posterior transverse groove.
Clitellum covers segments 13–18 inclusive. The ventral chætæ and intersegmental furrows are visible, though the skin is somewhat thickened across the ventral surface.
Genital Pores, &c.—A pair of large porophores on the 17th segment, in line with chætæ ab; and in front of the chætæ there is a single large median “copulatory tubercle,” in the form of an oval glandular pad, not very prominent, covering the whole of the ventral surface of segment 19, and the prechætal region of the 20th.
Dorsal pores are visible posteriorly.
Internal Anatomy.—In the bisected individual I made out the following points, sufficient to characterize the species: The dorsal blood-vessel is enlarged in segments 14, 15, and 16; the last hearts are in the 13th segment. The gizzard is quite minute, in segment 5. There is no œsophageal gland, though the tube is dilated in the 13th and 14th segments. The intestine commences in the 18th.
The Reproductive System.—The sperm-sacs lie in segments 9, 11, and 12. The prostate is a large, tongue-shaped gland, extending through the six segments 17–22, with a short muscular duct in the 17th. A couple of long chætal sacs underlie the prostate, extending back from the 17th to the 21st segment. Each sac contains three chætæ—a functional “penial chæta” and two smaller ones at different stages of development. The penial chæta (Plate XIV., figs. 4, 5) is long, delicate, and bent distally. The tip is curved, spoon-shaped, so that the side and top views differ. The surface is quite devoid of ornamentation.
There are three pairs of spermathecæ, in the segments 7, 8, and 9. Each (Plate XIV., fig. 1) is an ovoid sac, with a short muscular duct about half the diameter of the sac. It receives close to the body-wall a long cylindrical diverticulum, which in the specimen, studied is curved in an S-shaped manner. The tip is not dilated (till compressed), but the base is enlarged at its union with the duct.
Remarks.—In the possession of three pairs of spermathecæ this species resembles R. cockayni from Campbell Island, from which, however, it differs in its colour, dimensions, arrangement of copulatory tubercle, and form of penial chæta.

Art. XXI. —Note on the Occurrence of the Foraminiferan Genus Ramulina in the New Zealand Waters.
[Read before the Otago Institute, 13th September, 1904.]
During a dredging excursion in the summer of 1904, in which Mr. Charles Hedley, of the Australian Museum, Messrs. Suter, Cooper, Park, Murdoch, and others, took part, several interesting animals were obtained at a depth of about 100 fathoms, and amongst them the peculiar glassy foraminifer Ramulina globulifera. This has been described from the New Zealand waters by Brady, in the “Report of the ‘Challenger’ Expedition” (vol. ix.), but as it has escaped the notice of Mr. Hamilton, who drew up the admirable abstract from the “Summary,” and published by the New Zealand Institute under the title of “Deep-sea Fauna of New Zealand,” and as it is not enumerated in the “Index Fauæ Novæ-Zealandiæ,” it seems desirable to place on record its occurrences within the New Zealand area.
The specimens which were handed to me by Mr. Hamilton for identification were obtained at a depth of 110 fathoms, east of Great Barrier Island, lat. 36° 8 S., long. 175′ 55 E.
The following is quoted from Brady (loc. cit., p. 587):—
“Fam. Lagenid Æ.
“Subfam. Ramulin Æ.
“Test branching, composed of spherical or pyriform chambers, connected by long stoloniferous tubes.
Ramulina, Rupert Jones (1875).
“Test free, branching; consisting of a calcareous tube swollen at intervals so as to form more or less definite, often irregular, segments, from which lateral stolons or branches are given off. Texture hyaline.”
R. globulifera, Brady (1869) was obtained by the “Challenger” off the west coast of New Zealand in 145 and 275 fathoms (the latter was at station 166).
It has been met with on various stations in the Pacific, and also in the North Atlantic.
Figures of the species are given on pl. lxxvi, figs. 22–28, of vol. ix., “Challenger” Reports.

Art. XXII.—Further Notes on the Sipunculids of New Zealand.
[Read before the Otago Institute, 8th November, 1904.]
Plates XV. and XVI.
During the present year I have received specimens of two new species of Sipunculids, each representing a distinct genus not hitherto recorded from our coastal waters; and, in the absence of the requisite literature whereby to compare these with species described from elsewhere, I propose to give new names to them for the purpose, at any rate, of reference.
Phascolosoma,* Leuckart.
P. novæ-zealandiæ, n. sp.
A specimen was removed from the stomach of the dog-fish Mustelus antarcticus, which was being dissected in the Biological Laboratory.† The Sipunculid was partly macerated, but not so much as to prevent a study of its anatomy, sufficient, I believe, to characterize it.
The body-wall had been torn (by the shark's teeth, perhaps), and the skin had separated from the muscular coats, which; together with the viscera, protruded through the rupture.
Colour.—The skin is pale-yellowish; the posterior end and the introvert are pale-brown, probably darker in life.
Dimensions.—The total length of the skin is 310 mm., of which the introvert measures 75 mm.; the diameter of the body was about 10 mm.; and the base of the introvert about 4 mm. (Plate XV., fig. 1). These measurements are only approximate: in the first place the skin was softened and undoubtedly more extended than if it had been preserved in the usual manner; again, the skin was flattened so that the diameter had to be estimated.
General Description.
The skin is rough, with small brown tubercles and papillæ scattered more or less uniformly over the whole body and introvert (Plate XVI., fig. 8); but they are rather more densely arranged in the latter region, and also at the hinder end of the
[Footnote] * See my article in vol. xxxvi. of the Trans. N.Z. Inst., p. 172. The worm named by Hutton Phascolosomá annulatum belongs to the genus Physcosoma, as defined by Selenka.
[Footnote] † At the same time and from the same viscus a species of Echiurus was found which differs from E. novœ-zealandiœ, Dendy, in having two circles of hooks at the hinder end; but its internal organs were absent, so that no means of identification were to hand.

body. Here, too, the skin is wrinkled, forming irregular circular and a few obscure longitudinal ridges.
[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]
Each papilla as seen when the skin is examined under the microscope (Plate XVI., fig. 9) is somewhat ovoid in elevation, with a narrowed base of attachment; it is circular in plan, and about 1/10 mm. in diameter. It appears as a thick brown ring with a central light area. Further investigation by means of section shows that the epidermis has macerated away, and that these papillæ are entirely dermal, with a wall of connective tissue; and probably, in well-preserved material, this clear centre will not be visible (Plate XVI., fig. 10). In my sections the axial canal communicates with the subdermal spaces, and is traversed by what I take to be a nerve, which breaks up just before reaching the apex of the papilla into fine fibrils which are less stained than the nerve itself. Possibly, however, this axial thread is the remains of some secretion from gland-cells which have been destroyed (of. Shipley's figure of the skin of Oncchnesoma, pl. ix., fig. 6, Quart. Journ. Micr. Sci., xxxiii.). But it is unnecessary to discuss this matter here.
The tentacular crown was invaginated, so that the muscular wall of the introvert had to be slit open in order to ascertain the arrangement. There are numerous simple filiform tentacles springing from an undulating line of origin; and corresponding with four of these undulations are four well-marked thickenings or ridges along the œsophageal wall, of which two lie on each side, leaving a gap dorsally and ventrally. By pushing forward the wall of the œsophagus so as to cause the tentacles to occupy their proper position when fully extended, I have, as a result of the study of this preparation, been able to make a drawing of what is probably their appearance in life (Plate XV., fig. 2).
Internally, I was only able to note the following facts: The longitudinal muscles form a continuous sheet. There are two retractor muscles only, attached to the body-wall pretty far forward (my measurement gives 70 mm. below the tentacular crown: this was taken along the muscular wall, which, as above stated, was separated from the skin, hence the discrepancy between this statement and the “length of introvert”).
The alimentary canal still contains mud, and its wall is pretty firm, but had been thrown into a good deal of disorder by being ejected through the rupture in the body-wall—hence I can give no satisfactory account of its coils. Notwithstanding a certain degree of digestion to which the animal had been submitted, the dorsal blood-vessel still retains an orange-brown tint, and can be traced up to the base of the tentacles and along the dorsal ridge.
Remarks.—The only species with which I have been able

to institute any detailed comparison is the antarctic form, P. capsiforme, Baird,* and the varieties described by Michaelsen.† From this species it differs in several important respects, and is quite distinct from it.
Sipunculus, Linnæus.
S. maoricus, n. sp.
Two individuals were forwarded to me by Mr. Suter. They were found washed up on the beach at Tauroa Point, Ahipara, near Cape Maria Van Diemen. They had apparently dried up to some extent before being preserved, as the skin is tough and hard, and the internal organs are not sufficiently well preserved to be described.
Colour.—Pale-yellowish; perhaps somewhat translucent when alive.
Dimensions (Plate XV., fig. 3).—Total length, 116 mm., of which the introvert occupies 12 mm.; the diameter of the body at about the middle is 10 mm., and the base of the introvert measures 4 mm.‡ The anus lies 6 mm. below the base of the introvert. Thus the length is about 11½ times the breadth, and the introvert is about one-ninth of the total length.
General Description.
The body is, as always in this genus, marked with distinct circular and longitudinal furrows, dividing the surface into a number of rectangular areas (Plate XVI., fig. 13); the circular furrows and ridges being more conspicuous in that part of the body which is contracted, the longitudinal ones where it is less contracted. The hinder end of the body is slightly pointed, and here the longitudinal ridges are most conspicuous, converging to the mammilla-like tip.
The introvert, which is partly retracted in both specimens, is for the greater part of its extent covered with recurved hook-like tubercles, which are white (Plate XVI., fig. 11). These are not arranged in any very definite manner, though partly they form oblique rows. The tubercles are rather larger dorsally than ventrally, and also diminish slightly in size as the tentacles are approached, at the same time becoming softer. Between the tentacular crown and the tubercles is a naked area about 2 mm. or 3 mm. in extent (Plate XVI., fig. 7). Each of the hook-shaped tubercles§ has an irregularly rounded but wrinkled apex. It is
[Footnote] * Fischer, “Gephyreen,” Hamburg. Magalhaen. Sammelr., 1896.
[Footnote] † Michaelsen, “Die Gephyr. v. Sud-Georgien.” Jahrb. Hamburg. Wiss. Anstalt vi., 1889.
[Footnote] ‡ Michaelsen, “Die Gephyr. v. Sud-Georgien.” Jahrb. Hamburg. Wiss. Anstalt vi., 1889.
[Footnote] § The word “papilla” has been used for glands immersed in the body-wall; otherwise it would seem a more suitable word to use in the present connection.

covered by cuticle, which is thicker on the external convex surface than on the apex or inner face; below this are several “bicellular glands,” which project downwards from the epidermis into the dermis or fibrillated connective tissue (Plate XVI., fig. 12). The latter is excavated by a great space which at the base of the tubercle is in free communication, by means of a small aperture, with the general body cavity. In this space is a mass of spherical cells, each with a refringent body and a deeply staining nucleus. At first I took this for a specialised group of connective tissue-cells, serving as a skeletal axis to the tubercle; but examination of serial sections shows that this mass of cells is continuous with masses of granular cœlomic cells adhering to the body-wall, and in some cases the transition between the two appearances—refringent and granular—can be made out. Unfortunately, the material is not sufficiently well preserved to enable me to describe in greater detail this structure; but it appears to serve as a means of erecting these tubercles, for when the strong muscles of the body-wall contract this cœlomic fluid will be driven into these subdermal spaces.
On the body-wall itself, as opposed to the wall of the introvert, there are no tubercles: the usual lymph-spaces alternate with groups of bicellular glands, as has been described by previous writers on the histology of Sipunculus.
As the tentacular crown was invaginated, it is not altogether easy to reconstruct the appearance which it has when fully extended. On slitting open the œsophagus, five ridges, covered with tentacles, are seen (Plate XV., fig. 5): of these one is dorsal, two are lateral, and two latero-ventral; the dorsal ridge is the largest, and bears more numerous tentacles. The tentacles themselves are flattened, membranous, and truncated distally—quite unlike the more or less cylindrical tentacles of Phascolosoma and other genera; and it has been shown by various authors that in the genus Sipunculus the tentacles are really the jagged edge of a membrane: various species show different degrees in which the margin of the circumoral membrane is cut into, so that in some the “tentacles” are short and in others longer. In the present species the membrane is almost entirely “frayed” out into these tentacles. By artificially pushing the œsophagus and crown upwards we may cause the crown to assume, to some degree, the condition of eversion; and from a study of such a preparation, and of the ridges themselves, I venture to “reconstruct” this tentacular crown (Plate XV., fig. 4), which is very unlike that of the species of Sipunculus usually figured; but from the brief diagnosis of “Phattosoma” Levinsen, given by Yves Delage and Herouard,* there seems to be some resemblance
[Footnote] * Traité de Zool. Concréte, v., p. 21.

to the tentacular arrangement of that arctic form. Again, in the same text-book, a form named Stephanostoma by Danielssen and Koren is stated to have “six great groups of tentacles”; but this form is regarded by Selenka as a Phascolosoma and not a Sipunculus.
The appearance, then, of this tentacular crown will be, when fully everted, very different from that of either Dendrostoma or Physcosoma.
Internal Anatomy.—The longitudinal muscular coat consists of about twenty-seven distinct bands in the middle of the body: these run independently for considerable distances—i.e., anastomoses are very infrequent; necessarily they become more frequent posteriorly, where the number of bands decreases.
There are four retractors, all of the same length, arising from the body-wall about 20mm. below the base of the introvert. The dorsal retractors arise from four longitudinal muscle-bands, and the ventrals from three of them.
The intestine, filled with sand, has such a thin wall—probably due to post mortem changes—that it was impossible to trace the coils, or to detect the extent of the spindle muscle, for the wall burst on the slightest touch. The coils were attached to the body-wall by numerous delicate threads. The rectum runs close to the body-wall for a distance of 8mm., and appears to be adherent thereto.
I was unable to detect any posterior coeca, nor were the nephridia preserved.
Remarks.—I have given a new name to this Sipunculus, as it does not agree with the descriptions of any species in the small collection of literature available. Unfortunately I have not access to Selenka's monograph, so that it is possible that this article is a work of supererogation. Judging by the reference to the hook-like tubercles of S. australis, I expected that it would belong to this species; but the figure and description given by Shipley* do not seem to bear out this idea: it is true neither figure nor description is very detailed, but they are sufficient to indicate general differences. His account says nothing of the great axial cavity of the tubercle, and his figure shows gland and tubercle as distinct things (p1. xviii., fig. 5); and the brief diagnosis given by Quatrefages† does not incline one to refer my specimen to that species. The same negative result follows a comparison of the diagnoses of other Australian species given by the latter author.
[Footnote] * Willey's Zool. Results: Report on Sipunculoidea.
[Footnote] † Hist. Nat. des Annelés, ii., p. 619.

Note on Dendrostoma.
D. huttoni, n. sp.
In my previous article (Trans. N.Z. Inst., xxxvi., p. 180) I spent some time in trying to demonstrate the probability that Sipunculus lutulentus, Hutton, is a synonym of S. aeneus, Baird, and proceeded to state that I believed the species of Dendrostoma that I was describing was identical with the latter. I am now of opinion that I was altogether too precipitate in my conclusions. Some of the reasons that led me, at that time, to formulate these conclusions were: (1.) Two genera of Sipunculids, and only two, had been collected in recent times on our shores, and these at various spots from Auckland Harbour to Stewart Island. (2.) Both these had been described, so far as external features are concerned, and had been named, by Captain Hutton. (3.) In this Museum were certain tubes containing examples of two forms, with Hutton's labels upon them: one of them is undoubtedly the type of Physcosoma annulatum; the other I had reasons to think might be the type of his Sipunculus lutulentus. (4.) The latter, however, I found to belong to the genus Dendrostoma.
The proper proceeding would have been to have named this species Dendrostoma lutulenta, Hutton; but I went further, and, concluding that Hutton's species was identical with Baird's, gave it the name of D. aeneum.
Now, there are two very important lacunae in the evidence for this procedure: Firstly, we have no knowledge whatever, from the brief account of the external anatomy given by Baird, of the generic status of “S. aeneus.” Secondly, we are by no means certain that the specimen alluded to as coming from Cape Campbell is Hutton's type of “S. lutulentus”; for, although Captain Hutton wrote me that, so far as he remembered, he had only one specimen of this species and that came from Cape Campbell, yet the individual is less than half the size of S. lutulentus as described by him. That ought to have put me on my guard against identifying the Dendrostoma with Hutton's species.
Now, in his original account of P. annulatum Hutton states that he had obtained specimens from Dunedin and Cape Campbell: the former is in the Museum, as I stated in my article, fully labelled by Hutton; but I cannot find any specimen of this species with the latter locality attached. It has occurred to me that possibly this specimen referred to as “b” in my article was mistaken by Hutton for an individual of P. annu-latum: it is true it differs in various ways from that species, even externally, but in size and general form it is more like it than like his description of S. lutulentus. At any rate, I ought to have

been contented with the suggestion merely of the identity of the Dendrostoma with Hutton's S. lutulentus. But since writing that article I have received representatives of two more genera, and this leads me to doubt whether even this suggestion is probable. We must bear in mind that at the time Captain Hutton wrote the limitations of the genera of Sipunculids were by no means so clearly recognised as at the present day; and it is not quite certain to what genus Hutton's species belongs.
I propose, then, to name the Dendrostoma after our leading naturalist in New Zealand, so that it will now stand as Dendrostoma huttoni, Benham=Phascolosoma huttoni, Benham (Index F.N.Z.) = Dendrostoma aeneum, Baird (Benham, Trans. N.Z. Inst., xxxvi., p. 177).
Both Baird's and Hutton's species of Sipunculus must stand amongst our “species inquirendae.” I much regret the confusion to which my former article will perhaps give rise.
The figure (p1. vii., fig. 12) illustrating my account* of the tentacular crown of Dendrostoma huttoni does not do justice to the elegance of this organ: the lobes are too broad and the tentacles too short. I have therefore drawn another figure from a fully extended specimen, which is reproduced on Plate XV., fig. 6, for comparison with the corresponding organ of the genera described in the present communication.
Explanation of Plates XV. and XVI.
| Fig. 1. |
Phascolosoma novae-zealandiae: half natural size. |
| Fig. 2. |
The tentacular crown of the same: oral view; magnified; reconstructed. |
| Fig. 3. |
Sipunculus maoricus: natural size. a, the partially retracted introvert; b, contracted region of the body, where the circular ridges are very distinct. |
| Fig. 4. |
Tentacular crown of the same: oral view; magnified; reconstructed. |
| Fig. 5. |
One of the latero-dorsal ridges of the tentacular crown of the same, as seen on the wall of the opened introvert: stained and mounted; the arrow indicates the anterior end of the body; (camera, magnified). |
| Fig. 6. |
Dendrostoma huttoni: oral view of tentacular crown; magnified. |
| Fig. 7. |
Sipunculus maoricus: side view of the anterior end of the introvert; magnified. a, tuberculated region; b, smooth region. |
| Fig. 8. |
Phascolosoma novae-zealandiae: a piece of skin from the body-wall, as seen through a hand-lens, showing the characteristic papillae (p). |
| Fig. 9. |
The same: a papilla much enlarged. |
| Fig. 10. |
The same: a papilla in longitudinal section (camera × 80). ax, axial canal; ct, dermis; pg, pigment granules of the outer part of the dermis (the epidermis had macerated off); x, cord of granular material in the axial canal (? nerve or secretion). |
[Footnote] * Trans. N.Z. Inst., xxxvi., p. 177.

| Fig. 11. |
Sipunculus maoricus: piece of the wall of the introvert (seen under a low power), showing the hook-shaped tubercles; with (a) an axial mass of cells. |
| Fig. 12. |
The same: a tubercle in longitudinal section (X 80). a, axial mass of cells, in continuity with—c, a group of caelomic corpuscles; cm, circular muscles of body-wall; d, dermis; gl, epidermal glands; lm, longitudinal muscles. |
| Fig. 13. |
The same: a portion of surface of body (as seen through a hand-lens) showing the characteristic longitudinal and circular grooves marking out raised rectangular areas. |
Art. XXIII.—The Aquatic Larva of the Fly Ephydra.
[Read before the Otago Institute, 9th August, 1904.]
Plate XVII.
Amongst the organisms collected by Mr. J. A. Thomson in a saline pool at Barewood, Central Otago, which were submitted to me for identification by Mr. G. M. Thomson, were a number of small brown larvae, or rather “puparia,” having rather a peculiar form. This larva, which for reasons stated below I ascribe to the Dipteron genus Ephydra, resembles in general form and structure the “rat-tailed” larva of the hover-flies (Helophilus* and Eristalis †)—in the very reduced condition of the head, in possessing a series of paired groups of claw-shaped spines segmentally arranged along the ventral surface, and in the long posterior respiratory tube or “tail”; but in the two genera just named this tube is retractile, whereas in the insect herein described it is not retractile, and is, moreover, bifurcated.
In looking through the small amount of general literature on the Diptera available here, the only genus of flies that I was able to find in which the larva of this general form of structure has a bifurcated respiratory tube is Ephydra. No doubt a figure of this larva is to be found in one or more recent monographs, or in memoirs published in periodicals concerned with entomology; nevertheless, in the interests of those who are unable to consult these works, it seems to me worth while to present a brief illustrated account, so that naturalists in New Zealand may be able to identify the creature. My identification rests upon the brief accounts contained in Westwood's “Introduction to the Modern Classification of Insects” (1840), and in Packard's “Guide to the Study of Insects” (1872). The former author
[Footnote] * Hudson, N.Z. Entomology, p. 58, p1 vii., fig. 1.
[Footnote] †Miall's Nat. Hist. Aquatic Insects. p. 198 et sea.

gives a poor figure (vol. ii., p. 569, group 132, fig. 11) which shows the bifurcated tail, following ten claw-bearing segments. He states (p. 574), “Many species of Ephydra frequent salt marshy situations. The larva is cylindrical, without feet, and the terminal segment of the body very long, and terminated by a long fork, the prongs of which support spiracles at the tip; the puparium scarcely differs from the larva.” Packard, on p. 414, also represents, in a slightly more detailed figure, the “puparium” of another species. I find no reference to the genus in Miall's interesting book, nor in the Cambridge Natural History.
Description of the Puparium.
Amongst the dark-brown puparia I find one grey larva, which differs in a few points from the puparia, to which I paid more special attention; and since the larva, by hardening of the stem, becomes the puparium, the description of the latter will serve, with slight modifications, for the former. The puparium is a dark-brown hard-skinned object, measuring 9 mm. in total length, with a greatest diameter of 2 mm. It is cylindrical, bluntly pointed anteriorly, and produced into a narrow cylindrical tail occupying about one-third of the total length—viz., 3.5 mm. This terminates in a pair of short, narrow tubes, paler than the body, but tipped with dark pigment; each branch or tube is 1 mm. in length, and bears a spiracle at its apex. The entire surface—except these terminal branches—is covered with closely set blackish spines (as in Eristalis), groups of which at certain segments become larger and claw-like.
The body consists of a small “head,”* carrying a mouth, followed by ten distinctly marked segments, the last bearing the anus, behind which is the long tail.
There are no definite feet on this puparium, but on the ventral face of each of the eight segments 3 to 10 are paired groups of black claw-like spines, those on the last segment being carried on a prominent median papilla. In the majority of the specimens the ventral surface of the 9th segment is sunk below the general level, and, the body being abruptly curved ventralwards in this region, the claws of segments 8 and 10 are brought close together, and even in contact in some specimens, thus giving rise to clasp-ing-organ.
Westwood's figure shows nothing of this kind, all the segments being represented alike, and he places claws on all ten segments. Packard's figure, however, shows a slight prominence behind the 7th, and another larger one on the 10th. These, however, are wide apart, and do not suggest a clasping-organ.
[Footnote] * Possibly the true head is retracted.

The black claws are not arranged in circles, as they are in Eristalis, but in irregular groups, and the number differs in the different segments.
The first segment is armed on the ventral surface with a series of closely set black rectangular plates, each toothed along its hinder margin; these plates are in about half a dozen transverse rows, each row shorter than the preceding, giving rise to a somewhat triangular black patch. The anterior edge of this and of the second segment is also armed with a row of short black conical tooth-like spines.
The “head” bears on either side a small triramous organ, each branch of which is somewhat dilated at its extremity, and is apparently perforated by a minute aperture. Each is traversed by a brownish granular canal (in which I could not detect any rings), which at the base of the organ unites with the other two to form a single canal, which runs backwards for a short distance, then, near the hinder end of the head, dilates to form a hemispherical or conical saccule, which rests against and is partly embraced by a dilated tracheal tube, colourless and distinctly ringed as usual; from this a. trachea can be traced backwards into the body. This appears to be an “anterior spiracle” such as occurs in the pupa of several aquatic Diptera, such as the gnats and Eristalis, though in form it does not resemble the organ in the latter fly.
From each of the posterior spiracles, situated at the tip of each of the forks of the tail, a trachea can be traced forwards into the body, and there is no dilatation except the small one just referred to.
The Larva.
Amongst the dark-brown “puparia” I find one individual which is grey in colour, with a softer and flatter body; the spines covering the body are much longer than in the puparium; each of the ventral paired groups of “claws” is on a distinct prominence or transverse ridge, which appears to represent the paired “legs” of Eristalis, the ridge of segment 10 being more prominent than the rest. Further, round each segment, as well as on the “head,” is a series of tufts of small pale hairs, no doubt sensory in function. Each tuft consists of three hairs rising apparently from a common base; they are colourless, and much longer than the spines. These sensory hairs are also observable on the puparium, but are less conspicuous. The anterior end bears a pair of dorsal “eye-spots.” The mouth is armed on each side with two or three rows of stout spines, and within the cavity is a pair of short antennae; each appears to be 3-jointed, the terminal joint being brown, and to spring from a circular

convex disc, which I believe forms the anterior end of the real head, which is retracted, as has been observed to occur in Eristalis. The mouth leads into a short cylindrical tube, which soon dilates to form a pharynx, in the wall of which a number of sclerites are present, the most conspicuous of which is essentially a ring, with four backwardly directed processes, which no doubt serve for the attachment of muscles. Though built up on the same plan as that of Eristalis, it is very different in detail.
Some of the puparia were merely empty skins, and these were cleft horizontally on each side from the head backwards over two or three segments: no doubt the fly had emerged by this slit.
In one case the skin was ruptured somewhere near the middle of the body and exposed the pupa, which occupies only a small portion of this internal cavity; in length it extends through about four segments, and in breadth is only half the diameter of the puparium.
We have, then, the whole series of stages in the life-history of this creature, which I believe from the above—mentioned references to be Ephydra, both from its habitat in salt pools and from the resemblance in external form. Moreover, the genus has already been recorded by Captain Hutton as occurring in this country, for he described* E. aquaria from Christchurch, which is 5 mm. in length, and this agrees with the length of the pupa in the puparium above alluded to.
No doubt the general history agrees pretty well with that of Eristalis, and is as follows: The eggs are laid in the water of salt marshy ground; the larvae, issuing therefrom, burrow in the mud, where the spinous covering and the specialised claws serve as organs of locomotion, and the tail is directed upwards to the surface for respiration: this probably occurs only at intervals. The larva feeds on the organic matter in the mud, and as the time for pupation approaches it seems that it uses its hinder clawed segments 8 and 10 to cling to algae floating in the water, for in nearly all the individuals I find threads of algae held by these prominent segments; and at this time the anterior spiracles perhaps come into use. The skin now becomes hard and dark-coloured. The tissues, as in other cases of similar metamorphosis, become disintegrated, and later on become rearranged, if we may use the term, to give rise to the body of the pupa. The larval skin has become a cocoon or “puparium,” and it seems likely that the imago escapes through the anterior end—hence the object of the larva supporting itself by the algae, so as to allow the anterior end to emerge from the water.
[Footnote] * Trans. N.Z. Inst., vol. xxxiii., p. 90.

Explanation of Plate XVII.
| Fig. 1. |
Puparium of Ephydra, viewed ventrally, slightly from the right side; claw-bearing segments numbered. b, patch of black plates on segments 1. The spines covering the body are omitted from these figures. |
| Fig. 2. |
Side view of the hinder part of the body, showing the clasping-organ formed by the 8th and 10th segments. |
| Fig. 3. |
Ventral view, in outline, with part of the body-wall cut away to expose the pupa within, a, anterior triramous spiracle-bearing outgrowth; d, cavity within puparial skin; e, pupal envelope; f, imago within the pupal envelope, diagrammatically represented; g, paired group of claw-like spines on 8th segment; i, anus; m, mouth; p, pharyngeal skeleton (seen through the skin). |
| Fig. 4. |
Anterior end of a larva, showing the armature of the mouth and of the anterior margins of segments 1 and 2. a, anterior spiracular outgrowth; b, triangular group of spinose plates; m, mouth; n, the base of antennae (? = true head);'s, trifid sensory hairs. |
| Fig. 5. |
The mouth enlarged. showing the probably “true head” retracted within. n, one of the antennae; h, “true head.” |
| Fig. 6. |
Pharyngeal skeleton of the larva. m, mouth; p, pharynx. |
| Fig. 7. |
Anterior spiracular outgrowth, with its air-canals passing back to the trachea (t) At edge of figure is seen a few of the plates (marked b in fig. 4). |
Art. XXIV.—Notes on some Nudibranch Molluscs from New Zealand.
[Read before the Otago Institute, 11th October, 1904.]
Plate XVIII.
A Few years ago I sent a number of nudibranchs that I had collected at various places round our shores to the Danish specialist, Dr. R. Bergh. A description of these, and of others sent by Mr. H. Suter, has been published in an elaborate monographic serial, which is not likely to be available to naturalists in this colony; nor, indeed, would it occur to students of the New Zealand fauna to search the pages of Semper's “Reise im Philippinen” for information about new species limited, so far as is known, to our coasts. Nevertheless, in the sixth part of Bergh's “Malacologisch Untersuchungen,” which forms volume ix. of Semper's Expedition, may be found not only descriptions of our nudibranchs, but others from Tasmania and from Rarotonga; still others from Lamlash Bay, in Ireland; from the coast of Alaska; from South Carolina; and from Porto Rico!

In the interests of students of zoo-geography and local faunas, zoologists ought to protest against this sort of mixture of localities under a title which indicates, if words mean anything, that the contents were collected during an expedition to the Philippine Islands.
However, I have to thank Dr. Bergh for naming and describing these specimens, and for sending me proof-sheets of that part of his valuable “Untersuchungen” that deals with our species. It would have been useful to have reproduced his figures, but I understand from Mr. Hedley that the only figure of an entire animal is reproduced from a sketch of Atagema carinata that I forwarded to Bergh; the remaining figures refer only to differences of internal anatomy—many of them of microscopic detail.
In addition to six species the description of which is given in the translation below, Dr. Bergh named for me five other nudibranchs, of which one is Goniodoris castanea, Alder and Hancock, which is thus shown to have a very wide distribution, as it occurs on the British coast. Of the remainder, two new species of Chromodoris, a new species of Doriopsilla, and one of Aphelodoris were included in those I sent him, but, as I do not know whether he has yet published a description of these new species, I refrain from giving the specific names at present. These four new species and some of the others were obtained during the two trips of the s.s. “Doto” in which Mr. Ayson, Inspector of Fisheries, conducted a series of experimental trawlings in the years 1900 and 1901, when a considerable number of new and rare species of most groups of invertebrata were obtained.
Of the two new species of Chromodoris, one was obtained off the Mahia Peninsula, Hicks Bay, on the east coast of the North Island; the other between Kaipara and New Plymouth, where, too, the Aphelodoris was caught: these were carefully preserved by Mr. Hamilton. The Doriopsilla was dredged from 30 fathoms in Tasman Bay, in 1900, when Mr. G. M. Thomson was superintending the collection of invertebrates.
One of the most interesting of the molluscs described below is Homoiodoris novae-zealandiae, since the only other species of this genus occurs in the Japan Sea. We have several other similar facts of distribution amongst our marine invertebrata. I refer, for example, to the peculiar little enteropneust Dolichoglossus otagoensis, Benham,* with its grooved proboscis, which is only known in one other species (D. sulcatus, Spengel, from Japan). Again, the polynoid Physalidonotus squamosus, Qtfgs. (or Lepidonotus
[Footnote] * Trans. N.Z. Inst., vol. xxxiii., p. 90.

giganteus of Kirk) presents peculiarities, indicated by its generic name,* that have hitherto only been noted in two Japanese polynoids, recently described by Moore† as Lepido-notus branchiferus and L. chitoniformis, but which probably should be included in Ehler's new genus just mentioned. A second interesting form is Atagema carinata, which is now resuscitated from the oblivion that for seventy-five years has sorrounded it.
I proceed to give a translation of so much of Bergh's article as deals with the external features of our species (omitting details with regard to internal anatomy), and any remarks of my own will be enclosed in square brackets.
Nudibranchia Kladohepatica.‡
Fam. Tritoniidae.
Tritonia, Cuvier.
T. incerta, Bergh, n. sp., p. 24.
Of this form I have received from Professor Benham (Otago-University) a single specimen, obtained off the east coast of the North Island of New Zealand, between Kaipara and New Plymouth [during the trawling trip of the s.s. “Doto,” 1901].
[Colour.] The individual, in alcohol, was of a yellowish-white colour.
[Dimensions.] The length is 4cm., with a breadth (without the gills) of 1.5 cm., and a height of 1.3 cm. The breadth of the “buccal veil” is 10mm., its lobes 2.5 mm., and the tentacles 2 mm. The height of the rhinophore-sheath [hinder tentacles] is 4 cm.; the height of the gills is 4 cm.; breadth of the sole of the foot, 15 cm.; length of tail, 3 mm.
[Externals.] It has the form usual in the genus. The “buccal veil” carries altogether seven finger-shaped lobes. The tentacles have the usual form. The retracted rhinophores are 3.5 mm. high, with the usual structure. The back is finely tuberculated. The short arborescent gills, numbering 13–14 on each side, are nearly of equal size. The anus is situated below the 5th gill. The genital pore as usual. The foot is narrowed in front and behind; the margin relatively thin; the tail [meta-podium] short.
[p. 25.] This form can scarcely be identified with any of the Tritonias from the Pacific, hitherto sufficiently studied; it
[Footnote] * Ehlers, “Neuseelandische Anneliden,” Abhand. K. G. Wiss Got-tingen, 1904.
[Footnote] †Moore, J. P., Proc. Acad. Nat. Sci., Philadelphia, 1903, p. 405.
[Footnote] ‡Semper's “Reise im Philippinen,” ix. (Bergh, Malacol. Unitersuch vi., 1904).

is distinct from T. challengeriana and T. dioncedea, while from T. exsulans it differs in the buccal veil and the number of teeth at the radula.*
Nudibranchiata Holohepatica.
Dorididæ Cryptobranchiatæ.
Fam. Archidoridiæ.
Archidoris, Bergh, 1892.†
A. violacea, Bergh, n. sp. [p. 31]. PI. ii., figs. 36, 37; p1. iii., fig. 1.
This form was frequently obtained by Benham at a depth of 30–40 fathoms at various stations between Otago Peninsula and Oamaru. [I have obtained it in bucket-loads from fish-trawlers.] The living animal attains a length of over 6 in., and a breadth of 2½ in. The colour is a fine violet, with orange-coloured “tentacles” and foot.
[Dimensions.] The single specimen [that I sent him before I had ascertained its great abundance], when in alcohol, has a length of 4 cm.; breadth, 2.5 cm.; height, 1.7 cm. The mantle-edge is 5 mm. broad; the foot is 1.7 cm. broad, its margin 3.5 cm. broad; the tail is 5 cm. long. The height of the rhino-phore-sheath is 1 mm., and of the rhinophore itself 7 mm. The diameter of the entirely protruded gill is 1 cm.; length of individual branches, 5 mm. Height of anal papilla, 1 mm.
[I am ignorant as to the value or otherwise of these measurements for identification. It seems to me that many of them depend upon the mode of preservation. If slowly killed, with the various parts fully extended—as can be done by means of weak alcohol, cocaine, and so forth—the measurements will be very different from those taken on an individual of the same size killed by plunging into corrosive sublimate or strong alcohol. However, I give these details as they appear in the memoir.]
The colour of the preserved specimen is throughout yellowish-white. The back is covered all over with relatively closely set slightly protuberant rounded tubercles of whitish colour, and with a diameter of 4 mm. [These tubercles in life are violet.]
[Externals (p. 32).] The form is a longish oval, the back fairly arched. The tubercles towards the margin of the back are smaller and more crowded. The rhinophores are closely foliated. The gills, situated far back, are formed of eight tri-pinnate members, of which the hindmost are slightly smaller.
[Footnote] * Bergh, Opisthobranchen. Rep. of “Albatross,” Bull. Mus. Comp. Zool., Harvard, xxv., 1894, p. 150.
[Footnote] † It may be as well to state that this name replaces the term Doris, which is no longer used as a molluscan genus.

The anal papilla is nearly in the centre of the branchial crown. The mantle-flap* is of nearly the same breadth throughout; its under-surface smooth. The foot is rounded in front, with well-defined marginal groove; the margin is not very narrow; the tail short.
[Remarks (p. 33).] From its coloration this form probably represents a new species; possibly it is identical with one or other of the “species” recently described from the Pacific Ocean.†
[I confess I do not understand the above remark. If it is merely its coloration that distinguishes it, it seems scarcely probable that that would justify the formation of a new species. I may note that Sir G. Eliot, in his account of the “Nudibranchs of East Africa and Zanzibar,”‡ suggests that his A. africana may be identical with this species, and A. umia with A. nanntla. It is rather odd that Bergh does not refer to Abraham's paper in the Proc. Zool. Soc. 1877, in which several opisthobranchs from our coasts are described and figured. It is evident that all our nudibranchs require working over carefully, for it seems likely that the littoral forms have received more than one name. It will be seen, later, that in the case of other species there is the same hesitation or doubt as to its novelty or otherwise. Unfortunately, the valuable memoirs that Bergh has been publishing for some years past are very much too expensive for individual workers to purchase, and none of our libraries possess the work.]
A. nannula, Bergh, n. sp. [p. 33], p1. iii., fig. 2.
Of this form Benham has forwarded seven individuals, obtained at Port Chalmers, Otago Harbour. [I collected some dozens of this bright little species when the s.s. “Ringarooma” was in dry dock, after being at moorings for about a year in Deborah Bay, amongst the seaweeds, Bryozoa, sponges, &c., adhering to the bottom of the ship. I collected a considerable number of interesting animals, amongst others the Goniodoris castanea, referred to above.]
[Colour.] During life they are light—orange—coloured. In alcohol the seven individuals agree pretty well in colour, size, and form. They were entirely yellowish-white.
[Dimensions.] The length averages about 18mm., with a breadth up to 14mm., and a height of 3 mm.; the breadth of
[Footnote] * By this I translate “Mantelgebram.”
[Footnote] † R. Bergh: Chall. Repr., p. 85; Die Opisthobr., Alaska, 1894, p. 159 (A. kerguelensis); Die Opisthob. d. Sammulung Plate, Zool. Jahrb, Suppl., iv., 3, 1898, p. 501 (A. rubescens: A. (?) incerta); Ergebn. Reise. n.d. Pacific (Schauinsland), Zool. Jahrb., xiii., 3, 1900, p. 222 (A. nyctea).
[Footnote] ‡P.Z.S., 1904.

the mantle-flap is about 14mm. [sic.: probably a misprint]. The height of the gills is 4.5 mm.; breadth of the foot, 7 mm.
[Externals.] Form as usual. The back is entirely covered with almost hemispherical papillae, not very variable in size, but somewhat smaller at the mantle-edge. The club of the rhino-phore is much foliated, the foliae stiffened in the usual way with long spicules. The branchial region is in all the individuals much everted, and surrounded by the narrow lip of the distended gill-opening (which measures 6 mm. in diameter). The gill is formed of five to seven tripinnate members: the upright anal papilla completes the branchial circle posteriorly. The under-surface of the mantle-edge is smooth; the tentacles appear to be shortly conical; the foot is fairly broad, rounded in front, with a short tail.
[Remarks (p. 34).] This form is undoubtedly an Archidoris, but whether it represents a distinct species must remain for the present undecided.
Archidoris, sp. [p. 34].
The single specimen obtained by Benham at Kaikoura, on the east coast of the South Island, was, in life, pale-orange in colour.
[Colour.] The specimen in alcohol was much hardened, broken, somewhat contracted, and uniformly light-yellowish.
[Dimensions.] Length, 23 mm.; breadth, 16 mm.; height, 10 mm. Breadth of mantle-flap, 3 mm.; of the foot, 13 mm. The height of the rhinophore, 2 mm.; the diameter of the circular branchial aperture is 4 mm.; height of the gill, 3 mm.
The animal was somewhat hard and stiffened; the back fairly smooth, but finely granulated towards the margins; the rhinophores, lying far forward, has its club much foliated. The foot is large; its margin not broad; the tail short.
[Remarks (p. 35).] This form is probably identical with one of the recently described species of the genus.
Homoiodoris, Bergh, 1881.
Hitherto only one species of this genus, H. japonica, has been described, and this from the South Japan Sea.
H. novae-zealandiae, Bergh, n. sp. [p. 35], p1. iii., figs. 3–7.
Of this form five individuals were received from Mr. H. Suter, who collected them at Port Chalmers.
[Dimensions.] The specimens preserved in alcohol had a length varying from 12–20 mm. In the largest individual the breadth of the body is 13 mm.; the height, 7 mm.; the width of the mantle-flap is 3.5 mm.; that of the border of the foot is 3 mm.: the length of foot is 16 mm.; its breadth, 7 mm.;

the tail, 2 mm. The height of the (retracted) rhinophore is 2 mm.; length of tentacle, 1 mm.; height of the gill, 3 mm.; diameter of the branchial star, 7 mm.
[Colour.] The colour of the back is whitish or faintly yellowish white; the tubercles white; the head and foot inclining to yellowish; the clubs of the rhinophore and the gill are chrome-yellow. One individual had the under-surface of the mantle-flap spotted with violet-grey.
[Externals.] In consistency the animal is somewhat stiff. The form as in other Archidorids. The back is covered pretty closely with tubercles of dissimilar size, somewhat flattened on the apex (at least in the case of the larger ones): they attain a height and diameter of about 0.5 mm. Similar tubercles are also present at the edge of the rhinophore-pits and gill-aperture. [p. 36] The club of the rhinophore bears about 20 foliae; the branchial aperture is transversely oval; the gills number 5 or 6; the cylindrical anal papilla is relatively high (1 mm.). The under-surface of the mantle-flap is smooth; the sides of the body angular. The foot has a distinct groove at its anterior edge; the tail is short; the tentacles are short, thick, with grooves on the under-side.
[Remarks (p. 37).] Whether this form belongs to the genus Homoiodoris remains for the present undecided, since the most essential character—the armature of the vagina—was not established owing to the condition of preservation. If this armature noted [in the text] belongs to the sperm-duct, either a new genus will have to be formed, or it belongs to the genus Artachae,* in which, however, no prostate is developed. The vestibular glands and sac existing in the present species are absent from Homoiodoris and from Artachaea. From the hitherto known species of Homoiodoris, as well as of Artachaea, this new form is, however, specifically distinct, as is evident from the comparison of the radula—the innermost as well as the outermost teeth being quite unlike those of other species.
Atagema, Gray [p. 39].
Gray, fig. iv., 1850, p. 104; Guide, i., 1857, p. 209.
Form and internal structure as in an Archidorid. The back carries a median keel. As was so frequently the case, Gray erected a new genus (Atagema) on a drawing (by Quoy and Gaimard) of an animal presenting a somewhat peculiar appear-ance.
The animal described below belongs to the Archidorididae: it has the general body-form of the family, with granulated
[Footnote] * See Bergh, “System d. Nudibranch: Gasteropoda,” 1892, p. 1093.

back and a widish mantle-flap, with tri- or quadri-pinnate gills, small tentacles, and rather stout foot. The oral disc (Lippen-scheibe) is unarmed; the radula has a narrow naked rhachis, and numerous hook-like plates. Within this large family of Dorids it approaches most nearly the genus Archidoris. It shows well the short finger-shaped tentacles and the relatively small number of gills. The penis and vagina are both unarmed. Apparently the only peculiarity about the animal is the dorsal keel, and it-must for the present remain doubtful whether Gray was justified in founding a new genus on this character alone.
Atagema carinata (Q. and G.). Doris carinata, Q. and G., Voy. de 1'Astrolabe, ii., p. 254, pl. 16., figs. 14–16. Atagema carinata, Gray, fig. iv., p. 104. [Bergh figures it] pl. iii., fig. 8–11.]
An individual of this species lies before me, obtained by Dr. Benham from Dusky Sound, at the south-west corner of the South Island. The living animal is pure-white. The sketch made by Benham (fig. 8) is from an individual preserved in formol.
[I exhibited a specimen of this Dorid at a meeting of the Otago Institute in 1898,* and drew attention to the fact that it was the “D. carinata” of Quoy and Gaimard. Since sending the specimen to Dr. Bergh, I have received from Mr. R. Henry, at Dusky Sound, others of a larger size—viz'., length, 8–9 cm.; breadth, 6–7 cm.; height, 3.3 cm.
The specimen in alcohol is much arched, contracted, and hardened; it had been broken. It is now yellowish-white in colour.
[Dimensions (p. 40).] The length was [in an extended condition] probably 5–6 cm., with a breadth of 4 cm., and a height 2.5 cm. The dorsal keel has at its greatest length a breadth of 5–7 mm.; it is narrower in front and behind, while posteriorly to the protuberance its height is 5 mm. The width of the mantle-flap is at least 12 mm., and its thickness at the base is 7 mm.; the height of the rhinophore-sheath is 4 mm., and of the branchial papilla 4 mm.; the breadth of the foot is 22 mm., and of the margin of the foot 4 mm.
[Externals.] The consistency is leathery; the back is quite finely shagreened. The form is oval, with a somewhat arched back; the mantle-flap broad, and the foot projecting all round. The back carries a great median keel, which commences anteriorly between the rhinophores. It is at first somewhat low and narrow, then becomes thicker, and behind the middle
[Footnote] *See Trans. N.Z. Inst., xxxi., p. 743.

of its extent becomes elevated into a blunt point; thence it continues as a much narrower ridge to the gills. The projecting rhinophore-sheath has a circular aperture. The club of the retracted rhinophore is 4 mm. high, is closely foliated, and terminates in a small papilla. The branchial cone consists of four converging lappets (5 mm. high) with rounded ends. The deeply sunk gill is formed of six* tripinnate members, 7 mm. high, of which the two anterior are the larger. The relatively thin anal papilla, which is 3 mm. high, is curved forward, and subcentral in position. The retracted tentacle is 2 mm. high, short, finger-shaped with a groove on the underside. The foot is stout and broad, rounded in front, with a marginal furrow; the tail is relatively short.
[Remarks (p. 41).] The animal here described appears really to represent the Doris carinata of Quoy and Gaimard, which since the expedition of the “Astrolabe” in 1828 has not been met with. The French zoologists obtained the animal in New Zealand [in the estuary of the Thames, Hauraki Gulf]. The size (tres petite espece) of the individuals examined by them appears, however, to have been very much less [than the specimens now under consideration], the colour yellowish-white, and the number of gills only four.
[In view of the interest of this species I append figures of a specimen, natural size, in side view and from below (see Plate XVIII.).]
Art. XXV.—Note on the Function of the Last Pair of Thoracic Legs in the Whale-feed (Grimothea gregaria).
[Read before the Philosophical Institute of Canterbury, 30th November 1904]
Plate XIX.
The whale-feed, Grimothea gregaria (Fabr.), which is so common around the southern coasts of New Zealand at certain seasons of the year, belongs to a division of the Crustacea generally known as the Anomura, a group intermediate in many characters between the Brachyura (crabs) and the Macrura (crayfishes, lobsters, &c.). In the Anomura the thoracic legs of the fifth pair are generally small and weak, and thus markedly different, from the preceding pairs of legs. In giving the characters of
[Footnote] * In specimens in my possession I find the branchial cone to consist of five lappets or valves, and the gill consists of five members (Benham).

the Macrura anomala, a group including the majority of the Crustacea generally classed as Anomura, the Rev. T. R. R. Stebbing says,* “The fifth pair of legs are generally weak, not fiteither for walking, swimming, or grasping food or prey.” In many of the free-swimming forms, such as Grimothea, these legs have the joints of the limb folded against one another like the limbs of the letter Z, and the whole appendage is carried at the side of the carapace above the bases of the more anterior legs, and in this position they have the appearance of being quite useless; and I am not aware that any function has ever been assigned to the fifth pair of legs in this section of the Anomura, though in another family, the Lithodidm, where the fifth pair of legs are slender, chelate, and folded in the branchial chambers, Mr. Stebbing has suggested that they may be used to keep the branchial clear of parasites and thus be of advantage to the animal.†
During a short stay at the Marine Fish Hatchery and Biological Station at Portobello, in November, 1904, I had opportunities of observing living specimens of Grimothea gregaria, and one day while watching a small specimen under the dissecting microscope I was much interested to see the animal suddenly unfold the fifth pair of legs, stretch them forward over the anterior portion of the carapace, and with the tuft of setse on the terminal joints carefully brush away extraneous matter from the dorsal surface of the carapace, and particularly from the spaces between the rostrum, the eyes, and the bases of the antennae. The action was quite unexpected, and was almost ludicrously like that of a person engaged in brushing his back hair. For the purpose in question the fifth pair of legs of Grimothea gregaria seem well fitted—they are just long enough when unfolded to reach conveniently to the anterior portion of the carapace; and the two terminal joints (propodos and dactylus), which are bent nearly at right angles to the preceding joint, and when at rest are curved behind the bases of the fourth pair of legs, are supplied with numerous long setse projecting radially from them, one row of setae being curved and pectinate, and the whole forming a sort of circular brush specially adapted for sweeping out the spaces between the spines of the rostrum, around the, bases of the eye-stalks, &c. These two joints, moreover, form a chela or pincers with the fingers somewhat spoon-shaped, and are doubtless used to pick off substances that cannot be brushed away.
As the Grimothea gregaria swims rapidly backwards by means of alternate flexions, and extensions of its abdomen,
[Footnote] * “History of Crustacea,” p. 149. 21—Trans.
[Footnote] † l.c., p. 155.

any floating matter in the water would naturally tend to settle in these spaces, and would interfere with the proper use of the eyes; and there seems little doubt that the fifth pair of legs are of advantage to the animal in keeping these spaces clear, and in preventing parasites lodging therein.
It is comparatively seldom that one has an opportunity of observing the habits of marine animals, and the use that is made in this case of a pair of appendages that have all the appearance of becoming vestigial is perhaps worth placing on record.
Explanation of Plate XIX.
| Fig. 1. |
Grimotha gregaria, fifth thoracic leg (× about 15). |
| Fig. 2. |
Grimothea gregaria, extremity of the same (more highly magnified). |
Art. XXVI.—On the Occurrence of a Species of Cercaria in the Cockle.
[Read before the Philosophical Institute of Canterbury, 30th November, 1904.]
Plate XIX.
During the month of November, 1904, I spent a short holiday at the Portobello Fish-hatchery and Biological Station, and on the morning after my arrival the curator, Mr. T. Anderton, brought me a cockle (Chione stutchburyi) evidently infested with some parasite. On examination this proved to be the sporocysts of some species of Distomum containing numerous Cercarice, in most cases just ready to escape from the sporocyst. Subsequently the sporocysts were found in two or three other specimens of the cockle, but, though numerous in those specimens in which they were found, they were not often met with—certainly not in 1 per cent, of the cockles examined.
Similar Cercarice have been long known to occur in various molluscs in Europe, but so far as I am aware the only forms hitherto recorded from New Zealand are the two described by Professor Haswell from Mytilus latus;* and, as my specimens differ from these and from all the others that I can find descriptions of, I give here a brief description of its general form, mode of occurrence, &c. It appears to come very near to Cercaria villoti, Monticelli (= C. setifera, Villot),† but differs from that
[Footnote] * Proc. Linn. Soc. N.S.W., 1902, p. 497.
[Footnote] † See Korscheldt and Heider, “Embryology of Invertebrates,” part i. (English edition), p. 186; and Villot, “Annales des Sciences Naturelles,” Zool., viii., p. 33. (I am indebted to Professor Benham for these references.

species in the character and arrangement of the setse on the tail, and I propose to give it the name Cercaria pectinata.
Cercaria pectinata, n. sp. Plate XIX.
Similar to C. villoti, Monticelli (= G. setifera, Villot), but with the “setse” arranged in two rows along the sides of the tail, instead of forming rings around it; each “seta” consisting of a short vertical row of long bristles.
Hob. In Chione stutchburyi, Otago Harbour.
General Notes.
The sporocysts (fig. 1) were numerous in each cockle in which they were found, particularly in the first one, in which there must have been several hundreds. They were lying external to the body in the spaces between the mantle-lobes, the gills, &c. Each sporocyst is about 4 mm. long and 0.75 mm. broad, sausageshaped, but somewhat narrowed at each end, both ends being alike. By alternate contractions and expansions they were capable of creeping about somewhat slowly with a worm-like motion. The whole of the interior of each sporocyst was in nearly all cases filled with fully developed Cercarice just ready to escape, and through the semi-transparent walls of the sporocysts these Cercarice could be seen often moving about within. The number of Cercarice in each sporocyst varies to some extent, but usually there were a dozen or more. Swimming among the sporocysts were numerous Cercarice that had already escaped, and if a sporocyst were artificially ruptured the contained Cercarice at once began to swim freely about. In a few cases the Cercarice were seen escaping naturally through the walls of the sporocyst, and in these cases they made their way through the lateral wall and were apparently able to penetrate this at practically any point.
Occasionally the sporocysts contained, besides fully developed Cercarice one or two rounded or elongated masses of cells, presumably immature Cercarice, but in most of the cases observed they contained nothing but fully developed Cercarice.
The Cercaria (fig. 2) has a long powerful tail nearly twice as long as the head; this tail sometimes showed numerous fine transverse lines or striae, most evident anteriorly, but these were not made out in all cases. Along each side of the tail is a series of “setae,” and by the rapid vibrating action of the tail, combined with the use of these “setae” as fins, the Cercaria is able to swim rapidly through the water. When viewed from above during life each “seta” (see fig. 3) appears to be simple, but closer observation shows that each is made up of a number of fine long bristles arranged in a short vertical row and united together at the base so as to form a comb-like structure, the

teeth of which are, however, very long. These bristles are held together by fine transverse connections, and when in action during life are kept in the one plane and moved as a single structure, but in preserved specimens they become separated and spread out towards the distal end. The tail is also capable of considerable contraction and expansion, and its proportionate length and width accordingly varies greatly, but in the condition most usually maintained the width is about one-eighth of the length.
The head is usually more or less rounded when the Cercaria is swimming freely, but in addition to this mode of motion the Cercaria can creep along by means of the worm-like motions of the head caused by contraction and expansion, and accordingly the head then assumes other shapes and may be much elongated. In fig. 2 the anterior portion is shown in the act of contracting and consequently widening, while the posterior portion is still elongated. The general colour of the whole Cercaria is whitish, semitranslucent, but on the head there are usually a number of small irregular pigment-spots of a light-brown colour.
The general structure of the Cercaria appears to present the usual features: the buccal sucker is situated at the anterior extremity, and is the foremost portion when that part of the body is being extended; the posterior sucker is about equal in size to the anterior and is situated a little anterior to the middle of the head; in fig. 2 it appears rather nearer the anterior end owing to the contraction of that portion of the head. There is a well marked cuticle, and around the anterior sucker this bears a number of fine short setae. The internal organs are-well developed; during life the most conspicuous is a large V-shaped structure extending from the posterior extremity of the head nearly to the anterior end, and presumably representing the contractile bladder of the excretory system; in life this is darker than the rest of the head, and is filled with minute granules. The central portion of the tail presents much the same appearance, but whether the bladder actually extends into the tail or not I am not certain.
The alimentary system is also well developed and of the usual structure; the pharynx is small, nearly circular, and situated close up to the buccal sucker; the oesophagus narrow and short, and the two lateral branches of the intestine are also narrow.
The reproductive organs also appear to be well developed in most of the Cercarice examined, and in one which was accidentally ruptured while under examination I saw mature spermatozoa; in some the genital aperture could be seen about midway between the two suckers, and connected with it an oval structure extending back as far as the posterior sucker and apparently representing the rudiment of the genital atrium. I have, however, not

attempted to make out the arrangement of the different parts of the reproductive system. In specimens killed in osinic acid two small masses staining deeply with the acid are usually very prominent, one on each side, a little behind the posterior sucker.
As the Cercarice on leaving the sporocyst may readily escape from the cockle and swim freely in the sea-water, it may thus reach a great variety of animals, and in the meantime I am unable to offer any suggestion as to the host in which the adult fluke is likely to be found.
Explanation of Plate XIX
Cercana pectinata.
| Fig. 1. |
Sporocyst containing Cercarice (x about 10). |
| Fig. 2. |
Cercaria pectinata (× 60). |
| Fig. 3. |
Cercaria Pectinata: one of the “setca” from the tail (highly magnified). |
Art. XXVII.—Notes on Fruit-fltes, with a Description of a New Species (Dacus xanthodes).
[Read before the Auckland Institute, 27th February, 1904.]
These two-winged flies are, without doubt, the most dreaded of all orchard pests, and consequently strenuous efforts are made to prevent their establishment in New Zealand. The most important species from our point of view are the Mediterranean fly (Halterophora capitata) and the Queensland one (Tephritis tryoni).
The Mediterranean species is known to occur in widely distant countries, including Malta, Sicily, Azores, Madeira, Cape Verde Islands, Bermuda, Cape Colony, Mauritius, Cuba, and Australia. European records of its ravages extend as far back as the year 1826, when it was stated that “fully a third of the oranges shipped to London from the Azores were rendered unfit for use before they reached their destination through the presence of this maggot before they were packed.” In 1890 an article appeared in the American publication called “Insect Life” wherein this fly “was described, as a peach-pest at Bermuda. During 1892 Mr. J. H. Cooke stated that” all the oranges in Malta had been destroyed for the past few years by the maggot of this fly.” The opinions of competent observers and responsible officials only have been quoted here. Besides the fruits already mentioned, about fifteen others have been attacked.

The Queensland fly was, I believe, first recognised in that colony in 1887, but it was not described until 1897, the date of the report prepared by Mr. W. W. Froggatt for the Government of New South Wales. This insect in Australia does not appear to breed further south than Newcastle, New South Wales—about our 33rd parallel of latitude; but it has been plentifully found here in fruit imported from the tropical islands situated northeast of New Zealand.
Before proceeding further it may be stated that, relying solely on the results of the inspection of fruit landed at Auckland, the Mediterranean fly has not been reared here except from maggots found in fruit imported from Sydney, none at all having been bred from “island” fruit. As this species thrives in comparatively cold climates, and attacks almost all kinds of fruit, we must carefully guard against its introduction.
When dealing with importations of fruit infested with different kinds of insects an Inspector must apply special methods for their destruction before delivering such fruit to importers. In the case of scale insects, which are exposed on the outside of the fruit, the whole consignment is subjected to the action of hydrocyanic-acid gas within an air-tight chamber for an hour. This treatment destroys the insects, but does not remove their scales or waxy coverings. The fly-maggots, on the other hand, cannot be destroyed by artificial means, unless by burning the fruit itself, as the maggot until mature or full-fed is secure within the fruit, there being only a minute hole through its skin formed by the ovipositor of the female fly whilst inserting her egg. When an Inspector detects the presence of these maggots in a case of fruit, all the other cases forming that particular consignment and branded with the same mark are soon afterwards burned in a furnace. As many as 450 cases belonging to one lot and consigned to one importer have been destroyed in that way, thereby causing a serious loss to the importer or shipper. The only exception to this rule applies to the islands of the Cook Group, recently annexed to New Zealand. In this instance only a consignment found to be infested with these maggots may be picked over under the Inspector's supervision. Each fruit is examined separately: if infested it is put aside for burning, but those found to be free from this pest are handed over to the importer, who therefore only loses the infected fruit.
During the year from 1st April, 1902, to the 31st March, 1903, there were ninety-one different lots, comprising 4,119 cases, of fruit condemned at Auckland alone on account of these maggots. All these consisted of “island” fruit.
The maggot of the new species (Dacus xanthodes) does not differ materially from those of the other flies specially alluded to.

It was discovered here in one pineapple imported from Rarotonga on the 5th December, 1903, and on the 14th of that month Mr. G. Harnett, Assistant Inspector, and I again detected it in two cases of pineapples from Suva. We also found it on various occasions afterwards in oranges, grenadillas, and mammae-apples from Tonga and Rarotonga. The perfect flies were subsequently reared here from the maggots in considerable numbers, so that this new pest threatens to become as troublesome and injurious as the Queensland fly.
Dacus (Tephritis) xanthodes, sp. nov.
Imago—Length of body, 4½ lines; expanse of wings, 7½ lines. Body elongate, yellow, occasionally testaceous, extremity of abdomen blackish, head sometimes rufescent, tarsi infuscate; on the thorax, from base to apex, there is a central pale ivorylike streak, along each side there is a similar one; these lines after death become less conspicuous.
Head as broad as the thorax, smooth; on its back part there are four black outstanding setae, and between the eyes six finer ones. Antennas normal, their terminal joint elongate and rather darker than the others; at the tip of the preceding one arises a very long seta, which, though stout at its base, becomes very slender and darker towards the extremity. Eyes large, prominent, their inner edges straight and moderately distant from each other; they are finely facetted, and of a brilliant purple during life. Thorax cylindrical or subovate, nearly twice as long as it is broad, with two slight almost equidistant constrictions at each side; the surface bears numerous minute black granules, from each of these proceeds a fine dark hair; at the base, which is deeply emarginate, there are two long rigid conspicuous black setee directed horizontally backwards, four smaller ones are situated just before the posterior constriction, and about an equal number along each side. Hind-body quite the length of the thorax, its basal three segments, irrespective of the narrowed anterior portion, of the same length and breadth, thus forming a cylindric figure, fourth segment rather shorter and narrowed behind, the terminal elongate and tapering towards its apex, with very fine grey hairs; the preceding segments are minutely sculptured and bear many dark slender hairs.
Legs stout, moderately elongate, with short black hairs; posterior tibise somewhat arcuate. Tarsi elongate, pilose, their basal joint rather more than half of their entire length, fifth bilobed; claws black and rather small.
Halteres yellow, medially slender, clavate and triangular at the extremity.

Wings hyaline, unspotted, with pale-brown nervures corresponding in structure with those of Tephritis tryoni.
Underside flavescent, not maculate, except at the apex of the last ventral segment, which is piceous. The abdominal segments are concave, or so deeply hollowed that the sides and upper surface appear quite thin.
Female.—Hind-body elongate-oval, terminal segment rounded nd not prolonged; on each side of the uncovered second segment there is a small cluster of fine black setse: these do not occur in the other sex.
This species differs from Froggatt's Tephritis psidii in being larger, differently coloured and sculptured, and without dusky areas on the wings. From T. tryoni it is distinguished by the greater length of the body and expanse of wings, uniform coloration, without fuscous or yellow marks, dissimilar clothing and sculpture, less broadly oviform or wasp-like hind-body, and stouter antennal setse. In T. tryoni the flanks of the sternum are fuscous.
Art. XXVIII.—On some New Species of Lepidoptera.
[Read before the Wellington Philosophical Society, 2nd November, 1904.]
Plate XX.
Melanchra decorata, n. sp. Plate XX., fig. 2.
♂, ♀, 38 mm. Head and palpi grey-yellowish. Antennae simple in both sexes. Thorax with strong anterior bifid crest, less pronounced in ♀, grey yellowish mixed with brown; patagiae outwardly bordered with brown; a white line on collar followed by a black line. Abdomen with series of crests, grey -yellowish. Fore wings: Costa almost straight, termen slightly waved and faintly sinuate, pale brownish-yellow (in some examples tinged with pink) mixed with whitish on basal half of costa; a very dark reddish-brown shade from middle of base to termen above tornus, bordered beneath from about ⅓ by conspicuous white fascia which is bent sharply upwards at ¾; a short brown streak from base below middle; orbicular oblique, faintly outlined in brown; reniform large, pale, outlined in brown; space between orbicular and reniform dark reddish-brown, connected with costa by narrow projection; two brown spots on costa above reniform; a faint double waved line from costa at ¾; subterminal indistinct, pale, waved, bordered with bright reddish-brown; veins faintly

marked with brownish; cilia brownish-yellow, suffusedly barred with dark-brown. Hind wings fuscous, tinged with pink, lighter near base; cilia grey with dark line.
Very distinct; it is perhaps allied to M. diatmeta, but is not likely to be mistaken for that species. The single male I possess is not in good condition, and does not show the markings to the same extent as the female. The species is figured from a ♀ in fine condition lent to me for the purpose by Mr. Robt. Gibb, of Tuturau.
A rare species, taken in October at “sugar.” West Plains, Tuturau.
Melanchra levis., Plate XX., fig. 4.
♂, ♀, 35 mm. Head and palpi dull greenish. Thorax with strong thick pointed anterior crest, greenish-brown; collar broadly yellowish-brown; patagia brighter green. Abdomen dull - brownish, slightly greenish - tinged. Fore wings: Costa straight; apex rounded; termen straight, rounded beneath, brown faintly tinged with green, markings very obscure, blackish; a thin double line near base; a suffused patch at angle of dorsum near base; a faint curved line at ⅓, followed by oblique interrupted line not reaching dorsum; a fine bent line at½ posteriorly bordering orbicular and becoming obsolete at middle of wing; a similar line anteriorly bordering reniform; subterminal line pale-greenish, suffusedly and interruptedly bordered with blackish; cilia pale-greenish mixed with brown. Hind wings greyish, fuscous, paler towards base, cilia grey with obscure darker line
Belongs to the smaller exquisita-coeleno group, but does not closely resemble any species.
Taken at “sugar” sparingly in September and October. West Plains.
Leucania stulta, n. sp. Plate XX., fig. 1.
♂, ♀, 41 mm. Head, palpi, thorax, and abdomen pale-fawn; dark line on collar, suffusedly bordered with whitish. Antennae brown, basal portion whitish, shortly ciliated in male. Fore wings slightly dilated; costa uniformly arched; apex roundpointed; termen slightly oblique, rounded; pale-fawn; markings dark reddish-brown; a short streak from near base beneath to ¼; a suffused irregular streak from middle of base to termen above anal angle, almost interrupted before middle; a suffused streak from ⅓, attenuated anteriorly and much dilated towards termen, divided on lower portion by whitish borders of veins; one or two short apical streaks above this; cilia brownish on termen, fawn beneath. Hind wings fuscous; cilia whitish with dark line.

Not easily compared with any other Leucania.
October to December. Not common. I have not taken the male at West Plains, and have figured that sex from a specimen kindly supplied by Mr. Gibb, who has taken a few examples at Tuturau.
Leucania neuree, n. sp. Plate XX., fig. 5.
♂, 36 mm. Antennae strongly dentate, fasciculate; brown, with white spot at base. Head and thorax dull ochreous-brown. Abdomen pale greyish-brown. Fore wings: Costa straight, apex slightly rounded, termen straight, rounded beneath; dull-brown; veins sharply marked in black with scattered white scales; a white spot on vein 1b at¼; a conspicuous white spot at origin of veins 3 and 4; a curved series of white dots on veins from about ⅘ of costa to ⅘ of dorsum; cilia same colour as wing, tips whitish. Hind wings dull-fuscous, paler at base and dorsum.; cilia pale greyish-white with darker line.
Distinguished from L. micrastra, its nearest ally, by the smaller size, duller colour and paler cilia of the hind wings. Leucania toroneura differs from this species in the much lighter ground-colour and the absence of white spots.
Two males taken at “sugar”; date uncertain. West Plains.
Xanthorhoe imperfecta, n. sp. Plate XX., fig. 6.
♂, ♀ 29 mm. Head, thorax, and abdomen brownishochreous. Fore wings ochreous, in male often reddish on costa and about apex. The usual lines are in the male reduced to irregular blackish costal spots; in the female these costal spots give rise to very faint transverse waved lines. White spots follow most of the dark costal spots, and there is a fairly conspicuous pair at ⅔; cilia ochreous, barred with blackish. Hind wings ochreous; cilia greyish-ochreous mixed with darker.
Nearest to X. ceyrota, but easily distinguished by the brighter ground-colour and the white costal spots.
Fairly common in low-lying bush districts in January. West Plains; probably generally distributed in suitable localities.
Xanthorhoe recta, n. sp. Plate XX., fig. 3.
♂, 31 mm. Head, thorax, and abdomen ochreous. Fore wings moderate, greyish - ochreous, irregularly suffused with reddish especially towards costa, markings brownish; first line before ⅓, curved, anteriorly edged with grey; median band formed of three or four thin lines with narrow spaces of groundcolour between; the first of these is almost straight and rather darker than the others; the posterior edge of the last one is irregular with two strong projections below middle; this is followed

by a narrow band of pale-greyish, almost white in some examples; a broad band of brown from ⅘ of costa to tornus posteriorly waved and followed by a waved pale line; cilia long, brownish-pink. Hind wings long, narrow, pale-yellow; a terminal series of indistinct linear brownish spots; cilia pink.
Apparently allied to the clarata group, but not closely approaching any species.
Received from Mr. J. H. Lewis, Ida Valley. Mr. George Howes has also met with it near Dunedin.
Art. XXIX.—Notes on the Occurrence of some Rare Species of Lepidoptera.
[Read before the Wellington Philosophical Society, 6th July, 1904]
Dodonidia helmsii, Butler.
It will, I feel sure, be of interest to entomologists to hear that this beautiful butterfly appears to be on the increase. In February of 1903 Mr. H. W. Simmonds, of Wellington, took fifteen specimens on the birch-clad range above Silverstream, and saw some twenty-five or thirty on the wing. He tells me that it is a strong flier, and is very evasive of capture. It appeared to be very local in its habitat, and frequented for the most part the outskirts of a small patch of birch forest on the very ridge of the hill. Mr. Simmonds found it impossible to pursue it through the fern and scrub, but by posting himself at the opening of a glade in the bush he netted most of his specimens as they flew in, to seek, apparently, the shade. In February of the present year he again visited the same locality, but this time was less fortunate, for since the previous season the patch of bush that had been so productive had been partly destroyed by fire, and though he saw a few he was unable to capture any. To some extent this might have been accounted for by the fact of its being dull, cold weather and a little late in the month. He met with one on the low ground at the foot of the range, but with this exception they appeared to keep to the high level. I took one at the latter end of the same month at Lake Papaitonga, Ohau (Manawatu), on the flowers of the white Escallonia. It is the only occasion I have seen it on the wing, and I was much struck by its beauty as it hovered over the blossoms.
In passing, I would strongly recommend collectors to plant a few trees of the Escallonia (Escallonia montevidensis) in their gardens, as it possesses a singular attraction for insects of all kinds. This tree in particular, and which I had planted for the

purpose, seems, when in flower, to be the “Mecca” of the insect world in the neighbourhood. Throughout the day it is alive with Vanessa gonerilla, Chrysophanus salustius, C. enysii, Lycœna phœbe, Nyctemera annulata, and many varieties of Diptera; at dusk it furnishes a harvest of Noctuœ. It comes into blossom about February and flowers freely till the end of March; it grows readily, and is of hardy habit.
Of the seven specimens of Dodonidia in my collection, four were taken by Mr. Simmonds at Silverstream, one was taken on the Wainuiomata Range, one at Papaitonga, and another on the high lands in the Marlborough District.
Chærocampa celerio, Linn.
I think this will be the first record of the appearance of this handsome moth in New Zealand, and it will, I hope, take its place in our list of Sphingidœ, at present only represented by S. convolvuli. In March of the present year Mr. Creagh O'Connor took two very fine specimens at Titahi Bay (about fourteen miles from Wellington), and during the same month saw some ten or twelve others there. The two he netted were taken at dusk while feeding at the sweet-scented Christmas lily; they appeared to affect garden flowers generally, but were difficult to capture, being very active on the wing, and when once alarmed would not return.
Mr. G. V. Hudson tells me that he has recently received a specimen from Nelson. Noting the fact that it has appeared at two places on the West Coast, it is not unlikely to be an Australian species brought over to New Zealand by westerly winds, in view of the fact that the hawk-moth family are possessed of sustained powers of flight; indeed, I might mention that I have in my collection a fine Sphinx that flew on board the R.M.S. “Ruahine” when the vessel was some five hundred miles off the coast of South America. It is to be hoped that this species will become established here, for with its bars of gold on the thorax, its silver-striped upper wings, and the delicacy of its pink underwings, it will certainly be a very handsome addition to our list.
Sphinx convolvuli, Linn.
To the best of my knowledge, this species has so far been confined principally to the Auckland District, where I have taken it freely at the blossoms of the evening-primrose and trumpet-flower, but it appears now to be having a wider range. Five specimens were brought to me at the end of this summer, all taken in Wellington and its environs. Mr. O'Connor has also taken it, and I learn, too, that it has been seen freely in Nelson, and also at Ashburton (Canterbury).

Achæa melicerte, Meyr.
The first and, I believe, the only record of this moth in New Zealand appears in the Transactions of the Philosophical Society in 1876,* in an article contributed by the late Mr. Fereday. He states that a specimen was taken at Wellington by Mr. Liardet, and he describes and figures it as Catocala traversi. In his paper he mentions that one had been taken at Lyttelton two years previously, and that he understood it to be a common moth amongst the gum-trees in Australia.
Mr. Hudson tells me that, as far as he knows, this is the only record of its appearance, and until recently these have been the only two specimens known. (Mr. Meyrick has described it in the Transactions† as Achœa melicerte, but does not mention Catocala traversi as a synonym. From his description, however, it seems clear they are one and the same, and I have adopted Mr. Meyrick's nomenclature.)
It is now interesting to note that it has occurred somewhat plentifully at Titahi Bay, where, early in March this year, Mr. O'Connor took no less than eight or ten in one day, and saw as many more. He was collecting on a steep hillside shelving down to the beach, over which a fire had some time previously run, destroying most of the scrub but leaving here and there a few isolated bushes. On beating these the moths flew out, and, not knowing it to be such a rarity, he kept only five or six. He tells me that they would fly for fifteen yards or so and then settle, but would be off again on his approach. The day being very bright, this alertness rather points to the fact that they may be diurnal in their habits. A little later in the month he took one specimen at rest in a garden in Wellington.
The occurrence of this moth after being practically unknown for nearly thirty years is a matter of much interest and conjecture.
Utetheisa pulchella, Meyr.
This daintily coloured species has also been taken at Titahi by Mr. O'Connor. In January last he took ten in one day, some being at flower on the white rata, and the others he netted in the tussock-grass. He states that it has a feeble flight, and is easily captured. This was the only occasion on which he saw it.
The only New Zealand specimen I have seen is in Mr. Hudson's collection, and was taken by him at Wainuiomata in 1886. A specimen was taken at Petone by the late Mr. Norris; and these, I believe, until now, are the only local captures.
[Footnote] * Trans. N.Z. Inst., vol. ix., p. 459.
[Footnote] † Vol. xix., p. 37.

Art. XXX.—Notes on the Entomology of Mount Holdsworth, Tararua Range.
[Read before the Wellington Philosophical Society, 1st June, 1904.]
During the middle of February of this year I spent, in company with my wife, a fortnight in the vicinity of Mount Holdsworth. The object of this and two previous expeditions undertaken by me was to investigate the insect fauna of this well-known range of mountains; and, although the results at present arrived at cannot be regarded as complete, it is perhaps desirable that they should be placed on record, more especially as, so far as I am aware, the Tararuas have not yet been visited by other entomologists.
The Tararua Range is, of course, very familiar to all residents in Wellington, its snow-capped summits, which close in the head of the Hutt Valley, being a most beautiful and conspicuous feature in the landscape in winter-time. These peaks constitute the southern portion of the range, and Mount Holdsworth, which is situated further to the north, is hidden by them. The range is best approached from the other side—i.e., from the Wairarapa—and intending visitors should take the early train to either Carterton or Masterton, and thence proceed by trap to the junction of the Waingawa and Mangatariri Rivers. From this point there is a fair road for about four miles up the latter river, and a good camping-ground may be found at the termination of the road. To adequately explore the mountain it is necessary to make a permanent camp here, so that suitable weather may be selected for the ascent, as the summit of the mountain is nearly always enveloped in cloud. This is no doubt due to the fact that this range is situated in the centre of the southern portion of the North Island, and, being the highest land in the neighbourhood, attracts large quantities of cloud whenever there is the slightest atmospheric disturbance. In fact, much cloud is attracted with any wind, either north-west or south-east. For entomological or viewing purposes, the mountain should not be attempted except during the passage of the crest of an anti-cyclone—i.e., when the barometer is at its highest at the termination of a southerly wind, before the change to the north-west, the sky being, of course, absolutely clear at the time.
I have dwelt at some length on the meteorological aspect of the question, as it is a most important one. In fact, on two previous visits which I made to this locality, both a week in duration, I was unable to ascend the mountain, and the expeditions were almost fruitless in result, owing to unfavourable weather conditions,

although at the time the weather in other places would have been fairly satisfactory for entomological work.
From the above-described camping-ground at the end of the road, the top of the mountain may be reached after about six hours' hard climbing. The time and labour now necessary could, however, be reduced by at least one-half were a satisfactory track made through the bush, and this could no doubt be done for a very small sum. In the interests of botanists, entomologists, and others, it is surely possible that something may be done in this direction, when the exceptionally fine view which may be obtained from this mountain, its extreme richness in alpine plants, and its proximity to Wellington are all taken into account. Some steps ought to be taken to render such an interesting locality more readily accessible, and, in default of other means, a portion of the Research Fund of this Society might perhaps be so employed in thus aiding original biological research in the wilds of New Zealand.
The ascent of the mountain is not severe, the sole difficulty in the undertaking being due to the dense bush and undergrowth. There have been bush-fires from time to time at several points on the track, which have greatly increased this difficulty, and it is in these places that the track is so extremely difficult to follow. These fires in forest reserves, such as this, are much to be regretted, and any persons lighting such fires ought to be very severely punished. The land here is quite unsuitable for settlement, and hence the ranges have, I understand, been very wisely set aside as a forest reserve. The reserve is reached about one mile beyond the termination of the road, and at this point the Mangatariri River has to be forded. This is easily accomplished in fine weather, and, whilst fording, a beautiful view of a primeval forest stream of the purest water may be obtained. The track continues alongside the stream through very fine forest for about another mile, when an ascent of about 1,000 ft. brings us to what is called the “lower camp,” which is situated in the midst of wrecked bush, the site of what has evidently been one of the most disastrous fires on Mount Holdsworth. After this is passed the track passes for another mile through subalpine bush, 2,500 ft. There is a beautiful carpet of native grasses, and the trees are profusely festooned with long pendant mosses. For the next mile or so the track descends about 300 ft., traversing a broad swampy spur covered with low brushwood, chiefly manuka and birch. The vegetation is, however, largely subalpine, the mountain kiki and many grasses and mosses being frequently met with. At the end of this spur the “upper camp” is reached, which is situated at an elevation of about 2,300 ft., and is at the foot of the steep spur which leads to the summit of the mountain.

After this the track is very steep, and at about 3,800 ft. the birch-trees, which have become, as usual, very gnarled and stunted and thickly covered with lichens and mosses, suddenly end, and the open grassy country of the high mountain is reached, this elevation apparently representing the usual line of permanent snow.
I have seldom seen a mountain so richly covered with alpine plants as Mount Holdsworth, and I feel satisfied that it would be a locality of extreme interest to a botanist. Amongst many others, I observed the following familiar plants in great profusion: Mountain - lily (Ranunculus insignis?), spear - grass (Aciphylla squarrosa and A. colensoi or allied species), various species of Celmisia, &c. The only introduced plant I noticed was the “cape-weed” (Hypochœris), which was in profusion at about 4,000 ft. The exceptionally efficient means of dispersal with which its seeds are endowed no doubt explains its appearance so high on the mountain.
From the bush-line to the top of the mountain the distance is about three miles, and the ascent is very easy. The view obtained from the top is very fine, embracing Mount Egmont to the north-west, the Island of Kapiti, the Straits, and a considerable part of the northern portion of the South Island, including Mount Tapuaenuku, to the west, the opposite side of the ranges which close in the head of the Hutt Valley could be recognised to the south, and the long, low ranges of hills on the east coast, and beyond these the ocean could clearly be seen.
During my visit I made two successful ascents of Mount Holdsworth. On one occasion the weather was absolutely perfect, on the other somewhat cloudy and too cool for many insects to be about. It was, however, evident that I was too late in the season for many of the high alpine species, so that further visits are necessary earlier in the year, and the second week in January would probably be about the best time to find the greatest number of species.
I specially looked out for the two mountain butterflies, Erebia pluto and Erebia butleri, but could see nothing of them, and feel sure that, had they been present, some specimens would have been in evidence. I should mention, in support of this contention, that these butterflies are found as late as the middle of March on the mountains in the South Island, where they occur. I was not, however, altogether surprised at the absence of these insects, as they have never yet been recorded from the North Island. The same remarks apply to Argyrophenga antipodum, which is exclusively an alpine butterfly in the provinces of Nelson and Marlborough, but is found abundantly elsewhere in the South Island, both on mountains and in tussock country.

I was, however, much more surprised in not finding any species of Crambus or Orocrambus on Mount Holdsworth, and do not think that the total absence of both these genera, which are characteristic of all the mountains I have ever visited in the South Island, can be explained by the lateness of the season, though future visits must definitely determine this. In connection with the distribution of these forms, it would be interesting to know whether geologists have yet determined the northern limit of general glaciation in New Zealand. The absence of these characteristic alpine insects from the Tararuas may perhaps have some bearing on this question, as without the glaciation of the intervening lowlands such species might not have been able to travel from the mountains of the South Island to those of the North Island. The Tararua Range is an important one in connection with this inquiry, as it is the nearest high mountain-range to the South Island.
I will now give a list of the more interesting species observed on Mount Holdsworth and in its neighbourhood, with some special remarks on the rarer species met with. It must, however, be clearly understood that this list does not claim to be an exhaustive one, but subject to revision when the locality has been more adequately worked by entomologists.
Lepidoptera.
Vanessa gonerilla.
This butterfly was very abundant on Mount Holdsworth, from the bush-line to the summit. The specimens were very large, and in perfect condition. They, however, exhibited no divergence from the usual type.
Chrysophanus salustius.
A few specimens seen in the Mangatariri Valley.
C. boldenarum.
A distinct variety of this insect occurs in river-beds on the Wairarapa Plain.
Lycœna phœbe.
Mangatariri River; sparingly.
Nyctemera annulata.
Two specimens of this very common insect occurred on Mount Holdsworth at 4,000 ft. Had they not been actually netted, they might readily have been mistaken for one of the Erebias, and reported as such.
Leucania griseipennis.
At “sugar,” Mangatariri River. Rare.
L. purdii.
At “sugar,” Mangatariri River. Three specimens only.

L. atristriga.
At “sugar,” Mangatariri River. Extremely abundant.
L. unipuncta.
At “sugar,” Mangatariri River. Rare.
Melanchra insignis.
At “sugar,” Mangatariri River. Rare.
M. plena.
At “sugar,” Mangatariri River. Three specimens only. Not quite typical.
M. vitiosa.
At “sugar,” Mangatariri River. One only.
M. composita.
At “sugar,” Mangatariri River. Not so common as usual.
M. ustistriga.
At “sugar,” Mangatariri River. A few only.
M. lignana.
At “sugar,” Mangatariri River. Common.
Bityla defigurata.
At “sugar,” Mangatariri River. One only.
Cosmodes elegans.
One superb specimen, amongst grass near camp. I have not taken this species previously.
Tatosoma agrionata.
One specimen, in forest. Probably too late for this insect.
T. lestevata.
I secured a magnificent female specimen of this extremely rare species at “sugar” on 26th February; my other two specimens, both males, were taken, one at Nelson in 1885, and one at Wainuiomata in 1887. There are two specimens in the collection of the late Mr. R. W. Fereday, now in the Christchurch Museum. I am not aware of any others.
Elvia glaucata.
One specimen, in forest. Passed.
Hydriomena subochraria.
Very common in the Mangatariri Valley. This species is characteristic of the locality, as it is not a generally common species.
Asthena pulchraria.
In forest. A few specimens. Passed.
A. schistaria.
In forest. A few specimens. Passed.

Venusia undosata.
One specimen seen. Passed.
Asaphodes megaspilata.
Common as usual.
Xanthorhoe semifissata.
A few seen. Passed.
X. clarata.
Two specimens taken on Mount Holdsworth at 4,000 ft. Passed.
X. beata.
In forest. Passed.
X. œgrota.
One specimen only. Mangatariri River.
X. cinerearia.
Common as usual.
Dasyuris partheniata.
This fine and conspicuous insect was very abundant on Mount Holdsworth from 3,800 ft. to 5,000 ft.
Notoreas mechanitis.
Several taken on Mount Holdsworth at about 4,500 ft.
N. paradelpha.
Several taken on Mount Holdsworth at about 4,500 ft.
N. omichlias.
Extremely abundant on Mount Holdsworth from 3,800 ft. to 5,000 ft. This was much the commonest insect seen. It is a rare species on the mountains in the South Island.
N. brephos.
Some very small and vividly marked specimens were captured by Mrs. Hudson, in the river-bed of the Mangatariri.
Epirranthis alectoraria.
One specimen, at “sugar.”
Leptomeris rubraria.
Common as usual. Mangatariri River.
Selidosema fenerata.
Common as usual. Mangatariri River.
S. rudiata.
Common as usual. Mangatariri River.
S. suavis.
Common as usual. Mangatariri River.
S. productata.
Common as usual. Mangatariri River, and in birch forest up to 3,000 ft. A white variety of the female taken at “sugar.”

S. melinata.
Common as usual. Mangatariri Valley.
S. panagrata.
Common as usual. Mangatariri Valley.
S. dejectaria.
Common as usual. Mangatariri Valley.
Sestra humeraria.
Common in forest as usual.
Gonophylla fortinata.
One specimen, in forest.
Declana floccosa.
Common as usual.
Crambus ramosellus.
Common as usual. Mangatariri and Waingawa Rivers.
C. flexuosellus.
Common as usual. Mangatariri and Waingawa Rivers.
C. vitellus.
Common as usual. Mangatariri and Waingawa Rivers.
C. apicellus.
On swampy spur on track to Mount Holdsworth. Common.
C. xanthogrammus.
One specimen taken by Mrs. Hudson, Mangatariri River.
Mecyna marmarina.
A large pale variety of this species was common on Mount Holdsworth from about 3,800 ft. to 4,500 ft.
M. flavidalis.
Common as usual.
Scoparia philerga.
A few worn specimens in forest.
S. acharis.
A few worn specimens in forest.
S. hemicycla.
In stunted birch forest, Mount Holdsworth, 3,600 ft., common, but rather worn; evidently too late for it. This is a rare and interesting species.
S. characta.
One specimen seen at “sugar,” Mangatariri River.

S. epicomia.
In forest, Mangatariri River.
S. feredayi.
In forest, Mangatariri River. A few only, in poor condition.
S. crypsinoa.
Mount Holdsworth, 4,000 ft. Worn.
S. sabulosella.
Common as usual. Mangatariri River.
S. steropœa.
One seen. Mangatariri River.
S. astragolota?
Mount Holdsworth, in forest, 3,500 ft.
S. asterisca.
At “sugar,” Mangatariri River.
Nesarcha hybreadalis.
One seen. Mangatariri River.
Musotima aduncalis.
Common as usual. Mangatariri River.
Platyptilia haasti.
On swampy spur track to Mount Holdsworth. Locally common.
Pterophorus lycosemus.
Common in forest, Mangatariri River.
P. monospilalis.
Common in forest, Mangatariri River.
Clepsicosma iridia.
I took two specimens of this species in the dense swampy forest near the Mangatariri River.
Capua semiferana and Noteraula straminea.
Taken commonly by Mrs. Hudson amongst tutu, Mangatariri River.
Adoxophyes conditana.
In forest, Mangatariri River.
Simaethis.
Three specimens of this interesting genus occurred on Mount Holdsworth, from 3,600 ft. to 4,000 ft., as usual. These insects were flying very rapidly in the hottest sunshine.

Neuroptera.
The following species of Neuroptera were observed. The season was, however, too far advanced for these insects, and a visit earlier in the year would no doubt yield better results. The four common species of dragon-flies were abundant—i.e., Uropetala carovei, Somatochlora smithii, Lestes colensonis, and Xanthagrion zealandicum.
Stenosmylus latiusculus.
I secured seven specimens of this very rare insect at “sugar” in the Mangatariri Valley. They usually appeared singly each night, but the last night I was at the camp I secured four, two of them arriving at the “sugar” after 9 p.m. This species was described by Mr. McLachlan in 1894 from two specimens, one taken at Waitara and the other at Greymouth. I captured my first specimen in the Orongaronga Valley, to the east of Wellington Harbour, in January, 1892, and have not since met with the insect until this year. So far as I am aware, the above include all the captures which have been made.
Œconesus maori.
One specimen, at “sugar,” Mangatariri River.
Polycentropus puerilis.
At light, camp Mangatariri River.
Oniscigaster—– sp.?
One nymph taken in river in January, 1902.
Atelophlebia.
Several species observed.
Coleoptera.
This order was only worked in a desultory manner.
Mecodema scitulum.
Several specimens taken under stones on lower spurs of Mount Holdsworth, January, 1902. The forest at the foot of the mountain appears poor in beetles, but half an hour's beating yielded a fine anthribid.
Orthoptera.
Two or three fine species of grasshoppers occurred on Mount Holdsworth at elevations of 3,800 ft. to 5,000 ft., and an interesting stick insect on manuka - bushes at the “lower camp” (2,000 ft.), none of which have yet been determined.

Art. XXXI.—Epalxiphora axenana, Meyr.: a Species of Lepidoptera scarce in New Zealand.
[Read before the Wellington Philosophical Society, 1st June, 1904.]
Plate XXI.
This species of Lepidoptera I found of considerable interest. It occurs throughout New Zealand from early summer to winter, but seems to be scarce everywhere, though its relative scarcity probably depends to some extent upon climatic conditions. During the summer of 1901–2 sunshine was continuous throughout, and Epalxiphora axenana occurred in considerable numbers in the strip of sparsely wooded bush reserve where I had previously taken the species but sparingly. I do not know where the males rest, but have beaten them from brown dry leaves of tree-ferns, and also from the green leaves of the food-plant. I do not remember to have ever seen a male resting in an exposed situation; the females, however, rest exposed on the upper side of a leaf of Piper excelsum (its food-plant), where the coloration is rather conspicuous, but the shape of the insect with its wings closed is not mothlike—i.e., not conspicuously so. The position taken up by the females is not accidentally due to the drying of the wings on emergence from the pupa, which takes place during the afternoon. The insect flies at dusk, whereas my specimens have always been taken during the forenoon, and from two so taken I procured ova which were fertile, proving the insects had been flying the previous night.
Mr. Meyrick has twice published his diagnoses of the genus Epalxiphora.* His description of the species was made from a single ♂ specimen taken off a tree-trunk in Wellington, New Zealand. The sexual dimorphism of the species makes descriptions of the ♀ ♀ a necessity. The costa of anterior wings is in males curved, in females elbowed; the anterior-wing markings differ between the sexes: ♂ pattern is divided into basal and outer areas, with a characteristic buff apical tip; ♀ pattern is composed of transverse and longitudinal markings usually, but this sex is very variable. A bright-orange basal streak on costa of posterior wings ♂ (covered by anterior wings) is not present in ♀. The only mark of anterior wings which appears to be present in both sexes is a crescentic or angular mark about the middle of the inner margin, which, when the wings are closed together edge to edge over the back, forms a characteristic ocellated spot.
[Footnote] * Meyr., Proc. Linn. Soc. N.S.W., 1881, 648; Trans. N.Z. Inst., xvii., 147.

Epalxiphora axenana. Fig. 8.
25 mm. Forelegs dark-fuscous, ringed ochreous at joints; head above fuscous mixed with grey; thorax dark or light fuscous, tufts pitchy. Abdomen ochreous (anal segments testaceous) or wholly yellowish-grey. Anterior wings curved from base, apex falcate, outer margin obliquely sinuate. Basal area grey broadly mixed with dark fuscous dots to ⅓ costa, whence edged with indefinite dark fuscous transverse line to middle of wing, then with inner tooth and outward curve to½ inner margin it includes the marginal lunar mark, which is completed by a faint curve and dot; towards the inner margin the basal area shades to brownish - grey. Outer area dark - fuscous towards costa, reddish-brown or delicate violet hue towards inner margin; an elongate transverse costal spot of either colour at½ runs into discal black line, sometimes dividing it into two angular spots, in front a yellow or buff tip obliquely elongates to apex. Cilia ochreous with dark fuscous bars and basal line. Posterior wings grey or lighter with dark posterior mottling, cilia white with very fine dark-grey basal line.
Six ♂ ♂, others not retained: except in ground-colour and presence or absence of the violet hue there is no variation.
Epalixiphora axenana, ♀. Fig. 1.
27–30 mm. Head and thorax usually of same colour as ground-colour of wings, tufts pitchy only when some of the wing-markings are so. Abdomen yellowish - white. Anterior wings: Costa elbowed at ⅓, thence slightly curved to apex, outer margin obliquely sinuate, apex falcate. Ground-colour very pale-yellowish (bone-colour), at base mixed with dark dots, near inner margin suffused with greenish-fuscous. At ⅙ costa a dark olive-green transverse elongate spot to discal cell, below is a paler spot, and an indefinite longitudinal line angulated or curved to inner margin. At ⅓ costa a transverse greenish fuscous spot curved outward unites with a patch of similar colour covering the whole anal area, extending from before middle of wing longitudinally with irregular outline to near apex, its transverse outline deeply broken by the ground-colour forming the curve of lunar mark at½ inner margin. Nervures posteriorly thinly fuscous. A dilated sinuate costal streak of greenish-brown colour from ⅓ costa to near apex breaks the ground-colour into a longitudinal pattern. Cilia pale reddish-grey barred with fuscous. Posterior wings: Apex more acute than ♂, margin somewhat sinuate. Light-grey with posterior yellow tint, dark longitudinal mottling, cilia whitish, basal line fuscous.
I have selected this as the type form, it being the most abundant, though there is considerable colour - variation, chiefly as follows:—

α. Anterior wings: Ground-colour pale-yellow with greenish hue, markings pale-green, inner marginal lunar mark surrounded with dark-fuscous. Not common.
β. Anterior wings: Ground-colour reddish-yellow, markings ferruginous, the whole therefore having a reddish hue. Not common.
γ. Anterior wings: Ground-colour as type but less distinct on lower area, markings dark - fuscous without greenish hue, anal patch coppery-brown. A very handsome form; not common.
The following I prefer to term “aberrations” rather than “varieties,” since one needs to breed from ova a larger proportion of any given form to term it a definite variety, otherwise subspecies.
Ab. albo-suffusa, n. ab. Fig. 2.
Anterior wings: Markings longitudinally divided from inner base to apex, the line of division deeply dentate at ⅔; ground-colour white above, at ⅙ costa a dark greenish brown elongate transverse spot; a very faint costal mark at ⅓, costal streak at ⅔ to near apex is fuscous mixed with lighter; lower half similar to type of markings; colour fuscous posteriorly dusted with white, nervures thinly marked, marginal lunar mark distinct. Some specimens are suffused with white over a larger area restricting the anal patch, the dark costal mark at ⅙ normal; inner margin has thin fuscous basal streak to lunar mark wider beyond and shading to grey at outer margin. Some have inner colour coppery.
Ab. brunnei-lineata, n. ab. Fig. 3.
Anterior wings: Dark fuscous streak extends from base to below apex, with a parallel line above it of reddish-yellow ground-colour heightening the linear effect, being suffused with fuscous on costa, with normal fuscous costal marks; below the colour is suffused with dark - fuscous from base to lunar mark and beyond. Posterior wings grey, with posterior reddish hue. One specimen
Ab. purpurascens, n. ab. Fig. 4.
Anterior wings: Pattern composed of two colours separated by an almost straight dividing-line from base to near apex; costal area is unicolorous, pale-yellow (bone-colour), costal marks faintly dusted in grey; lower area is purplish-red, marginal lunar mark scarcely perceptible. Cilia reddish-yellow. A specimen has pale fuscous costal marks, lower area of wing deep-purplish-black with some indistinct markings. Rare; three specimens.

Ab. obsoleta, n. ab. Fig. 5.
Anterior wings: Ground-colour pale-yellow with an orange suffusion at base, longitudinally and posteriorly; no other markings. One specimen.
Ab. obscura, n. ab. Fig. 6.
Anterior wings wholly covered with fine sheeny dark-fuscous dots obscuring all markings except faint costal marks at ⅙ and ½, and faint lunar mark; apex of anterior wings tipped pitchy; thoracic tufts pitchy. One specimen.
Ab. nigra-extrema, n. ab. Fig. 7.
Anterior wings black, costal area to ⅓ light-fuscous with black dots, and spot at ⅙; fuscous mottling at ⅔ in middle of wing, similar mottling from lunar mark to anal angle. Several specimens, one having purplish-red ground-colour instead of black.
The female deposits a circular semitransparent mass numbering some twenty-nine ova, symmetrically imbricated, closely appressed. If laid on a green leaf, reflection of the natural tint of the leaf through the ova must afford greater protection than any colour of the egg-shell could do. Ova deposited 22nd January, 1902, hatched 5th February = 14 days. Ova deposited 20th February, 1903, hatched 5th March = 13 days.
Ovum (figs. 9, 10, 11): Each ovum is oval in outline, slightly convex above and covered with a rather large crystalline pattern composed mostly of pentagonal but some hexagonal figures. Examined some hours after they were deposited each ovum had inner and outer circumferences, the pale-greenish egg-contents not reaching the outer wall of the ovum; the micropyle, forming a rosette of small elongate figures, is situated towards one end of the ovum on this rather wide marginal area. The ovum is always so placed in relation to the egg-mass that the micropyle is outwards and not covered by other ova. The first ovum laid forms the centre of the egg-mass to the disadvantage of the ensuing larva, which may—as it did in one instance—fail to eat through the overlapping edges of surrounding ova.
For two days the ovum contained only yelk spherules and the protoplasmic fluid whence the embryo ultimately develops: no cellular activity was observed.
In seven days the embryo reached an advanced stage, having formed, though neither caput nor thoracic segments could be detected, and at the head area, towards the micropyle, a large quantity of outer (greenish) spherules remained; the ocelli of the caput were distinct, the thoracic legs exceedingly long, outline

of the abdominal segments quite distinct, abdominal feet and claspers well developed, even the anal comb could be distinguished; the outer spherules about these parts were almost used up. The number of developing embryos within the egg-mass reminds one of a dish of opened oysters. Two days later internal anatomy was well formed, and all external tubercle setæ and the jaws of caput becoming chitmous.
On the eleventh day the caput was very distinct, assuming at first a faint violet hue which gradually darkened to brown, the jaws appearing bright-reddish; spherules immediately in front of caput were not quite used up. No movement of jaws, but internal pulsation was observed. The larva does not emerge through the micropylar area.
The newly hatched larva does not eat the egg-shell. Young larvæ feed on the under-surface of a leaf beneath a few threads of silk; later two leaves are drawn together, or, failing this, the leaf is folded over. Piper excelsum is the staple pabulum of Epalx. axenana: the leaves are broad and succulent. In normal seasons it is difficult to find leaves of P. excelsum which are not riddled with holes: one suspects these are made by slugs. The larvæ of Epalx. axenana are easily alarmed and drop to the ground; they are seldom found feeding between leaves which have holes in them: the slugs or whatever cause the holes are probably responsible for a high mortality among them by alarming them away from their food. During the genial season referred to very few leaves of P. excelsum had holes in them (it was a very dry season), and the larvæ were plentiful on that plant, and on several other plants and shrubs on which I never found them at any other time.
The larva (figs. 12, 13, 14) when newly hatched has a very dark-brown head, otherwise it is transparent pale-greenish. The head is flat-elongate, and the mouth parts protrude forwards; tapering somewhat anteriorly and posteriorly, abdominal segments 1 to 3 are widest, and the 9th narrowest. The whole skin is covered with long spicules; all setæ are smooth. Prothorax has no scutellum; on either side of medio-dorsal line are three setæ anterior marginal and three setæ posterior semimarginal, all equidistant. Posterior spiracle circular, prespiracular tubercle bears three setæ, tubercle above legs two setæ. Abdominal segments: Trapezoidals normal, i. seta short, ii. seta longer, supra-spiracular single long seta, spiracle circular placed on a midlateral swelling, the subspiracular is anterior and higher than the post-subspiracular tubercle, single seta each, basal tubercle bears two setæ. Abdominal feet have a single row of dark widely spaced hooklets, about twelve in number.
The larva in its second stage is very pale-greenish colour,

head yellowish with a suspicion of mottling. The mouth-parts are normal, not protruding as in previous stage; spinneret is short and stout. Prothorax as previous stage. Mesothorax: One below the other are two setæ in a dorsal depression, below which a large swelling bears two similar setæ, a lower posterior swelling bears one seta, an anterior swelling bears two setæ; tubercle above legs bears two setæ. Abdominal segments: All tubercles bear single seta, i. and ii. normal, iii. above the spiracle, which is well sunk into a lateral depression, anterior subspiracular almost below spiracle, post-subspiracular quite beneath, vi. anterior, basal setæ three in number. Abdominal feet have a single row of closely placed terminal hooks.
The adult larva is semitransparent green, with no appreciable markings except on the head, which is yellowish with characteristic brown mottling on each lobe. External structure as in second stage.
The length of a larva at fifteen days is 10 mm., after which it grows rapidly; at thirty days it is 28 mm., remaining attenuated in breadth. Duration of larval existence, thirty to thirty-two days.
The skin has a reticulation. From the centre of each figure—hexagonal or what not—of which it is composed a small boss rises tipped with a spike: these spicules persist throughout the larval existence, and practically cover the whole skin; but around the base of setæ a space exists without the spicules. The anal comb is seen with difficulty in a newly hatched larva, when the prongs are like setæ. In an adult larva (figs. 15 and 16) the comb consists of eight strong prongs each terminating in two points; at either side are incipient prongs. The prongs rise from a small pad on the underside of anal flap. Judging from the direction of the prongs in different specimens examined, the comb may be elevated or lowered at will. The function of the comb is probably in connection with the removal of excrement, and the position of comb is best seen when the anal flap is distended at the passing of same. Spicules are numerous in the region of the comb.
I once found three pupæ each in a leaf of Urtica ferox (tree-nettle). These leaves are covered with stinging spines, and had been rolled into a cylinder. It is doubtful whether the larvæ fed on the Urtica leaves.
The pupal shroud is made by the larva first with an outer series of silk threads apparently placed irregularly, but designed to hold together the two leaves or the fold of the leaf and prevent any alteration in the curvature which might subsequently affect the pupa disadvantageously. Within this outer series of threads a definite closely woven elongate cocoon is made,

having a slender neck which extends to the edge of leaf or leaves. The pupa is suspended horizontally within the body of the cocoon. In emergence the pupa projects rather more than the thoracic segments beyond the neck of the cocoon, a silken cable secured to its anal armature preventing it from overpassing the point of security and falling to the ground. I did find one pupa, which had either slipped its cable or broken it, with its anal armature amongst the outer silk at the mouth of the cocoon; the pupa, having passed quite out of the cocoon, was empty when found.
The pupa at first is unicolorous, green with darker green mediodorsal line on abdomen. The first colour-change is noticed in the eyes, which become red, then a reddish colour spreads over the face-parts, next the thorax and wings show imaginal markings. Duration of pupal stage, seventeen to twenty-nine days.
The pupa is 9 mm. long, 3 mm. wide at mesothorax, anterior end rounded; the segments taper dorsally and ventrally from mesothorax; the 9th abdominal is ventrally rounded, 10th produced to a long blunt point carrying two lateral hooks on each side, two ventro-posterior hooks, two posterior hooks—in all eight hooks, which are not the modified prongs of the larval anal comb, but a distinct pupal armature. Wing-cases extend to the posterior edge of 3rd abdominal segment; antennæ follow the costal curve from back of eyes to tips of wings; proboscis and two pairs of legs fill the space to antennal tips. Abdominals 2 to 7 carry two rows of dorsal spines. Some portions of the segments have a reticulation similar to that on the larval skin, but without bosses or spikes except in the region of the obsolete anal orifice, where a minute point appears in the centre of many figures of the reticulation.
On dehiscence the head separates and carries the antennæ, which are separated throughout their length from the other organs; leg-cases remain attached at their tips; wing-cases are loosened at their suture with abdomen. The dorsum of thorax splits centrally.
The imagines of first brood were all males with one exception, a ♀ type like its parent. Imagines of second brood were mostly males, but there were two typical females, and one like the parent, type form β.
Parasites on Epalx. axenana: Frequently the one true home of the larva of Epalx. axenana is occupied by a spider; sometimes a living larva and a living but very small, possibly juvenile, spider have occupied the same roof-leaf, and pupæ have been similarly situated. Sometimes the pupa contains the grub of a parasite which assimilates the internal anatomy

of its host, and emerges from the head of the pupa-case a fly—an Ichneumon, Triclistus, Forst., sp. nov., of which Mr. Claude Morley says, “certainly very closely allied to Trilistus nigritellus, Holmyr., from which it differs mainly in its unicolorous antennæ, dark tegutæ, black hind tibiæ, and larger size.”
Explanation of Plate XXI.
| Fig. 1. |
Epalxiphora axenana, ♀ type (nat. size). |
| Fig. 2. |
" ♀ Albo-suffusa, n. ab. (nat. size). |
| Fig. 3. |
" ♀ Brunnei-lineata, n. ab. |
| Fig. 4. |
" ♀ Purpurascens, n. ab. |
| Fig. 5. |
" ♀ Obsoleta, n. ab. |
| Fig. 6. |
" ♀ Obscura, n. ab. |
| Fig. 7. |
" ♀ Nigra-extrema, n. ab. |
| Fig. 8. |
" ♂ type. |
| Fig. 9. |
Ova, imbrication, and ovum at two days (× 75). |
| Fig. 10. |
Ovum at seven days (× 75). |
| Fig. 11. |
Micropyle, and sculpturing of ovum (× 250). |
| Fig. 12. |
First larval stage, dorsa of posterior segments (× 250). |
| Fig. 13. |
" dorsum of caput (× 250). |
| Fig. 14. |
" 3rd abdominal segment, lateral view (× 250). |
| Fig. 15. |
Second larval stage, anal comb, anal flap distended (× 250). |
| Fig. 16. |
Ultimate stage, anal comb, ventral view (× 50). |
Art. XXXII.—On the Occurrence of Graucalus melanops, Latham, in New Zealand.
[Read before the Philosophical Institute of Canter bury, 2nd November, 1904.]
On the 11th June last the Museum received a specimen in the flesh of the shrike-thrush from Mr. E. A. Radford, of Gebbie's Valley, who stated that he had picked it up dead on Rabbit Island. The bird is in the young plumage, and proved to be a male.
The first recorded occurrence of the species in New Zealand was a specimen in the Nelson Athenæum, which had been shot in an apple-tree at Motueka in 1869 or 1870. Another specimen was shot near Invercargill on the 8th April, 1870. Both were in the immature plumage. I was also informed by the Hon. W. Mantell that he had seen one at Port Chalmers in 1842. Mr. W. T. L. Travers also told me that he had seen the bird at his run at Lake Guyon, and Captain Fraser told me that he had seen it at Lake Hawea. In all these instances the birds appear to have been in immature plumage also. This seems very remarkable.

Graucalus melanops.
G. melanops, Vigors and Horsford; Buller, “Birds of New Zealand,” 1st ed. p. 148, 2nd ed. vol. i. p. 66; Cat. Birds in British Museum, vol. iv., p. 30. Colluricincla concinna, Hutton, Cat. Birds of New Zealand, p. 15 (1871).
Light ash-grey; abdomen and lower tail-coverts white; a broad band of black from the bill through the eye. The two middle tail - feathers dark - grey, lateral ones brownish-black, tipped with white. Quills brownish-black, the outer webs edged with white. Bill and legs black. Length of the wing, 8 in.; of the tarsus, 1.1 in. In the adult bird the forehead, sides of the face and neck, the throat and fore-neck, are black, with a greenish gloss.
The immature bird from Australia is described as having wavy bars of dusky black on the throat, but there are none on any of the three New Zealand examples which I have seen.
Art. XXXIII.—A Rare Saurian.
[Read before the Auckland Institute, 12th September, 1904.]
From time to time over a number of years reports have been brought in by surveyors, bushmen, and others of the existence of a large lizard on the Waoku Plateau. This is an extensive tableland, about 2,000 ft. above sea-level, lying between the Hokianga and Kaipara districts. It is covered with forest, and here and there are several shallow lagoons. It is about these lagoons that the lizard is supposed to make its home.
So far as I am aware, no specimens have been captured; or, if they have, they have not been preserved. A dead specimen was, however, washed down the Waima Creek, a stream leading from the plateau, about thirty-five years ago, on the occasion of the hahunga or official reinterment of the bones of Arama Karaka, when it was seen by several European visitors, and was recognised by the Maoris, who were much frightened at its appearance. Being in a partly decomposed condition, however, no attempt, I believe, was made at preservation. From the appearance of this specimen, and from such other slight details as have been gathered from the reports above mentioned, it has been concluded, I understand, that the animal is a species of salamander hitherto undescribed.
As I have occasional opportunities of visiting the neighbourhood of the Waoku Plateau, as well as the Waima Valley, where the decomposed specimen was seen, I have made it my business to seek for any information that was to be had on the subject.

Beyond the repetition of certain vague second-hand rumours, however, I had until lately never met with any success. The lizard, like the taniwha, was apparently a creature of whom all had heard but none had seen, and if it had not been for the well-authenticated specimen of Waima I should have been tempted to classify it amongst the fabled monsters of imagination.
During a recent visit to Hokianga, however, in the course of a conversation with Mr. John Webster, of Opononi, the ultimate referee in all Hokianga matters, this gentleman informed me that shortly after his coming to the district, some fifty years ago, he had seen an animal which may have been one of the sought-for species. He was exploring in a rocky part of the Wairiri Creek—a tributary of the Hokianga River—when he saw what he described as a lizard about 18 in. long of a yellowish colour. While looking about for a stick or stone wherewith to despatch it the reptile slipped down into the water, and quickly disappeared amongst the boulders in the bottom of the stream. Whether it was one of the same species there is not sufficient evidence for exact proof, but we have at least the fact established of a lizard of about the required dimensions, of amphibious habits, and found in a locality connected with the Waoku Plateau by a continuous forest. In the absence of any proof to the contrary, therefore, I think it may be fairly assumed that the species are identical, and that at that period at least the animal was pretty widely distributed.
My object in presenting these very sketchy notes is the hope that the information, scanty as it is, may lead to further research. Although a road has been made over the plateau, the place is still very much in its primitive condition. Some attempt at occupation was made some ten or fifteen years ago, but though a good deal of bush was cut down no very effective settlement took place. The swampy nature of the soil, which is nothing but a thick layer of vegetable humus, and the too abundant rains, made a successful “burn” impracticable, and after some more or less futile efforts to clear the land the attempt at settlement was generally abandoned. The clearings are fast reverting to bush, and the primæval forest which still occupies the greatest part of the surface remains practically intact. If, as seems to be the case, the animal is amphibious and probably a tree-climber, these qualities should help to secure it from the ravages of the wild pig, which would otherwise have probably exterminated it before this, as they have done in the case of the tuatara wherever they have had access.
Taking all things into consideration, therefore, I think there is little doubt that a careful and intelligent search would result in the addition of an interesting species to the list of the New Zealand fauna.

Art. XXXIV.—A Revision of the Species of the Formicidæ (Ants) of New Zealand.
[Read before the Wellington Philosophical Society, 1st June, 1904.]
Chigny près Morges, Switzerland, 3rd April, 1904.
Dear Sir,—I have received the list of the Hymenoptera of New Zealand by P. Cameron, published in your Transactions, vol. xxxv., pp. 290–299. This list contains such a large number of errors that I think it necessary to send you a revision of the New Zealand species of Formicidœ, as now known. Of the species named in the list Aphœnogaster antarcticus, Smith = Monomorium antarcticum, White, Smith named it Atta antarctica, while Cameron made an Aphœnogaster of this false Atta—the Aphœnogaster being more erroneous than the Atta. In the following paper I have given the necessary synonymy.
Mr. Cameron has forgotten to include in his list many new species discovered since 1892, and has added species that do not exist. He gives no Lasius in New Zealand, and Formica zealandia = Melophorus advena, Smith. Emery suggested that my Strumigenys antarctica may be a synonym of Orectognathus perplexus, Smith, because he thinks Smith has miscounted the number of joints of the antennæ; but, as it is only a suspicion, it must wait for more material.
I have no doubt that the North Island will furnish further new species after careful search, especially in vegetable mould and the hollows of trees.—Truly yours,
Prof. Dr. Forel.
Revision of New Zealand Species of Formicidæ.
1. Subfamily Ponerinæ, Lep., 1836.
Ponera, Latr., 1804.
(1.) P. (Mesoponera) Castanea, Mayr., Reise “Novara,” Zool. ii., 1, Formicidæ, 1865.
P. castanea, Smith, Trans. Ent. Soc. Lond., 1876, p. 489.
P. castaneicolor, Dalla Torre Cat., 1893, p. 38.
(2.) P. Antipoda, Forel, Ann. Soc. Ent. Belg., 1895.
Ectatomma, Smith, Cat., 1858.
(3.) E. (Acanthoponera) Brounii, Forel, Mitt. Schw. Ent. Ges., 1892, p. 330.

Discothyrea, Roger, Berlin, Ent. Ges., 1863.
(4.) D. Antarctica, Emery, Trans. N.Z. Inst., 1895, p. 635.
Amblyopone, Erichson, Arch. f. Natg., 1841.
(5.) A. Cephalotes, Smith, Trans. Ent. Soc. London, 1876, p. 490.
(6.) A. Saundersi, Forel, Mitt. Schw. Ent. Ges., 1892, p. 336.
2. Subfamily Myrmicinæ, Lep., 1836 (p. 169).
Orectognathus, Smith, 1854.
(7.) O. Antennatus, Smith, Trans. Ent. Soc. London, 1854, p. 228.
(8.) O. Perplexus, Smith, Trans. Ent. Soc. London, 1876, p. 491.
Strumigenys, Smith, 1860 (p. 72).
(9.) S. Antarctica, Forel, Mitt. Schw. Ent. Ges., 1892, p. 338.
Huberia, Forel, Ann. Soc. Belg., 1890.
(10.) H. Striata, Smith.
Tetramorium striatum, Smith, Trans. Ent. Soc. Lond., 1876, p. 481.
Huberia striata, Forel, Ann. Soc. Ent. Belg., 1890.
(11.) H. Braumi, Forel, Ann. Soc. Ent. Belg., Jan., 1895.
(12.) H. Striata, var. Rufescens, Forel, Revue Suisse de Zool., 1902, p. 447.
Monomorium, Mayr., 1855.
(13.) M. Antarcticum, White.
Atta antarctica, White, Zool. “Erebus” and “Terror,” part ii.
Atta antarctica, Smith, Cat., 1858, p. 167.
Monomorium fulvum, Mayr., Reise “Novara,” Zool., ii., 1865, p. 93.
Monomorium fulvum, Mayr., Verh. Zool. Bot. Ges. Wien, 1886, p. 360.
(14.) M. Nitidum, Smith.
Tetramorium nitidum, Smith, Trans. Ent. Soc. Lond., 1876, p. 480.
Monomorium nitidum, Mayr., Verh. Zool. Bot. Ges. Wien, 1886, p. 363.
(15.) M. Suteri, Forel, Mitt. Schw. Ent. Ges., 1892, p. 340.
(16.) M. Smithii, Forel, Mitt. Schw. Ent. Ges., 1892, p. 342.
(17.) M. Integrum, Forel, Ann. Soc. Ent. Belg., 1894, p. 228.
(18.) M. Antipodum, Forel, Ann. Soc. Ent. Belg., 1901, p. 377.

3. Subfamily Camponotinæ, Forel, 1878.
Melophorus, Lubbock, Journ. Linn. Soc. Zool., 1883, p. 51.
Prolasius, Forel, Mitt. Schw. Linn. Ent. Ges., 1892.
(19.) M. Advena, Smith.
Formica advena, Smith, Trans. Ent. Soc. Lond., 1862, p. 53.
Formica zelandica (?), Smith, Trans. Ent. Soc. Lond., 1878, p. 6 (♀).
Prenolepis advena, Mayr., Verh. Zool. Bot. Ges. Wien, 1886.
Prolasius advena, Forel, Mitt. Schw. Ent. Ges., 1892.
Lasius advena, Dalla Torre Cat., 1893.
Melophorus (Lasiophanes) advena, Emery, Act. Soc. Scien. Chili, 1895, p. 16.
Prenolepis, Mayr., 1861.
(20.) Pr. Longicornis, Latr.,* Hist. Nat. Journ., 1802.
Formica longicornis, Latr.
Prenolepis longicornis, Roger, Verzeichn J. Formic., 1863.
Summa.—Twenty forms (nineteen species and one variety).
Art. XXXV.—On some New Species of Macro-lepidoptera in New Zealand.
[Read before the Wellington Philosophical Society, 2nd November, 1904.]
Plate XXII.
Caradrinina.
Orthosia pallida, n. sp.
Two specimens of this insect were taken by Mr. H. W. Simmonds at Napier in April, 1903.
The expansion of the wings is nearly 1½ in. The fore wings are pale-cream colour; there is a very obscure wavy grey transverse line near the base, another at about ⅓, and another at about ¾, the space between the second and third lines being slightly shaded with brown. The hind wings are almost white, slightly shaded with grey near the termen. The head and thorax are cream-colour, and the abdomen whitish-ochreous.
This is an obscure species, and may perhaps ultimately prove to be a bleached or pale variety of some other species already described. Mr. Simmonds, however, assures me that he has no reason to doubt his specimens represent a distinct species.
[Footnote] * A cosmopolitan species (transported by ship).

Grammodes pulcherrima, Lucas. Plate XXII., fig. 4.
A single specimen of this well-known Brisbane species was captured by Mr. Creagh O'Connor at Titahi Bay, near Wellington, in March last, and so far as I am aware has not been previously found in New Zealand.
The expansion of the wings is 1⅜ in. All the wings are dark blackish-brown with white markings. The fore wings have an oblique transverse band from about ¼ on the costa to about ½ on the dorsum; another band, very slender and curved near the dorsum, from a little more than ½ on costa to about ¾ on dorsum. There is a conspicuous black spot, partially ringed with pale-yellow, near the tornus, a small oblique white mark a little before the apex, and a fine oblique shaded line from the termen below the apex ending just before the black spot. The hind wings have a very broad curved white band near the base, several spots on the termen, and a blackish blotch in the middle of the termen. The cilia of the fore wings are grey; of the hind wings white, except near the black blotch, where they are black. The head and thorax are dark-grey, the abdomen pale-grey. The underside is much paler, with the white markings much broader than on the upper surface.
Kindly identified by Mr. Lyell, of Gisborne, Victoria, from a drawing taken for that purpose from Mr. O'Connor's specimen.
Notodontina.
Chloroclystis minima, n. sp.
A single specimen of this little species was found by Mr. J. H. Lewis at Ida Valley in November, 1902.
The expansion of the wings is ⅝ in. All the wings are dark-grey, stippled and striped with blackish. The fore wings have five rather ill-defined transverse bands, which are more distinct on the costa and termen than near the centre of the wing. The hind wings are shaded with blackish near the termen, but are otherwise destitute of distinct markings. The cilia of all the wings are grey, barred with black. The head, thorax, and abdomen are dark-grey.
In general appearance this species closely resembles some of the smaller varieties of Xanthorhoe cinerearia, but may be readily distinguished from that species by its smaller size and darker coloration. Its structure and wing-outline clearly indicate its affinity with Chloroclystis.
Dichromodes ida, n. sp. Plate XXII., fig. 2.
This interesting species was discovered by Mr. J. H. Lewis at Ida Valley, Central Otago.
The expansion of the wings is ⅞ in. The fore wings are very pale greenish-blue, speckled and marked with black. There is an

ill-defined wavy black stripe near the base, another at about ⅓; this is followed by a large central clear space containing a conspicuous discal spot above middle. There is a conspicuous very jagged black stripe from a little more than ½ of costa to about ¾ of dorsum, followed by a very conspicuous pale-ochreous line; beyond this are two somewhat ill-defined black bands. The cilia are black, mixed with pale bluish-green. The hind wings are ochreous tinged with reddish and speckled with black, especially towards the base and termen. There is a conspicuous black discal spot and a clear rather wavy yellowish band about ¾ from base to termen. The cilia are blackish. Head and thorax black dotted with pale bluish-green; abdomen yellowish.
Mr. Lewis has favoured me with the following note relating to the transformations of this interesting new species. “The specimen was bred from a pupa found in a cleft of rock: a chamber had been formed by cementing moss - dust and silk together. From the fragments of caterpillar-skin remaining, I judge that the larva was one I had tried unsuccessfully to rear a few weeks ago, found feeding openly on lichen, remarkable for its fimbriated aspect, each segment being produced into irregular lobed processes at the edges—very protective amongst lichen.”
Lythria fulva, n. sp. Plate XXII., fig. 3.
This species was captured by Mr. J. H. Lewis near Mount Ida, Central Otago, at about 3,500 ft. above the sea-level.
The expansion of the wings is ¾ in. The fore wings are dull greyish-brown, greenish-tinged. There is a wavy darker band near the base; a broad median band with a strong rounded projection towards the termen above middle; a dull-greenish band on termen preceded by a row of pale dots on veins, cilia grey with blackish bars. The hind wings are dull reddish-ochreous. There are three very obscure blackish transverse lines. The cilia are grey, faintly barred with darker.
This is a very distinct species. It may be at once distinguished from L. eudidiata by the absence of the conspicuous red, black, and yellow markings on the underside of the fore and hind wings.
Porina minos, n. sp. Plate XXII., fig. 5.
This species was discovered by Mr. J. H. Lewis at Ophir.
The expansion of the wings is 1⅛ in. The fore wings are dull yellowish-brown. There are two rather large white spots near the base, a wavy irregular chain of white spots at about ⅓, an oblique chain of white spots from about ¾ of costa to ⅔ of termen. The outer chain has two spots between veins 7 and 8 and

8 and 9. The spots between veins 6 and 7 on both series are much elongated and almost touch each other, thus forming together an interrupted oblique white line on the upper portion of the wing. There are four small white spots on the termen near the tornus. The hind wings are pale yellowish-brown. The head and thorax dark-brown and very hairy; abdomen paler.
This species is easily recognised by its small size.
Explanation of Plate XXII.
| Fig. 1. |
Chærocampa celerio. (See page 359.) |
| Fig. 2. |
Dichromodes ida. |
| Fig. 3. |
Lythria fulva. |
| Fig. 4. |
Grammodes pulcherrima. |
| Fig. 5. |
Porina minos. |
Art. XXXVA.—On Macro-lepidoptera observed during the Summer of 1903–4, including a Note on the Occurrence of a Hawk-moth new to New Zealand.
[Read before the Wellington Philosophical Society, 6th July, 1904.]
Plate XXII.
Anosia erippus.
Mr. R. I. Kingsley records having seen this butterfly in his garden at Nelson on the 6th January, 1904.
Diadema bolina.
This butterfly, which is usually very rare in New Zealand, occurred in considerable numbers in various parts of the country during the past summer. The Rev. Alex. Doull, of Otahuhu, to whom I am indebted for the fine series of specimens exhibited this evening, informs me that during the end of February and beginning of March he saw no less than fifteen specimens, and succeeded in capturing seven on a flowering shrub (Escallonia floribunda) in his garden. The occurrence of Diadema bolina at Wanganui in March was reported by Mr. Ritchings Grant, and at Nelson by Mr. R. I. Kingsley, who stated that he had seen seven specimens, and heard of others.
Chrysophanus salustius.
I have noticed, since my garden at Karori has become more sheltered through the growth of trees, that this butterfly is much more abundant there than formerly. Last summer it was very common, and one warm night in January I discovered, by means of a lantern, no less than six individuals

asleep. The butterflies were simply perched on the outside foliage of a macrocarpa tree, and were quite readily seen, though the bright-yellow colouring of the closed wings suggested a faded leaf. This resemblance would, however, afford the insect very efficient protection whilst resting asleep amongst the foliage of many of the native shrubs, the faded leaves of which are, in many instances, yellow in colour. In such situations its destruction by nocturnal enemies would, no doubt, thus be largely obviated.
Sphinx convolvuli.
This insect is generally very irregular in its appearance. Last summer, however, it was reported by Mr. Grant as very plentiful at Wanganui, and by Messrs. F. G. Gibbs and R. I. Kingsley as abundant in the Nelson District.
Chærocampa celerio, Linn. Plate XXII., fig. 1.
Four specimens of this insect, which, so far as I am aware, has not been previously met with in New Zealand, were taken in Nelson last summer—one specimen, apparently the first, by Mr. Kingsley the week before Christmas, another by Mr. Frank Whitwell shortly afterwards, a third by Mr. Edward Mules on 24th February, and a fourth by Mr. Gibbs a few days later. The occurrence of this insect in New Zealand is of excessive interest, as prior to this discovery the family Sphingidœ was only represented in this country by a single, almost cosmopolitan, species, i.e., Sphinx convolvuli. It will be interesting to see if the newcomer is permanently established, and collectors should be specially on the watch to detect further specimens.
The expansion of the wings is about 3 in. The fore wings are brownish-ochreous, with short black and silvery longitudinal lines, and a shining silvery wavy streak, divided by two fine brownish lines running from the base of the dorsum to the apex, below which are whitish longitudinal lines running along the dorsum. A little below the middle of the costa is a black spot in a pale ring. The hind wings are rose-colour, with the termen and a central streak broadly black. The intermediate rosy band is also divided by black veins. The larva is green or brown, with black eye-like markings on the fifth and sixth segments, with white pupils, and enclosed in slender yellow rings. The horn is slender and long and straight. The larva feeds on the vine.” (Kirby.)
C. celerio occurs rarely in Britain south of the Caledonian Canal, and in the north of Ireland, but only as an occasional immigrant. It is a very widely distributed species, otherwise ranging through west, central, and southern Europe, south Asia, Africa, and Australia.

Selidosema rudiata.
An additional food-plant for the larva of this insect is the wharangi (Brachyglottis repanda). (See “New Zealand Moths and Butterflies,” page 83.)
Gonophylla ophiopa.
The following is the life-history of this species: The egg is oval, flattened at one end, pale sea-green, covered with numerous very slight hexagonal depressions. It is highly polished, with a large oval depression on each side of its long axis. As the enclosed embryo develops, small irregular reddish-brown patches appear on the surface of the egg-shell.
The larva, which feeds on tree-ferns (Dicksonia), is, when full-grown, about 1¼ in. in length and of uniform thickness throughout. The general colour is pale rusty-brown with an obscure pale-brown dorsal line, stronger on the thorax and at the commencement of each segment. There are two similar obscure lateral lines. The head is yellow, thickly dotted with dull-red, and the entire larva is thickly dotted with dark-brown dots and clothed with pale-reddish hairs. There are several obscure marks near the spiracular region.
This larva is very sluggish and grows slowly. It probably emerges from the egg in the autumn, hibernates during the winter, and feeds from September till the middle or end of December. It then finally buries itself in the earth and changes into a pupa, the moth appearing a month or six weeks later. (See “New Zealand Moths and Butterflies,” p. 98.)
Paradetis porphyrias.
I find that my report of the occurrence of this species at Wainuiomata, recorded in the “Transactions of the New Zealand Institute,” vol. xxxii., p. 11, is erroneous, having been founded on a mistaken identification.
Chloroclystis antarctica.
The larva of this species, which feeds on the common Veronica in December, is, when full-grown, about ¾ in. in length, rather attenuated anteriorly, almost uniform, dark reddish-brown, darker on the sides. The head is reddish, and there are traces of several longitudinal lines in younger larvæ. Others are dull yellowish-brown, with the lines plainer and the prolegs pale-yellow; but as the larva is so extremely variable a detailed description hardly appears possible. The pupa is enclosed between two leaves of the Veronica, fastened together with silk. The moth emerges at the end of January. (“New Zealand Moths and Butterflies,” p. 42.)

