
Fam. Sphæriidæ.
Cardinal teeth not exceeding two in each valve, and exhibiting a cessation of development at an early stage. (Dall.)
The New Zealand species of Sphærium and Corneocyclas are not easily separated with regard to their generic position, as both are about to the same extent inequilateral, though in Corneocyclas the anterior part is mostly more attenuated and the nepionic shell sometimes more conspicuous than in Sphærium. The cardinal teeth are very variable, and it is a tedious job to ascertain the genus from these alone, but they assist in the general diagnosis. To be absolutely certain living specimens should be examined, Sphærium having two siphonal tubes, Corneocyclas only one; but with the specimens to be described that was out of the question. Fortunately I have specimens of both genera from New Zealand in my collection, which I examined when alive, and these were of great help to me for the study of the various specimens collected by Messrs. Lucas and Hodgkin.
Much useful information was obtained from the chapter on the family Sphæriidæ in the classical work of Dr. W. H. Dall, “Tertiary Fauna of Florida.”
The indices I calculated for the Sphæriidæ are part of those used for the Unionidœ: iii. = index of height; vii. = index of diameter; x. = umbonal index.
Genus Sphærium, Scopoli (1777).
Subgenus Sphærium, Scopoli, s.s.
Type: S. corneum, Linné.
The nepionic shell passing into the adult without any distinct demarcation; the anterior end shorter; the ligament subinternal; the two right cardinals widely divergent and coalescent at their adjacent or upper ends, thus apparently forming but one tooth, but which if it had continued in development would have separated into two; the widening of the ventral angle causes the A-shape to disappear; the nepionic shell (and consequently the beaks) is finely concentrically striate or even nearly smooth and rather convex. (Dall.)
Sphærium novæ-zelandiœ, Deshayes (1853).
Cat. Conchif. Brit. Museum, p. 272; P.Z.S., 1854, p. 342.
(1.) Lake Rotoiti (Stat. 15—From muddy bottom and weeds in 6 ft.).—Three specimens of nearly equal size, suborbicular, subequilateral; colour grey, near the ventral margin yellowish; beaks covered with a ferrugineous incrustation. Nepionic shell

hardly to be distinguished from the later growth, obtuse; anterior side of shell slightly produced, rounded, posterior side broadly convex; the whole of the valves concentrically finely striate. Ligament inset. Left valve with a straight lamellar upper cardinal, and below and slightly in front of it a curved stouter tooth. In the right valve the two cardinals are united, forming an oblong squarish tooth with three denticulations on the lower margin. Laterals smooth. Fig. 5 illustrates the cardinal teeth of the two valves. Length, 5 mm.; height, 4¼ mm.; diam., 2¾ mm. Umbo 3 mm. from anterior end. The indices are—iii. = 85; vii. = 55; x. = 60.
This is no doubt the same as S. lenticula, Dunker, as Hochstetter obtained the specimens from the lakes Rotoiti and Taupo, and I consider it as a synonym of S. novæ-ze-landiæ.
(2.) Lake Taupo (Stat. 4—Dredged with weeds in 20 ft., and from bottom of coarse pumice in 100 ft.).—Four specimens of straw colour, the nepionic shell rather distinct, but otherwise not differing from the typical form. Right valve with two minute cardinals, the anterior oblong, the posterior short, triangularly oval, elevated. Left valve with two minute teeth, broadly rounded behind, pointed at the anterior end, one in front of the other. Fig. 6 shows the form and position of the cardinals. Length, 4½ mm.; height, 4 mm.; diam., 2½ mm. Umbo 2¾ mm. from anterior end. Indices—iii. = 84–89; vii. = 56; x. = 61.
(3.) Lake Taupo (Stat. 34—With weeds from 20 ft. to 80 ft.).—Six small specimens, a little more oval and compressed, but otherwise the same as the examples from Stat. 4. Length, 4 mm.; height, 3 mm.; diam., 1¾ mm. These are the dimensions of Dunker's S. lenticula. Indices—iii. = 75; vii.—44.
(4.) Lake Waikaremoana (Stat. 33—Dredged in 10 ft. to 20 ft.).—There are twenty-three small to minute specimens. The larger examples are yellowish-grey, the small ones straw-colour. In the right valve there are two lamellar cardinal teeth meeting at an obtuse angle; in the left valve there is one crescent-shaped

rounded cardinal, and a strong bifid crenulated cardinal below and slightly in front of it. Fig. 7 illustrates the cardinals of the valves. The lateral teeth are smooth; the ligament inset. Dimensions of two specimens—Length, 4¼ mm.; height, 3½ mm.; diam., 2 mm.; beak 2½ mm. from anterior margin: length, 4 mm.; height, 3½ mm.; diam., 2 mm.; beak 2¼ mm. from anterior margin. The mean of the indices is—iii. = 85; vii. = 49; x. = 58.
Genus Corneocyclas, Férussac (1818).
(= Pisidium, Pfeiffer, &c.)
Subgenus Corneocyclas.
Nepionic shell convex, concentrically striated; hinge with two separate cardinals in the left, and a single compound, usually arcuate, cardinal in the right valve. (Dall.)
Sec. Corneocyclas, s.s.
Type: Tellina pusilla, Gmelin.
Nepionic valves passing into the mature disc without any strong demarcation; the anterior cardinal and lateral adjacent and retaining traces of their original connection; ligament internal. (Dall.)
Corneocyclas novozeelandica, Prime (1862).
P.Z.S., 1862, p. 3.
(1.) Lake Waikare (Stat. 29—From weeds in 1½ ft. to 2 ft.).—Nine specimens of different size; colour greyish-yellow, shining, inequilateral, with fine concentric lines, which are crossed by close microscopic radiate striæ. Nepionic shell convex, concentrically striated, well delimited, but passing without any change into the disc of the adult valve. Ligament inset. Right valve with the two cardinals remaining united, crescent-shaped; four laterals. Left valve with two cardinals, the posterior bifid, the anterior triangular; two laterals. The accompanying fig. 8 shows the characters of the hinge.

Four specimens were measured—Length, 2¾–5 mm.; height, 2½–4 mm.; diam., 1¼–2½ mm.: and the mean indices are—iii. = 83; vii. = 48; x. = 59.
These specimens are very much like those I collected in the River Avon, Christchurch.
(2.) Lake Waikare (Stat. 29F—From weeds in 1½ ft. to 2 ft.).—Four specimens, dead when collected. They do not differ much from those of Stat. 29, except being slightly more globular. The dimensions of three specimens measured are—Length, 5 mm.; height, 4½ mm.; diam., 3¼ mm.; umbo 3 mm. from anterior margin: length, 4¼ mm.; height, 3½ mm.; diam., 2¼ mm. umbo 2½ mm. from anterior margin: length, 3 mm.; height, 2½ mm.; diam., 1½ mm.; umbo 1¾ mm. from anterior margin. The mean indices are—iii. = 85; vii. = 56; x. = 58.
(3.) Lake Wakatipu (Stat. 17—Dredged with weeds in 20 ft. to 100 ft.).—One minute specimen, rounded, yellowish-white, subequilateral. Length, 2¼ mm.; height, 2 mm.
This small specimen has the aspect of a Sphærium, but the nepionic shell is so distinct that I prefer placing it in Corneocyclas for the present. I tried to separate the valves, but could not do it without running the risk of breaking the valves. Without ample material, and especially examining the living animal, it is impossible to be quite certain about its generic position.
(4.) Lake Wakatipu (Stat. 34—Dredged in 10 ft. to 30 ft.).—One adult and four young specimens. The adult is light-brown, the nepionic shell distinctly limited; the young examples are grey to yellowish, more inequilateral and tapering in front. The dimensions of two specimens are—Length, 5¼ mm.; height, 4½ mm.; diam., 2½ mm.; umbo 2¾ mm. from anterior end: length, 3½ mm.; height, 1¾ mm.; diam., 1½ mm.; umbo 2 mm. from anterior end: and the mean indices are—iii. = 83; vii. = 46; x. = 55.
There is one specimen, covered with a black coating, which is suborbicular, much compressed, inequilateral, with inconspicuous beaks, the anterior side broadly rounded. The dimensions are—Length, 4 mm.; height, 3¼ mm.; diam., 1½ mm.; umbo 2½ mm. from anterior end: and the indices are—iii. = 81; vii. = 38; x. = 62. I have seen similar abnormities amongst specimens I collected in the River Avon.
(5.) Lake Manapouri (Stat. 13—Dredged in shallow water near the shore).—Three young specimens, which are at once recognised as being Corneocyclas, being more inequilateral, compressed and attenuated anteriorly than adult specimens. Had I not seen the same forms from other localities, collected together with adult examples, I might have been inclined to consider

them a distinct species. With few exceptions the young shells of C. novozeelandica show the characters just indicated. The colour is light-grey, and the nepionic shell is distinct. The dimensions of two specimens are—Length, 3¼ mm.; height, 2¾ mm.; diam., 1½ mm.; umbo 2 mm. from anterior end: length, 2¾ mm.; height, 2 mm.; diam., 1 mm.; umbo, 2 mm. from anterior end. The mean indices are—iii. = 78; vii. = 41; x. = 66.
Corneocyclas hodgkini, n. sp. Fig. 9.
(1.) Lake Waikaremoana (Stat. 18—Dredged in 800 ft.).—Two specimens. They are very small, oval, much compressed, the anterior side produced and attenuated; posterior and ventral margins regularly rounded; colour yellowish-white; beaks obtuse, unconspicuous; nepionic shell distinct, very finely concentrically striated, passing without change into the adult valves, which are irregularly finely concentrically striate. Posterior part with a ferrugineous coating. The dimensions are—Length, 2¾ mm.; height, 2 mm.; diam., 1 mm.; and the indices—iii. = 72; vii. = 36: length, 2 mm.; height, 1¾ mm.; diam., ¾ mm.; and the indices—iii. = 87; vii. = 50.
Named in honour of Mr. G. L. Hodgkin, who so ably assisted Mr. Lucas in his arduous work.
Type in my collection.
(2.) Lake Taupo (Stat. 10—Dredged in 280 ft. and 320 ft., muddy bottom).—Six small specimens, thickly coated all over with ferrugineous earth, so that the form of the shell is unrecognisable. I succeeded in cleaning one specimen, and found it to agree with the specimens just described from Lake Waikaremoana. Dimensions of the cleaned specimen—Length, 3 mm.; height, 2¼ mm.; diam., 1 mm.

