Go to National Library of New Zealand Te Puna Mātauranga o Aotearoa
Volume 39, 1906

Transactions
of the
New Zealand Institute,
1906.

Art. I.—The Fungus Flora of New Zealand.—Part II.

[Read before the Wellington Philosophical Society, 5th April, 1906.]

Communicated by A. Hamilton.

Plates I and II.
PolyporeÆ.

The one constant morphological feature of the present family consists in the hymenium, or spore-bearing surface, being developed over the entire inner surface of tubes or pits. In the most typical forms, included in such genera as Boletus and Polyporus, the tubes collectively forming the hymenium are frequently 2 cm. or 3 cm. long, and vary in different species from 0.5 mm. to 5 mm. in diameter. In the highest types the tubes are cylindrical, and are packed compactly side by side, as seen in a vertical section, whereas when viewed in the entire plant the pores or openings of the tubes only are seen. The form of the pores is circular when the tubes are cylindrical; in other instances the pores are polygonal, or sinuous, when somewhat elongated and wavy or flexuous.

In certain genera the tubes are so very shallow that they merely resemble circular, polygonal, or sinuous pits or depressions.

The bordering walls of the tubes, which bear the hymenium on their free surface, are called “dissepiments.”

The elements of the hymenium consist of basidia and paraphyses, and in many instances large cystidia are also present in considerable numbers.

Texture, size, and form vary exceedingly in the present group. In Boletus the species are fleshy, soft, and putrescent, completely disappearing within one or two weeks after their

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first appearance. On the other hand, the species of Fomes are all perennial, hard, and woody, and form a new hymenial layer of tubes each year; consequently in old specimens the tubes are described as “stratose,” each stratum of tubes corresponding to one year's growth.

As illustrating the variety of form included in the family, in Boletus, also in some species of Polyporus and Polystictus, there is an agaric type of structure, consisting of a pileus and central stem, pores replacing gills on the under surface of the pileus. In other genera the stem is lateral or altogether absent, the fungus being fixed by a broad base and forming bracket-or hoof-like horizontal projections. In other genera, again, as Poria, the entire fungus is quite thin, forming more or less extended patches firmly fixed to the matrix, and having the free surface completely covered with the hymenium. Some species are quite minute; others, again, are veritable fungal giants, certain species of Polyporus and Fomes numbering amongst the largest of known Fungi.

The family includes a considerable number of destructive parasites; species of Polyporus, Fomes, Merulius, and Poria being more especially destructive to forest and orchard trees; worked timber also suffers unless special precautions are taken.

Boletus alone, so far as is known, contains a few edible species, the members of the other genera being either woody or coriaceous, Strobilomyces, a genus closely allied to Boletus, contains several species from Australia, some of which may prove to be edible.

It is somewhat remarkable that no specimens representing the genera Boletus or Strobilomyces have as yet been recorded as occurring in New Zealand. It would appear highly improbable that representatives of these genera should be entirely absent, considering their comparative abundance in Australia.

Analysis of the Genera.

36.Polyporus. Pileus smooth, fleshy, flesh soft; tubes sharply defined but not separable from the flesh, not stratose.

37.Fomes. Pileus smooth, flesh thick, woody; tubes woody, stratose.

38.Polystictus. Pileus hirsute or silky, flesh quite thin; tubes short, not stratose.

39.Poria. Entirely resupinate; flesh usually very thin; tubes short.

40.Trimetes. Pileus corky, sessile; tubes penetrating different depths into the flesh, not stratose.

41.Dædalea. Pileus corky, sessile; tubes elongated and sinuous, walls thick, elastic.

42.Favolus. Tubes elongated and lamellose, radiating from point of attachment of pileus.

43.Laschia Stipitate or dimidiate; tubes as in Polyporus, but along with the pileus soft and subgelatinous.

44.Merulius. Resupinate, subgelatinous; pores very shallow, irregular, often reduced to irregular wrinkles or folds.

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36. Polyporus, Mich.

Stem central, lateral, or absent; pileus fleshy, flesh soft and tough at first, becoming firmer, externally more or less glabrous, not sulcate nor zoned; tubes not separable from flesh of pileus, never stratose; pores rounded or angular, often more or less torn at the margin. Annual.

Polyporus, Micheli, Gen. Pl., p. 129 (in part).

Certain central-stemmed fleshy species of Polyporus closely resemble Boletus, but are at once distinguished by the tubes not being readily separable from the flesh of the pileus. Fomes differs in the woody consistency of the entire fungus, concentrically ridged pileus, and stratose pores. Polystictus is separated by the thin flesh and silky or hirsute, usually zoned, pileus.

The great majority of species grow on wood. A few central-stemmed species, leading up to the genus Boletus, grow on the ground.

I. Stipitate; stem central, or nearly so.

Polyporus arcularius, Fries, Hym. Eur., 526; Hdbk. N.Z. Flora, p. 607; Sacc., Syll. vi, no. 4903; Austr. Fung., p. 113.

Stipitate; pileus 2–4 cm. across, almost flat, or slightly depressed at the centre, margin more or less incurved, yellowish-brown, squamulose with small darker scales which are most persistent at the margin but eventually entirely disappear, flesh very thin; tubes very shallow; pores large elongatohexagonal, elongated radially, pale-wood colour, entire, 1.5–2 mm. long by 0.5–1 mm. wide; stem central, 2–3 cm. long, slender, coloured like the pileus, squamulose, becoming bald.

On trunks, &c. New Zealand. Australia, Tasmania, Mauritius, Cape of Good Hope, Natal, India, Java, China, Formosa, Madagascar, Ceylon, central and southern Europe, United States, Brazil.

A very beautiful fungus; when in full vigour the margin is beautifully fringed, and the pileus dotted with minute squamules, as is also the stem; eventually, however, every part becomes bald. Distinguished by the large, entire, radially elongated pores.

Polyporus melanopus, Fries, Hym. Eur., p. 534; Sacc., Syll. vi, no. 4958; Austr. Fung., p. 115. Syn., Polyporus leprodes, Rost.

Pileus 5–10 cm. across, flattish at first, then becoming depressed or even irregularly funnel-shaped, rather thin, pliant when growing, delicately flocculose when young, whitish, then dingy yellowish-brown, margin often wavy or lobed; stem varying in position from lateral to being almost central, rather slender, short but variable in length, tapering upwards, blackish-

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brown; pores very shallow, whitish, minute, decurrent for a short distance down the stem.

On dead wood, branches, and on the ground. New Zealand. Victoria, Queensland, Europe.

Distinguished from allied species by the black stem tapering upwards. The flesh is white, softish, and not at all woody. When growth is very vigorous the margin of the pileus is often beautifully waved and crisped, and under such circumstances the pores are often larger than usual.

II. Stem lateral when present, often almost obsolete, but pileus attached by a narrowed base.

Polyporus phlebophorus, Berk., Fl. N.Z., ii, p. 177, tab. cv, fig. 3; Hdbk. N.Z. Flora, p. 607; Sacc., Syll. vi, no. 4999.

Entirely pure white; pileus about 2.5 cm. long and broad, irregularly fan-shaped, and narrowed behind into a very short, wrinkled, stem-like point of attachment; glabrous; cuticle subgelatinous, then becoming dry and cartilaginous; pores very minute; dissepiments thin, minutely toothed.

On decaying logs. Tarawera, Northern Island, New Zealand.

A very beautiful and distinct species, which does not appear to have been collected since the original specimens on which the species was founded were found by Colenso.

Polyporus xerophyllus, Berk., Fl. N.Z., ii, p. 178, tab. cv, fig. 2; Hdbk. N.Z. Flora, p. 607.

Pileus suborbicular or reniform, 2–3 cm. across, rufousbrown, marked with radiating wrinkles, minutely scabrid, or rough with raised points; stem lateral, up to 0.1 cm. long, black, wrinkled, very minutely velvety; hymenium whitish, pores very minute, dissepiments entire.

On dead wood. New Zealand.

An endemic species discovered by Colenso. Distinguished from allies by the black, minutely velvety stem and radially wrinkled pileus.

Polyporus grammocephalus, Berk., Hook. Lond. Journ., 1842, p. 148; Sacc., Syll. vi, no. 5005; Cooke, Austr. Fung., p. 117. Syn., Polyporus emerici, Kalchbr., Grev., x, pl. 145, fig. 125; Polyporus russiceps, Berk. and Broome, Journ. Linn. Soc. (Bot.), xiv, p. 48; Polyporus incompletus, Cesati, Myc. Borneo, p. 4; Polyporus platotis, Berk. and Broome, Linn. Trans., ser. ii, vol. i, p. 401; Polyporus fusco-lineatus, Berk. and Broome, Linn. Trans., ser. ii, vol. i, p. 401.

Pileus very thin, 2–3 mm. thick, obovate, irregularly circular or reniform, horizontal, margin sometimes more or less wavy,

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nearly flat, varying from pale-ochraceous through reddish-brown to umber, with very delicate lines radiating from the point of attachment to the margin, otherwise glabrous, 3–8 cm. broad; pores rather large, irregular, coloured like the pileus; stem usually lateral, very short, 3–4 mm. long by 2–3 mm. thick, scarcely discoid, sometimes quite central, at others almost absent.

On rotten trunks, fallen wood, &c. New Zealand. Queensland, New South Wales, New Guinea.

A very variable species in minor points, but characterized by the thin flesh, very short lateral stem, and the pileus streaked with very delicate, crowded, radiating ridges.

Polyporus borealis, Fries, Syst. Myc., i, p. 366; Hdbk. N.Z. Flora, p. 609; Cooke, Austr. Fung., p. 124; Sacc., Syll. vi, no. 5187.

Horizontal, subspathulate or reniform, either narrowed behind into a short more or less distinct stem, or thick and sessile, 4–10 cm. across, whitish then dingy-yellow, spongy then corky, compact, hairy; flesh thick, whitish, composed of parallel fibres; tubes 5–8 mm. long; pores unequal, flexuous; dissepiments thin, white, torn; spores hyaline, subglobose, 4 μ diameter.

On stumps and trunks of conifers, &c. Otago, Middle Island, New Zealand. Asiatic Siberia, Europe, United States.

Pileus often radiately wrinkled, rigid and more or less incurved when dry. When a stem is present the pores are more or less decurrent. Fleshy when young, becoming corky with age.

Polyporus colensoi, Berk., Fl. N.Z., ii, p. 178; Hdbk. N.Z. Flora, p. 607; Sacc., Syll. vi, no. 5016.

Main branches numerous, rather slender, springing from a common basal mass, and dividing into many smaller branchlets, each terminated by a small, fan-shaped, depressed pileus, brownish, nearly smooth; hymenium pale; pores shallow, large, often elongated, decurrent; dissepiments thin, edge acute, often toothed.

On trunks, Tarawera, Northern Island, New Zealand.

“Forming a mass more than 1 ft. across; main stems slender, distinct, somewhat elongated, repeatedly dichotomous. Pilei extremely numerous, flabelliform, expanded, depressed above, brownish, smooth, or nearly so, with a few raised lines. Pores pale, often very much elongated, decurrent; dissepiments thin; edge extremely acute, often toothed, sublamelliform. This is a noble species, and evidently differing from every form of P. intybaceus in its distinct dichotomous branches and the constantly acute dissepiments. Some of the figures of Hydnum

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coralloides give a better idea of the ramification of the species than any of those of P. frondosus.” (Berk.)

Polyporus lactus, Cooke, Grev., xii, p. 16; Sacc., Syll. vi, no. 5047; Cooke, Austr. Fung., p. 119.

Imbricated, much divided, rather thin, tough, tawny-orange or rusty above; pilei dimidiate, coalescing, entire, surface broken up into adpressed scaly zones, converging behind into a narrowed point of attachment, margin acute, sometimes tinged crimson or purple, 7–15 cm. broad; pores large, irregular in form, dissepiments thin, pallid; flesh thin, fibrous, orange-rusty.

On decaying trunks. Northern Island, New Zealand. Victoria.

Closely allied to the European Polyporus giganteus, differing in the bright colour of the pileus and flesh, and in the larger pores.

III. Pileus sessile, attached by a broad base.

Polyporus plebius, Berk., Fl. N.Z., ii, p. 179; Hdbk. N.Z. Flora, p. 608; Cooke, Austr. Fung., p. 126; Sacc., Syll. vi, no. 5247.

Pallid or pale-wood colour, imbricated, sessile, attached by a broad base, more or less semicircular, 3–8 cm. across, up to 1 cm. thick behind, becoming thinner towards the margin, which is sometimes rather thick and blunt, at other thinner and acute, not zoned, minutely pubescent when growing, usually even, sometimes with a groove or more or less rugged; flesh corky; hymenium concave; pores minute, ⅙–⅛ mm. diameter.

On dead wood. Northern Island, New Zealand. Himalayas, Queensland, Victoria, Cuba.

The varieties indicated by Berkeley, depending on the acute or obtuse margin of the pileus, are not constant features, both occurring in the same group of specimens.

“In the New Zealand form the older parts are inclined to assume a spuriously laccate appearance.” (Berk.)

Polyporus scruposus, Fries, Epicr., p. 473; Fl. N.Z., ii, p. 178; Hdbk. N.Z. Flora, p. 608; Sacc., Syll. vi, no. 5130; Cooke, Austr. Fung., p. 122.

Pileus sessile, attached by a broad base, semicircular or sometimes almost triquetrous, 1 cm. or more thick at the base and becoming thinner towards the acute margin; surface with raised concentric zones, rough with raised points as if minutely corrugated, brown, margin paler; flesh rather thick, orange-brown; tubes 2–5 mm. long; pores very minute, rounded, umber, often with a tinge of purple.

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On dead wood. Northern Island, New Zealand. Victoria, New South Wales, Queensland, Western Australia, Tasmania, Pegu, Nepal, Island of Aru, United States, Mexico, Cuba.

Hard and woody, solitary or imbricated; known by the sulcate, rough pileus, and coloured flesh.

Var. iridioides. Syn., Polyporus iridioides, Berk., Hook. Lond. Journ. Bot., ii, p. 515; Fl. N.Z., ii, p. 178; Hdbk. N.Z. Flora, p. 608 (called by a slip iridioides); Cooke, Austr. Fung., p. 122; Sacc., Syll. vi, no. 5131.

Agreeing in habit and size with the type form; differing more especially in having the pileus rough, with elongated bristly nodules behind.

Although Dr. Cooke retains Polyporus isidiodes as a species in the “Handbook of Australian Fungi,” after giving the diagnosis he says, “only a variety of P. scruposus,” a statement with which I entirely agree.

Polyporus dichrous, Fries, Syst. Myc., i, p. 364; Sacc., Syll. vii, no. 5152; Austr. Fung., p. 123; Hdbk. N.Z. Flora, p. 608.

Thin, tough, soft, sessile, effuso-reflexed, often imbricated, 2–3 in. wide, sometimes much larger, slightly silky, white; pores very shallow, minute, rounded, cinnamon-colour, 4–5 in the space of 1 mm.

On trunks. Northern Island, New Zealand. Victoria, South Africa, Europe, United States.

Often covering a large extent of surface, more or less resupinate, with numerous free, spreading margins or lobes. Almost like a Polystictus, but soft and not zoned.

37. Fomes, Fries.

Pileus hard, covered with a rigid, crustaceous, zoneless, often concentrically grooved cuticle; tubes stratose. Perennial.

Fomes, Fries, Nov. Symb., p. 31. Polyporus of old authors.

The woody pileus with a cartilaginous cuticle not ornamented with coloured zones, and the stratose tubes, stamp the present genus.

I. Stem lateral, sometimes very short.

Fomes lucidus, Fries, Nov. Symb., p. 61; Fl. N.Z., ii, 177; Hdbk. N.Z. Flora, 607; Cooke, Austr. Fung., p. 128; Sacc., Syll. vi, no. 5305.

Pileus horizontal or oblique, flabelliform, subreniform, or irregular in form, laterally stipitate, 8–15 cm. across, corky, then hard and woody, sulcato-rugose, deep chestnut-red, or

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sometimes almost blood-red, polished, shining; tubes 1–1.5 cm. long, pores minute, whitish, then cinnamon; stem very variable in length, irregularly wrinkled, coloured and polished like the pileus; spores 7x5 μ, tinged brown.

On trunks, &c. Northern Island, New Zealand. Common in all tropical, subtropical, and temperate regions.

Very variable in size and form; the stem is sometimes almost or even quite central. The lacquered appearance of the pileus and stem is due to the exudation of a thick glutinous liquid which covers the surface, where it soon dries, giving to the surface a perfectly smooth and polished aspect.

II. Sessile, attached by a broad base.

Fomes igniarius, Fries, Syst. Myc., i, p. 375; Fl. N.Z., ii, p. 179; Hdbk. N.Z. Flora, p. 608; Sacc., Syll. vi, no. 5412; Cooke, Austr. Fung., p. 131.

Pileus at first irregularly globose, even, with a delicate brownish nap clothing the surface, then becoming hoof-shaped, rust-colour, changing to opaque dingy-brown, cuticle very hard, uneven, 6–18 cm. across; margin blunt, paler; flesh ferruginous, zoned, very hard; tubes 2–5 cm. long, very small, stratose, cinnamon, filled with white mycelium when old, general surface of hymenium convex; pores ¼–⅓ mm. across, rounded, at first hoary; spores subglobose, hyaline, 6–7 μ; diameter; cystidia scanty, 10–25 × 5–6 μ.

On trunks of various trees, living and dead. Bay of Islands, Northern Island, New Zealand. Victoria, Queensland, New South Wales, South Australia, Western Australia, Tasmania, Ceylon, India, Siberia, Europe, United States.

Sometimes very large, thick, and in section more or less triangular, hence hoof-shaped. Allied to Fomes fomentarius, differing in being a perennial plant, very hard cuticle and flesh, and in the hyaline spores. A destructive wound-parasite, attacking many different species of trees, dissolving and destroying the heart-wood.

Fomes australis, Fries, Hym. Eur., p. 536; Hbdk. N.Z. Flora, p. 608; Sacc., Syll. vi, no. 5394; Austr. Fung., p. 130.

Normally bracket-shaped, sessile, dimidiate, concentrically zoned or irregularly wavy or tuberculose, glabrous, dark-brown, external crust exceedingly hard, 8–20 cm. across; flesh not very thick, brown, with a tinge of purple; tubes very long, stratose, brown, substance hard; pores at first whitish, then umber, very minute, about 4 in the space of 1 mm.; margin sterile, often slightly thickened.

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On dead trunks, stumps, &c. Common in New Zealand, also widely distributed over southern tropical and subtropical regions.

A well-marked but at the same time exceedingly variable species. At times the surface of the pileus is marked with more or less deeply indented concentric furrows, at others only slightly concentrically zoned, whereas in other forms the pileus is irregularly rugged or tuberculated. The pileus is sometimes thin, almost semicircular, and bracket-like; at others it becomes elongated and almost cylindrical. Tubes distinctly stratose in thick specimens. The principal features are the very rigid woody cortex, almost too hard to cut with a knife, and the very minute pores.

Fomes hemitephrus, Cooke, Grev., xiv, p. 21; Sacc., Syll., vi, no. 5497. Polyporus hemitephrus, Berk., Fl. N.Z., ii, 179; Hdbk. N.Z. Flora, p. 608 (incorrectly written hemitrephius).

Pileus bracket-shaped, often with a boss near the point of attachment, usually with coarse concentric ridges, glabrous, brown, sometimes paler when young, when the rounded margin is whitish, hard, up to 12 cm. across; flesh 3–4 cm. thick, wood-colour, hard; tubes wood-colour, imperfectly stratified; pores very minute, rounded; hymenium concave, whitish.

On trunks of trees. Northern Island, New Zealand. Victoria, India, Gold Coast.

Allied to Fomes fraxineus, Fries.

Fomes salicinus, Fries, Syst. Myc, i, p. 376; Fl. N.Z., ii, p. 179; Hdbk. N.Z. Flora, p. 608; Cooke, Austr. Fung., p. 132; Sacc., Syll., vi, no. 5429.

Often broadly effused, woody, very hard, the greater portion usually resupinate, with a narrow, wavy, smooth, blunt, spreading free margin, cinnamon then greyish; pores minute, rounded, rusty-cinnamon like the flesh; spores 5 × 3 μ; cystidia plentiful 12–35 × 6–8 μ.

On trunks, living and dead, especially species of Salix. Dusky Bay, Middle Island, New Zealand. Queensland, South Africa, Europe, United States.

Pileus 1 ft. or more across; entirely resupinate, or on vertical trunks having the upper margin free and reflexed. Allied to Fomes fomentarius and F. igniarius, differing in not being hoof-shaped, but thinner in the flesh, and in being more effused over the matrix. Sometimes acts as a destructive wound-parasite.

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Fomes [ unclear: ] hauslerianus, P. Henn., Hedw., 1896, p. 305.

Pileus rigid, somewhat woody, reniform or flabellate, radiately venosely rugulose, zoneless, at first covered with olive mealy down, then naked and blackish, 2.5–3 cm. long; margin thin, rigid, waved, crenate; stem short, lateral, olive-primrose; flesh pallid with a yellow tinge; hymenium greyish; tubes short; pores punctiform, minute; spores subglobose

On trunks. New Zealand; Auckland, Ohaupo.

38. Polysticus, Fries.

Pileus thin, coriaceous, cuticle fibrous, silky or hirsute, often with coloured zones; tubes short. Annual.

Polystictus, Fries, Nov. Symb., p. 54. Polyporus of old authors.

Often imbricated or growing in superposed tiers.

I. Pileus stipitate, stem central.

Polystictus oblectans, Berk., Hook. Journ., 1845, p. 51; Fl. N.Z., ii, p. 177; Hdbk. N.Z. Flora, p. 607; Austr. Fung., p. 138; Sacc., Syll. vi, no. 5545.

Entirely bright cinnamon-brown, stem usually darkest; pileus 2–5 cm. across, thin, coriaceous, depressed, margin often torn and wavy, zoned, strigosely silky, shining; pores very short, minute, dissepiments torn; stem central, 2–3 cm. long, 2–4 mm. thick, velvety.

On the ground. Northern Island, New Zealand. Victoria, Queensland, Western Australia, Tasmania, Ceylon, India, Brazil.

Polystictus cinnamomeus, Jacq., is closely allied to the present species, differing chiefly in the larger angular pores.

II. Pileus sessile, attached laterally.

Polystictus sanguineus, Fries, Epicr., p. 444; Hdbk. N.Z. Flora, p. 609; Sacc., Syll. vi, no. 5631; Austr. Fung., p. 141.

Pileus reniform or somewhat fan-shaped, sessile, or narrowed behind into a very short stem-like base, attached by an expanded disc, 3–10 cm. broad, glabrous and polished, sometimes concentrically zoned, vermilion, bleaching almost white when old; flesh thin, compact, 0.5 cm. or less thick, margin thinnest, frequently lobed or wavy; tubes very short; pores minute, rounded, 3 in space of 1 mm., deep and persistent vermilion.

On trunks, stumps, &c. New Zealand. Australia, Tasmania, Lord Howe Island, India, Java, Borneo, Philippines, Malacca, Sumatra, Friendly and Society Islands, South America, Central America, Cuba, United States

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A showy fungus, superficially resembling Polystictus cinnabarinus. For distinction between the two see note under the last-named species.

Polystictus cinnabarinus, Fries, Syst. Myc., i, p. 371; Hdbk. N.Z. Flora, p. 609; Sacc., Syll. vi, no. 5711; Cooke, Austr. Fung., p. 146.

Pileus semicircular, somewhat narrowed behind at the point of attachment; convexo-plane, thickest behind, where it is 1–2 cm. thick, margin thin, 6–10 cm. broad; pileus corky, often slightly zoned or rugulose, at first downy, then glabrous, vermilion, bleaching almost white with age; flesh spongy or fibrous, red; tubes 2–3 mm. long; pores vermilion, roundish, 2, rarely more, in the space of 1 mm.

On dead trunks, &c. New Zealand. Australia, Tasmania, India, Ceylon, Sumatra, Cape of Good Hope, Europe, United States.

A beautiful fungus, superficially resembling Polystictus sanguineus. The latter, however, differs in the thinner substance of the pileus, which is glabrous and polished at all stages, is attached to the matrix by a distinct disc, and has smaller pores.

Polystictus versicolor, Fries, Syst. Myc., i, p. 368; Sacc., Syll. vi, no. 5741; Austr. Fung., p. 146. Polyporus versicolor, Fries, Hdbk. N.Z. Flora, p. 609.

Pileus thin, coriaceous, flat on both surfaces, often slightly depressed behind, upper surface densely velvety, shining, with variously coloured concentric bands, 3–10 cm. across; pores very short, minute, white, becoming tinged buff or cream-colour; dissepiments thin, becoming torn; pores about ¼ mm. across.

On trunks, stumps, branches, &c. Northern Island, New Zealand. Victoria, Queensland, New South Wales, Tasmania, Europe, America.

Distinguished by the silky shining pileus being marked with concentric zones of various colours.

Polystictus velutinus, Fries, Syst. Myc., i, p. 368; Fl. N.Z., ii, p. 178; Hdbk. N.Z. Flora, p. 609; Cooke, Austr. Fung., p. 147; Sacc., Syll. vi, no. 5763.

Horizontal, attached laterally by a more or less narrowed base, and imbricated, flat, or attached by a more or less central point and remaining flattened; thin, pliant when growing, then rigid, 4–10 cm. across; pileus velvety, indistinctly zoned, dull, dingy-white to pale yellowish-white, sometimes tinged brown; tubes very short; pores subangular, white, minute, often dis-

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appearing towards the margin; spores broadly elliptic-oblong, obliquely apiculate, 5–6 × 4 μ.

On trunks, stumps, &c. Northern Island, New Zealand. Victoria, Queensland, India, Borneo, Java, Philippines, Japan, Europe, Asiatic Siberia, United States, Cuba, South America.

Flaccid when young and growing, shrinking and curling inwards when dried; about 3–4 mm. thick. Allied to Polystictus versicolor, from which the present differs in the dull, opaque—not silkily shining—pileus, which is whitish, and not variegated with zones of deep colours.

Polystictus hirsutus, Fries, Syst. Myc., i, p. 367; Sacc., Syll. vi, no. 5760. Polyporus hirsutus, Fries, Hdbk. N.Z. Flora, p. 609.

Pileus thin, both surfaces almost plane, more or less semicircular in outline, often imbricated, upper surface generally whitish, sometimes tinged yellow, coarsely hispid or strigose, often concentrically zoned, 3–7 cm. across; pores roundish, small, about ⅓ mm., greyish-buff; spores narrowly elliptical, 4–5 × 2 μ.

On stumps, dead wood, &c. Middle Island, New Zealand. Victoria, Queensland, New South Wales, Europe, Siberia, United States.

Differs from Polystictus versicolor and P. velutinus in the coarsely strigose hymenium and the greyish pores.

Polystictus tabacinus, Mont., Flor. Juan Fernandez, no. 15; Fl. N.Z., ii, p. 178; Hdbk. N.Z. Flora, p. 609; Sacc., Syll. vi, no. 5876; Cooke, Austr. Fung., p. 151.

Pilei imbricate, entirely dark-brown, sessile, irregularly semicircular or shell-shaped, concentrically zoned, silky and shining, very thin, rigid, 2–6 cm. long, 2–4 cm. broad; tubes very short, rather small, the dissepiments becoming torn or toothed.

On dead wood. Northern Island, New Zealand. New South Wales, Mauritius, Island of Aru, Chili, Juan Fernandez.

A very beautiful fungus; flesh very thin, as is the whole fungus; pileus glistening with a silky sheen. Distinguished by the umber-brown colour of every part. Much incurved and very rigid when dry.

Polystictus sector, Fries, Epicr., p. 480; Sacc., Syll. vi, no. 5900.

Pileus 2–4 cm. across, wedge-shaped or fan-shaped, sessile, imbricated, thin, coriaceous, downy, becoming glabrous, zoned, striate, pale-brown; pores shallow, small, brown, sometimes with a tinge of purple, dissepiments thin.

On branches, &c. Northern Island, New Zealand. Tasmania, Cuba, Brazil.

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Pileus sometimes almost triangular, narrowed behind, tomen-tose when young, margin fibrillose, often densely imbricated.

Polystictus adustus, Fries, Hym. Eur., 549; Sacc., Syll. vi. no. 5146; Austr. Fung., p. 123, Hdbk. N.Z. Flora, p. 607.

Effuso-reflexed or entirely resupinate, forming large patches 5–25 Cm. across; flesh, thin, whitish, flexible when moist; pileus greyish, downy, indistinctly zoned, often rugulose; tubes very short; pores minute, rounded, whitish, then dingy-grey or lead-colour, blackish when dry, averaging 3–4 to 1 mm. of space; margin sterile, whitish; spores colourless, 4–5 × 2.5 μ.

On stumps, &c. New Zealand. Australia, Tasmania, India, Europe, United States, Cuba.

Very variable, sometimes entirely resupinate and resembling a Poria, at other times the margin becomes free. Distinguished by the grey pores and white margin. The pores become dark when bruised.

Polystictus catervatus, Berk., Fl N.Z., ii, p. 180, tab. cv, fig. 1; Hdbk. N.Z. Flora, p. 609; Sacc., Syll. vi, no. 5918.

Usually densely crowded; pileus about 1 cm. across, fan-shaped or irregularly reniform, margin often lobed or torn, narrowing below into a short, slender, stem-like base, white, silkily fibrillose; pores very shallow, minute, irregular, white; dissepiments thin, edge minutely toothed.

On split stems of Podocarpus spicata, Mission Station; on bark, Wellington, New Zealand.

Very frequently the adjoining pilei grow together at the margin, and form a continuous membrane. Very thin and delicate, altogether white. An endemic species, and apparently rare.

39. Poria, Pers.

Entirely resupinate; flesh usually very thin, attached throughout to the substratum; tubes usually short.

Poria, Persoon, Syn., p. 542.

In all probability many species included under the present genus may prove to be nothing more than degraded resupinate conditions of species of Fomes, Polystictus, or Polyporus.

Poria vaporaria, Fries, Syst. Myc., i, p. 682; Fl. N.Z., ii, p. 180; Hdbk. N.Z. Flora, p. 610; Cooke, Austr. Fung., p. 155; Sacc., Syll. vi, no. 6035.

Broadly effused, thin, inseparable, the white mycelium penetrating the matrix; pores large, angular or sinuous, white, then cream-colour, forming a continuous stratum.

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On dead trunks and branches. Northern Island, New Zealand. Victoria, Queensland, Western Australia, Malacca, Ceylon, Europe, North America.

Often broadly effused, or almost entirely covering fallen branches, inseparable from the matrix; flesh almost none; pores very variable, large, angular, or sinuous; often irregularly torn and more or less oblique, appearing as if sunk in the matrix, which is usually bark, whitish or pallid, becoming pale-ochraceous when dry. Pores often reaching 1 mm. in diameter. This species is sometimes destructive to worked wood, forming a white, spreading mycelium resembling the early stage of “dry rot.”

Poria mollusca, Fries, Syst. Myc., i, p. 384; Sacc., Syll. vi, no. 5936; Austr. Fung., p. 153.

Effused, thin, soft, white, margin fibrillose and giving off radiating strands; pores very shallow, minute, angular, dissepiments very thin and unequally torn, occupying the central portion of the patch, or scattered here and there in groups, ¼–⅓ mm. diameter.

On rotten wood, and on heaps of dead leaves. New Zealand. Victoria, Europe, United States.

Sometimes broadly effused; known by the fringed fibrillose margin; the partitions of the pores are very thin, and usually toothed or torn. Sometimes tinged with yellow. At first forming a mere byssoid margin, which gradually acquires moderate, rigid, subrotund and angular pores.

Poria hyalina, Berk., in Hooker's Flora Tasm., ii, p. 255 (1860); Sacc., Syll. vi, no. 5938; Cooke, Austr. Fung., p. 153.

Resupinate, very thin, white, more or less hyaline, circumference sterile, membranaceous, margin not byssoid; pores, very shallow, irregular in form, ¼–⅙ mm., dissepiments very thin.

On dead wood. New Zealand. Tasmania.

Very delicate, not thicker than paper; hymenium becoming much cracked, due to shrinkage during drying. The somewhat broad sterile border remains firmly attached to the matrix.

Poria leucoplaca, Berk., Fl. N.Z., ii, p. 180; Hdbk. N.Z. Flora, p. 609; Sacc., Syll. vi, no. 6092.

Entirely pure white, resupinate, thin; following the irregularities of the matrix, margin distinctly defined, every part covered by the small pores about ⅓ mm. diameter, dissepiments rather thick, edge pulverulent under a lens; flesh almost none.

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On dead branches. Northern Island, New Zealand. Malacca.

About 2 mm. thick, forming well - defined white crusts 2–10 cm. long.

Poria corticola, Fries, Syst. Myc., i, p. 385; Sacc., Syll. vi, no. 6093; Austr. Fung., p. 156.

White, then pallid, inseparable, often forming broad; thin, firm patches; not unfrequently more or leas sterile (without pores); pores naked, very shallow, small, roundish.

On dead bark. New Zealand. Queensland, Victoria, South Australia, Europe, United States, Brazil.

Position uncertain.

Polyporus diffissus, Berk., Fl. N.Z., ii, p. 180; Berk., Hdbk. N.Z. Flora, p. 610.

Fleshy, red, at length separating from the matrix, pores small, dissepiments thin, the membranaceous edge minutely toothed.

In the charred inside of a Fagus. New Zealand.

“Resupinate, effused, fleshy, of a bright red, at length tearing away from the matrix and leaving part of the substance behind; pores small; dissepiments thin; edge membranaceous, slightly toothed. This is probably a resupinate form of some anodermeous species, which has not at present been observed. Its bright colour, however, makes it very remarkable, on which account it is inserted here, though the specimen is by no means in a satisfactory state.” (Berk.)

No specimen exists at Kew, hence Berkeley's account, given above, cannot be supplemented. If this should prove to be a good species it would have to be known as Poria difissa.

40. Trimetes, Fries.

Pileus corky or woody; tubes penetrating unequally into the flesh of the pileus; pores roundish or more or less elongated radially.

Trimetes, Fries, Epicr., p. 488.

Trimetes is intermediate between Dœdalea and Fomes, differing from the former in the rounded or only slightly elongated pores, and from the latter in the tubes running up into the flesh of the pileus at different levels.

Trimetes epitephra, Berk., Journ. Linn. Soc. (Bot.), xiii, p. 165 (1873); Sacc., Syll. vi, no. 6240; Austr. Fung., p. 159.

Imbricated; pileus hoof-shaped, with a few more or less prominent ridges, ashy-brown, coarsely velvety; becoming almost

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smooth with age, margin whitish, hard, 1–2.5 cm. broad; pores pallid, very much decurrent or running down the bark, more or less elongated, rather large; dissepiments very thick.

On trunks and decaying wood. New Zealand. Adelaide, South Australia.

Remarkable for the very thick dissepiments or walls separating the pores. Might with almost equal propriety have been placed in the genus Dœdalea.

41. Dædalea, Pers.

Pileus woody or corky; pores elongated and irregularly sinuous; dissepiments or walls of tubes thick, flexible.

Dædalea, Persoon, Syn., p. 449.

Distinguished by the wide labyrinthiform pores, with thick, corky, and elastic walls.

Dædalea pendula, Berk., Fl. N.Z., ii, p. 180, tab. cv, fig. 4; Hdbk. N.Z. Flora, p. 610; Sacc., Syll. vi, no. 6394.

Imbricated; pileus 3–6 cm. broad and high, sessile, attached by the back and base, pendulous, irregularly cup-shaped, with the opening downwards, thin, flexible, strigose, reddish-grey; hymenium lining the cavity, pinkish-lilac., sparingly and vaguely scattered with tooth-like projections, and irregular shallow pores.

On dead wood. Ngawakatatara, New Zealand.

“Imbricated, coriaceous. Pilei 1 ½ in. long, pendulous, bursÆ-form, pale reddish-grey, tinged with lilac, sparingly zoned, clothed with short, strigose, matted brown hairs; margin tomentose. Hymenium tinged with lilac and reddish-grey, sparingly porous, with irregular tooth-like dissepiments, which are finely setulose. This, if fully grown, is scarcely a Dædalea in its characters, having more the hymenium of a Radulum; but it is evidently allied to such species as D. unicolor; and though the dissepiments are irregular, there are very evident pores, while in some parts there are as evident teeth. The species is at any rate undescribed, whatever may be thought of the genus.” (Berk.)

From the foregoing quotation it will be learned that the fungus under consideration is very imperfectly known, and, as it has not been collected since the type was found by Colenso, much remains to be learnt before its systematic position can be determined with certainty.

Dædalea confragosa, Pers., Syn., p. 501; Fl. N.Z., ii, p. 180; Hdbk. N.Z. Flora, p. 610; Sacc., Syll. vi, no. 6347.

Pileus sessile, horizontal, semicircular or subreniform, attached by a stout base, almost flat above, reddish-brown, in-

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distinctly zoned, roughened, 4–12 cm. across; flesh pale-wood colour, corky, 1–2 cm. thick at the base, becoming thinner towards the margin; tubes elongated; pores rounded or more frequently elongated and sinuous, grey, then brownish; dissepiments thick, flexible.

On dead wood, more especially Salix. Bay of Islands, Northern Island, New Zealand. Europe, United States.

Variable in size; the pileus is sometimes rough with large irregular elevations, at others only scabrid.

42. Favolus, Fries.

Pileus thin, tough, dimidiate or substipitate; pores large, elongated, extending radially from the point of attachment of the pileus.

Favolus, Fries, Elench., p. 44.

The pores appear to be formed from radially arranged lamellÆ or gills which anastomose and are connected by numerous lateral ridges. Perhaps most closely allied to Cantharellus, but more woody in texture.

Favolus intestinalis, Berk., in Hook. Journ. Bot., iii, p. 167 (1851); Hdbk. N.Z. Flora, p. 610; Sacc., Syll. vi, no. 6471.

Pileus thin, soft, irregularly reniform, margin variously undulate or lobed, attached by a very short lateral stem, which is sometimes almost obsolete, extending horizontally, 4–7 cm. long, upper surface very delicately pubescent when fresh; pores inferior, large, shallow, polygonal, up to 0.5 cm. across; spores broadly elliptical, hyaline.

On dead wood, among moss, &c. Northern Island, New-Zealand. India.

Entirely white when fresh, becoming very thin, translucent, and dingy-ochraceous when dry. Berkeley writes as follows of this species, which was described from Indian specimens: “A very singular esculent species, looking like a piece of tripe. The substance dries up so completely that the pores are visible from the upper side, as in some other species.”

43. Laschia, Fries.

Subgelatinous and tremelloid, thin, rigid when dry; under surface irregularly honeycomb-like in structure.

Laschia, Fries, Linnea, v, p. 533.

Distinguished by the flaccid subgelatinous texture, and the irregularly, hexagonal indentations on the under fertile surface; there are frequently protuberances on the upper surface of the pileus corresponding to the indentations on the lower surface.

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Laschia thwaitesii, Berk. and Broome, Journ. Lion. Soc. (Bot.), xiv, p. 58; Austr. Fung., p. 167; Sacc., Syll. vi, no. 6508.

Cæspitose; subgelatinous when moist; pileus convex, often oblique, thin, even, orange or yellowish-ochraceous, very minutely silky when young, 5–8 mm. across; pores rather large, irregular, yellowish; spores elliptical, obliquely apiculate, white, 6–7 × 4 μ; stem variable in length, up to 1.5 cm. long, sometimes very short, slender, whitish.

On dead stems of Rhipogonum. Pohangina River, New Zealand. Ceylon, Queensland.

A pretty and distinct species, growing mixed with Marasmius subsupinus, Berk., and sent to Kew, along with many other species, by Kirk. The pores are rather irregular, and in some specimens almost resemble gills connected by high transverse ridges. Differs from Marasmius in being subgelatinous when moist.

44. Merulius, Hall.

Resupinate, or with the margin more or less free and reflexed; substance usually somewhat soft and inclined to be gelatinous; pores very irregular in form, often formed from slightly raised, wavy, and anastomosing wrinkles.

Merulius, Hall., Helv., p. 150.

Differs from Poria in consistency and in larger and irregularly formed pores. Some species are destructive parasites; others, as M. lacrymans (“dry rot”), destroy worked wood.

Merulius corium, Fries, Elench., p. 58; Sacc., Syll. vi, no. 6532; Austr. Fung., p. 168.

Effused and resupinate, forming patches 3–10 cm. across, upper margin usually free and reflexed, substance very thin, flexible and tough; pileus whitish, silky; hymenium reticulatoporous, from pale-ochraceous to tan-colour, sometimes tinged lilac; spores oblong, 8–10 × 3–4 μ.

On trunks and branches. New Zealand. Australia, Tasmania, India, South Africa, Europe, United States.

Substance thin, often separable from the matrix. The hymenium is furnished with very slightly elevated ridges which anastomose to form an irregular reticulation that disappears towards the flat and sterile margin.

HydneÆ.

The sequence of general form is the same in the present family as in the PolyporeÆ. In the highest types there is a more or less thick or fleshy pileus supported on a central stem; next we descend to species having a lateral stem; then, the sessile

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bracket-shaped pileus; and finally to the resupinate condition where the entire fungus forms an incrusting more or less widely extending mass, inseparable from the matrix. The hymenium, instead of consisting of tubes having the inside lined with the hymenium as in the PolyporÆ, consists of closely packed solid spines or teeth, the entire surface of which bears the hymenium.

In the genus Hydnum the spines are slender and pointed, or awl-shaped, and vary in length in different species from 3 cm. to less than 1 mm. In other genera, as Radulum, the teeth are more or less flattened, obtuse, and often very irregular in size and form, whereas in some of the simpler resupinate genera, as Grandinia, the teeth are very much reduced, the hymenium being densely covered with very minute warts or granules.

The spines or teeth are very often more or less fringed or feathered at the tip when seen under a pocket-lens

Representatives of the family are apparently rare in the Southern Hemisphere, the most highly developed species being most abundant in the forests of northern Europe.

Some of the large fleshy species are edible.

A few species are destructive parasites, attacking timber and fruit-trees.

Analysis of the Genera.

45. Hydnum. Spines rounded, acute, distinct from each other at the base.

46. Irpex. Teeth obtuse, springing from anastomosing ridges.

47. Phlebia. Hymenium covered with delicate radiating folds or wrinkles.

48. Grandinia. Hymenium crowded with very minute warts.

45. Hydnum, Linn.

Hymenium inferior in stipitate and dimidiate species, superior in resupinate forms, covered with acute spines or teeth that are perfectly free from each other at the base.

Hydnum, Linn., Gen. Pl., no. 968.

Distinguished from allies by the acute awl-shaped or spine-like teeth arising free from each other.

Hydnum clathroides, Pallas, Russ. Reis., p. 2, fig. 3; Hdbk. N.Z. Flora, p. 611; Sacc., Syll. vi, no. 6683

Entirely grey, very much branched, trunk divided from the base, branches fasciculate and anastomosing laterally to form an irregular network; upper surface of branches papillose, under surface densely crowded with filiform spines 2–3 mm. long.

On wood (Knightia sp.). Northern Island, New Zealand. Asiatic Russia.

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Specimens imperfect, and identification hence doubtful; Native name, “pekepeke rione” (Coll.). Berk., in N.Z. Fl.

Hydnum coralloides, Scop., 2, p. 472; Sacc., Syll. vi, no. 6677; Cooke, Austr. Fung., 171.

Forming large tufts 5–12 in. across; white, becoming pallid with age; originating from a knob, which at once becomes divided into several tapering, crooked, pendulous branches which are ½ in. or more thick at the base, narrowing to about a line thick at the tip; spines springing from one side of the branches, pendulous, 3–8 lines long, awl-shaped, entire; spores hyaline, globose, 4–6 μ diameter.

On rotten wood, inside hollow trunks, &c. New Zealand. Queensland, Europe, Asiatic Siberia, United States.

“When old it forms tufts a foot or more in length, with flexuous angular branches, beset with incurved ramuli, bearing spines on the under side.” (Cooke.)

Hydnum udum, Fries, Syst. Myc., i, p. 422; Sacc., Syll. vi, no. 6795; Cooke, Austr. Fung., p. 173.

Patches thin, effused, inseparable from the matrix, sub-gelatinous, flesh-colour then dull-yellowish; spines crowded, unequal, 2–3 mm. long, awl-shaped or compressed, simple or toothed, coloured like the subiculum.

On dead wood. New Zealand. Queensland, New South Wales, Tasmania, Europe, United States.

Sometimes forming dingily coloured subgelatinous patches several inches long. Yellowish towards the margin when dry, the central part pale-fawn or dingy flesh-colour.

Hydnum niveum, Pers., Disp., tab. 4, f. 6–7; Sacc., Syll. vi, no. 6815.

Entirely resupinate, inseparable from the matrix, very thin and delicate, pure white, becoming pallid when dry, patches 2–8 cm. long; margin delicately byssoid; spines crowded, very minute, equal, glabrous.

On dead wood. New Zealand. Europe.

A very delicate species, resembling a mere film; spines or teeth very delicate, but under a lens are found to be acute, very uniform in size, glabrous.

Hydnum scopinellum, Berk., Fl. N.Z., ii, p. 181; Hdbk. N.Z. Flora, p. 611; Sacc., Syll. vi. no. 6825.

Effused, white, subiculum interwoven; spines tomentose at the base, tips penicillate.

On dead wood.

“Widely effused; subiculum composed of delicate inter-

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woven threads, which make the base of the aculei tomentose, tips penicillate.”

The above is the account of the species as given by Berkeley, and as there is no specimen in Berkeley's herbarium I am unable to add to the description. The species is placed in the resupinate section.

46. Irpex, Fries.

Pileus dimidiate or resupinate; teeth springing from irregularly arranged ridges or folds.

Irpex, Fries, Elench., p. 142.

The teeth ate somewhat irregular in form, and not so uniformly spine-like and pointed as in Hydnum.

Irpex brevis, Berk., Fl. N.Z., ii, p. 181; Hdbk. N.Z. Flora, p. 611; Sacc., Syll. vi, no. 6887.

Sessile, more or less fan-shaped or sometimes attached by a broad base, dimidiate, horizontal or slightly pendulous, about 1 cm. long by 1.5–2.5 cm. broad, very thin; pileus at first whitish, then brownish, more or less zoned and fibrillose; teeth, flattened, often irregularly divided, 2–3 mm. long, pale.

On dead bark, often growing among moss. New Zealand.

Apparently not uncommon, having been sent to Kew by Colenso on several occasions. An endemic species. Very variable in mode of growth. Sometimes several more or less fan-shaped pilei are crowded in an imbricate manner, at others they extend for a distance of several centimetres, attached by a broad base, the free portion overhanging and slightly drooping. The teeth are often decurrent for some distance below the free portion of the pileus. Finally the fungus is sometimes entirely resupinate, without a trace of free margin anywhere, the central portion being furnished with irregular plates or pores, almost resembling a Poria.

47. Phlebia, Fries.

Resupinate; hymenium covering the entire free surface, somewhat gelatinous, everywhere covered with fine radiating wrinkles or folds.

Phlebia, Fries, Syst. Myc., i, p. 426.

Distinguished by the closely crowded series of corrugation or ridges radiating irregularly from centre to margin of the hymenium.

Phlebia reflexa, Berk., in Hook. Journ. Bot., iii, p. 168 (1851); Sacc., Syll. vi, no. 6964; Cooke, Austr. Fung., p. 176.

Densely imbricated or superimposed, reflexed, thin, free portion 2–4 cm. broad, flaccid and tough when moist, rigid

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when dry; pileus covered with a dense coat of short down, purplish-brown towards the margin, often greyish-white near the line of attachment, irregularly zoned; hymenium dark-brown, often tinged purple, corrugated behind, almost even towards the margin; spores subglobose, about 4 μ diameter.

On logs of Fagus, &c. Northern Island, New Zealand. Victoria, Queensland, New South Wales, New Guinea, China, Sikkim, Himalayas, Africa.

At first resupinate and spreading widely, but easily detached, then broadly reflexed and dimidiate, 4–6 cm. long, 2–4 cm. broad; very tough and pliant when moist. Distinguished from species of Auricularia by the hymenium being covered with short, irregularly arranged, radiating ribs, giving it a wrinkled or corrugated appearance.

48. Grandinia, Fries.

Resupinate; thin, incrusting, hymenium minutely papillose or granulose, covering the entire exposed surface.

Grandinia, Fries, Epicr., p. 527.

When examined under a low power of the microscope, the tips of the minute granules covering the hymenium are usually found to be indented.

Grandinia crustosa, Fries, Epicr., p. 528; Sacc., Syll. vi, no. 6973; Austr. Fung., p. 176.

White or with a pallid or yellow tinge, irregularly effused, sometimes for several inches, closely adnate, thin, crustaceous, rather mealy at maturity; warts crowded, subglobose, minute, often collapsing at the apex, unequal.

On dead bark, wood, and on other fungi. New Zealand. Victoria, Ceylon, Europe.

ThelephoreÆ.

The fungi constituting this group show less differentiation and division of labour than is observable in the preceding groups. In the simplest forms the entire plant is resupinate or attached to the matrix at every point, and the upper surface is everywhere covered by the hymenium, with the exception of the silky or fibrous margin, which is the youngest or growing portion. When the fungus becomes partly free from the matrix or substance upon which it is growing, then the differentiation of the usually more or less membranous expansion is obvious. The under side, which is turned away from the light, is covered with the hymenium or spore-bearing portion, whereas the upper surface is completely sterile and usually more or less velvet or

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hispid. In many of the higher forms, as Stereum, Hymenochœte, &c., the sterile surface or pileus is brightly coloured and zoned.

The principal feature of the present group is the even surface of the hymenium, strictly confined to one side of the pileus, and the one-celled or non-septate basidia, as compared with the warts, spines, pores, or gills over which the spore-bearing surface is disposed in other groups. In some of the resupinate species of Corticium and Coniophora, where the flesh of the fungus is very thin, the hymenium often presents a waited or wrinkled surface, but on examination it will be found that this unevenness is due to the very thin substance of the fungus following the irregularities of the wood or other substance upon which it is growing.

Among the hymenial elements cystidia are very frequently highly differentiated, and are of importance in fixing the limits of genera. In Peniophora cystidia are very prominent, colourless, and the portion projecting above the general surface of the hymenium often coated with particles of oxalate of lime. In Hymenochœte, on the other hand, the cystidia are very thick-walled, rigid, and coloured brown. In the two genera just mentioned cystidia are so numerous in the hymenium, and project so much above the level of the basidia, that the surface presents a velvety appearance when seen through a pocket-lens.

In the most highly developed species there is a distinct central stem supporting a pileus which is usually depressed or funnel-shaped, thus resembling in general build an agaric, differing, however, in the absence of gills on the under surface of the pileus, which is quite even and more or less polished.

In some species of Stereum and Lachnocladium the pileus is cut up into numerous narrow strips or shreds, and superficially resembling certain species of Clavaria. In the last-named, however, the hymenium completely surrounds the branches, whereas in Stereum and Lachnocladium one side only of each narrow branch bears the hymenium, the opposite side being sterile and velvety or hairy.

Analysis of the Genera.

A. Spores coloured.

* Spores smooth.

49.Coniophora. Resupinate; surface dry and pulverulent.

     Spores warted or echinulate.

50.Thelephora. Substance dry and fibrous; hymenium often irregularly rugulose or nodulose.

51.Soppittitella. Subgelatinous, effused or variously incrusting twigs, grass, &c.

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B. Spores colourless.

* Hymenium minutely setulose with projecting cystidia.

52.Peniophora. Cystidia colourless.

53.Hymenochæte. Cystidia coloured.

     Hymenium glabrous.

54.Corticium. Entirely resupinate; hymenium usually cracked when dry.

55.Stereum. Effuso-reflexed; pileus silky or strigose; hymenium even.

56.Lachnocladium. Erect, narrowed to a stem-like base; pileus cut up into many very narrow segments; fertile on one side only.

57.Craterellus. Large, erect, funnel-shaped. Terrestrial.

58.Cyphella. Minute, cup-shaped, mouth open. On plants.

49. Coniophora, DC.

Broadly effused, resupinate, margin determinate or indeterminate; hymenium powdered with the smooth coloured spores; cystidia absent.

Coniophora, DC., Flor. Fr., vi, p. 34.

Forming broadly expanded, minutely powdery expansions on bark or wood.

Coniophora sulphurea, Mass., Journ. Linn. Soc. (Bot.), xxv, p. 133; Cooke, Austr. Fung., p. 196. Syn., Corticium sulphureum, Fries, Epicr., p. 561; Sacc., Syll. vi, no. 7535.

Broadly effused, margin bright sulphur-yellow, often fibrillose and running out in cord-like radiating strands; hymenium thickish, compact, almost waxy, brownish with a yellow tinge, cracking when dry; spores broadly elliptical, brownish-yellow, 11–12 × 8–10 μ.

On wood, bark dead leaves, &c. Northern Island, New Zealand. Tasmania, Ceylon, Europe, United States, Cuba.

Often sterile, and then very showy, as the mycelium and margin is clear yellow. Often extending for many inches.

50. Thelephora, Ehrh.

Varying from central-stemmed, through dimidiate, to resupinate; pileus usually fibrillose or strigose; hymenium usually wrinkled; spores coloured, warted, or echinulate. No cystidia.

Thelephora, Ehrh., Crypt., p. 178.

Differs from Coniophora in rough spores, which usually have a vinous or pale-purple tinge. Stereum differs in colourless spores.

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Thelephora vaga, Berk., Fl. N.Z., ii, p. 182; Hdbk. N.Z. Flora, p. 611; Sacc., Syll. vi, no. 7181.

Resupinate, variously incrusting, dry, dingy-brown; mycelium byssoid, creeping, loose; spores vinous-brown, irregularly globose, minutely warted, 6–8 μ.

On wood, heaps of dead leaves, &c. Ashburton, New Zealand.

“It grows under pine-trees, chiefly Pinus insignia. I have observed it growing on bare soil, among beds of dead pine-leaves, which become matted into a mass, and also at the roots of Dactylis glomerata, growing under P. insignis.” (W. W. Smith, Ashburton, New Zealand.)

51. Soppittiella, Mass.

Whitish at first, soft and subgelatinous, then becoming rigid, incrusting, form very variable; hymenium collapsing when dry and often tinged brown; spores coloured, spinulose.

Soppittiella, Mass., Brit. Fung. Fl., i, p. 106.

Distinguished by the soft substance when growing. Often creeping up living tufts of grass or other plants in an irregularly shaped fringed mass.

Soppittiella fastidiosa, Mass., Brit. Fung. Fl., i, p. 107 (1892). Thelephora fastidiosa, Berk., Outl., p. 268; Sacc., Syll. vi, no. 7161.

Whitish; forming broadly effused, incrusting, amorphous, or forming irregularly flattened branches; hymenium irregularly papillose, becoming rufescent with age or when bruised; spores broadly elliptical, rough, almost colourless, 6–7 × 4–5 μ. Smell of entire plant very fœtid, especially when bruised.

New Zealand. Europe.

White, becoming cream-colour, running as a thin soft film over everything in its way, sometimes forming free flattened branches. Silky or byssoid when young.

52. Peniophora, Cooke.

Resupinate, or with the extreme margin free and more or less raised; hymenium with projecting colourless spines or cystidia; spores colourless.

Peniophora, Cooke, Grev., vii., p. 20.

Differs from Hymenochœte in the cystidia being colourless; the projecting portions of the cystidia are often incrusted with particles of lime. Corticium differs in having no cystidia.

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Peniophora velutina, Cooke, Grev., viii, p. 21, pl. 125, fig. 15; Sacc., Syll. vi, no. 7701.

Broadly effused, rather fleshy, inseparable, margin running out into long branching strands; hymenium minutely velvety, cream-colour, often slightly tinged with pink or buff; cystidia cylindrical or attenuated upwards, 60–80 × 10–15 μ; spores elliptical with a minute apiculus, 10 × 5 μ.

On wood and bark. New Zealand. Europe, United States.

Often forming patches 5–10 cm. long. When perfectly developed the hymenium bristles with projecting cystidia when seen under a lens; these are more cylindrical and less incrusted with lime than usual. In some specimens the hymenium is very much cracked, in others quite continuous. The marginal radiating strands of mycelium often extend for many inches and connect several distinct fertile patches.

Peniophora ochracea, Mass., Journ. Linn. Soc. (Bot.), xxv, p. 150. Syn., Corticium ochraceum, Fries, Epicr., p. 563; Sacc., Syll. vi, no. 7600.

Broadly effused, inseparable, magin radiato-byssoid, soon disappearing; hymenium ochraceous, sparkling with very minute crystals of oxalate of lime when fresh, cracked when dry; cystidia fusoid, 40–60 × 18–22 μ; spores elliptical, hyaline, 10 × 5 μ.

On dead bark and wood. New Zealand. Europe, United States.

Closely resembling in habit and general appearance, colour, and in the presence of sparkling atoms on the hymenium Coniophora olivacea, but distinguished by the cystidia and smaller colourless spores.

Peniophora papyrina, Cooke, Grev., viii, p. 20, pl. 124, fig. 9; Austr. Fung., p. 191, fig. 82; Sacc., Syll. vi, no. 7688. Syn., Stereum papyrinum, Mont., Hdbk. N.Z. Flora, p. 612.

Very broadly effused, margin usually reflexed, very thin, coriaceous, strigose, grey, concentrically grooved, margin acute, tawny; hymenium umber, becoming purplish, minutely velvety; setÆ fusoid, 80–90 × 12–14 μ; spores subglobose, 6 μ diameter.

On bark and wood. Northern Island, New Zealand. Cuba.

Forming broadly effused, very thin patches, which follow the irregularities of the bark.

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53. Hymenochœte, Lév.

Pileus with a central stem, dimidiate or entirely resupinate; hymenium minutely setulose with projecting coloured cystidia; spores hyaline or coloured.

Hymenochœte, Lév., Ann. Sci. Nat., ser. 3, 1846, p. 150.

Distinguished at once by the numerous brown cystidia projecting from the surface of the hymenium. These can be easily seen with an ordinary pocket-lens.

Hymenochœte tabacina, Lév., Ann. Sci. Nat., ser. 3, vol. v, p. 152; Sacc., Syll. vi, no. 7428; Cooke, Austr. Fung., p. 189; Mass., Mon. The]., Journ. Linn. Soc. (Bot.), xxvii, p. 112; Mass., Brit. Fung. Fl., i, p. 117.

Subcoriaceous, thin, flaccid when moist, margin often reflexed, silky below, at length smooth, subferruginous, intermediate stratum and margin bright golden-yellow; hymenium cinnamon or rusty, usually with a tinge of purple, often cracked, minutely velvety; cystidia conico-acuminate, coloured, 80–130 × 10–14 μ; spores elliptical, olive, 5–6 × 3 μ.

On trunks, branches, &c. New Zealand. Victoria, New South Wales, Malacca, Europe, North and South America.

Distinguished by the golden-yellow margin of the hymenium and the coloured spores. Sometimes almost completely covering the under surface of fallen logs. When moist dirty ferruginous passing to mulberry-colour, rigid when dry, adnate, margin more or less free all round, often lobed, or free and reflexed above, rugulose. Hymenium often cracked when dry in lines radiating from the centre, or from several starting-points in broadly effused specimens.

Hymenochœte rhabarbarina, Cooke, Grev., viii, p. 148; Sacc., Syll. vi, no. 7467. Syn., Corticium rhabarbarinum, Berk., Fl. N.Z., ii, p. 184.

Effused, inseparable, hymenium minutely velvety, rusty-orange, margin paler, indeterminate, 8–12 cm. broad; setÆ acuminate, 30–40 × 7–9 μ; spores pale-olive, oblong-ellipsoid, 8 × 4 μ.

On bark. Northern Island, New Zealand.

The present species proves to be a true Hymenochœte, having the hymenium furnished with projecting acute setÆ or cystidia.

Hymenochœte kalchbrenneri, Mass., Journ. Linn. Soc. (Bot.), xxvii, p. 116; Cooke, Austr. Fung., p. 190.

Resupinate, rather dingy brown throughout, submembranaceous, broadly effused, loosely adnate to the matrix, margin

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rather well defined, wavy; hymenium minutely velvety; spores elliptical, hyaline, 7 × 5 μ; setÆ cylindrical or subclavate, often rough, 80–90 × 6–8 μ.

On dead trunks. New Zealand. Victoria, Queensland.

There is sometimes a violet tinge on the hymenium. The entire plant is sometimes almost separable from the matrix.

Hymenochœte phœa, Mass., Mon. Thel., Journ. Linn. Soc., xxvii, p. 98 (1891); Cooke, Austr. Fung., p. 188. Syn., Stereum phœum, Berk., Hdbk. N.Z. Flora, p. 612.

Pileus dimidiate, sessile, thin, coriaceous, flexible, zoned, velvety, bay, concentrically grooved, the grooves forming corresponding ridges on the rust-coloured minutely setulose hymenium, 6–10 cm. broad; setÆ scattered, conico-acuminate, 30–50 × 6–7 μ; spores subglobose, 4 × 3 μ.

On bark and wood. Northern Island, New Zealand. Tasmania.

Laterally attached, usually by a broad base, concentrically grooved, blackish-umber when dry; strigose with alternating dark and pale zones, margin crisped; hymenium umber.

Hymenochœte mougeotii, Mass., Sacc., Syll. vi, no. 7449; Austr. Fung., p. 189.

Broadly spreading, forming conspicuous blood-red or dull-red thin patches, sometimes there is an indication of a tinge of purple; closely adnate, margin determinate, dry, very minutely velvety under a lens, cracking when old and dry; cystidia conical, coloured, 30–75 × 5–8 μ; spores elliptic-fusoid, olive, 6–7 × 3–5 μ.

On dead trunks of pine and other wood. New Zealand. Victoria, Tasmania, Ceylon, India, central Europe.

Distinguished by the bright colour of the hymenium. The patches are sometimes 20–30 cm. long.

54. Corticium, Fries.

Entirely resupinate, or rarely with the extreme margin free; hymenium smooth, waxy, polished, becoming variously cracked when dry; spores colourless.

Corticium, Fries, Epicr., p. 556.

Forming resupinate inseparable patches on wood, bark, &c.

*Hymenium dingy flesh-colour.

Corticium nudum, Fries, Epicr., p. 564; Sacc., Syll. vi, no. 7609; Austr. Fung., p. 194.

Often forming thin patches 3–7 cm. long, waxy, margin determinate, glabrous; hymenium flesh-colour, becoming pale,

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cracked when dry, minutely pulverulent under a lens; spores elliptical, 12–13 × 4 μ.

On dead bark. New Zealand. Queensland, South Africa, Europe.

Distinguished by the pale-flesh-coloured minutely pulverulent hymenium.

Corticium polygonium, Fries, Hym. Eur., p. 655; Berk., Hdbk. N.Z. Flora, p, 613; Sacc., Syll. vi, no. 7611.

Closely adnate, inseparable, outline sharp, extreme margin byssoid, soon becoming hard and rigid, 5–10 cm. broad; hymenium dingy flesh-colour, primrose, usually much cracked or nodulose; spores narrowly elliptical, 14–16 × 5–7 μ.

On dead bark and wood. Northern Island, New Zealand. Europe, United States.

Usually extending under the form of small, distinct, Tubercularia - like pustules, which eventually usually become confluent, thick, separating from each other more or less when dry; giving the patch a cracked or tesellated appearance; sometimes continuous, and then the surface is more or less tuberculose; margin thin, adnate, byssoid; hymenium primrose, pinkish, black, or dingy-ochraceous.

Hymenium white at first, sometimes becoming pale-tan or pale-rose colour.

Corticium auberianum, Montag., Crypt. Cuba, p. 372; Sacc., Syll. vi, no. 7552; Austr. Fung., p. 194.

Closely adnate, at first orbicular, several patches soon becoming confluent or growing into each other and forming broadly extended patches, 5–10 cm. long and broad, very thin, at first snow-white and minutely primrose, finally glabrous and tinged with dingy-yellow or grey, finely crooked when dry; margin persistently minutely floccose or fibrillose; spores elliptical, 6–7 × 4 μ.

On bark, &c. New Zealand. Australia, Patagonia, Cuba, United States.

Readily distinguished by the hymenium, which is snow-white and primrose when young. When the fungus is old it sometimes partly peels away from the matrix.

Corticium, albidum, Mass. Syn., Aleurodiscus albidus, Mass., Grev., xvii, p. 55; Cooke, Austr. Fung., p. 193, fig. 83.

At first concave, rather fleshy, white, outside and margin tomentose, at first incurved then becoming extended and flattened, up to 6 mm. diameter, often confluent and forming rather large patches; hymenium white, minute mealy, cracking slightly when dry; spores elliptical, 10–12 × 9 μ.

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On branches. Northern Island, New Zealand. Queensland.

This species was at first incorrectly determined as Aleurodiscus oakesii. The last-named is not known to occur in New-Zealand.

Corticium scutellare, Berk. and Curt., Grev., ii, p. 4; Brit. Fung. Fl., i, p. 121; Sacc., Syll. vi, no. 7647.

Broadly effused, thin, quite inseparable from the matrix, margin indistinct, whitish, then dirty pale-tan-colour or tawny, waxy, smooth, very much cracked into polygonal portions, interstices white, silky; spores elliptical, 5 × 3 μ.

On wood, dead herbaceous stems, &c. Recognised by the brownish areolately cracked hymenium, and small spores.

Corticium leve, Pers., Disp., p. 30; Sacc., Syll. vi, no. 7530; Cooke, Austr. Fung., p. 194; Fl. N.Z., ii, p. 184; Hdbk. N.Z. Flora, p. 613.

Forming very thin patches from 5–10 cm. in diameter; hymenium smooth and with a more or less polished appearance, old-ivory colour often suffused with a flesh-colour or rosy tinge, becoming cracked when dry, the interstices silky, margin byssoid; spores elliptical, often slightly curved, 10–12 × 6–7 p.

On dead wood and bark. Northern Island, New Zealand. Victoria, Queensland, Tasmania, Ceylon, Europe, North and South America.

Some forms of this species closely resemble, superficially, Peniophora rosea, but can at once be recognised under the microscope, or even under a pocket-lens, by the absence of projecting cystidia in the hymenium.

Hymenium tinged green.

Corticium viride, Berk., Fl. N.Z., ii, p. 184; Hdbk. N.Z. Flora, p. 613.

Olive-green, crustaceous, effused, cracked; margin very thin, membranous, scarcely byssoid, livid; spores large, elliptic or subglobose.

On dead bark and wood. Northern Island, New Zealand.

Effused, forming small confluent patches of a yellow olivaceous green, with a very thin, membranous, scarcely byssoid, livid margin; hymenium cracked; spores subglobose or elliptic, very large 1/1750 in. long (= about 14 μ). Analogous to Hydnum viride. When old it acquires a darker tinge. There is no specimen of this species at Kew.

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Corticium terreum, Berk., Fl. N.Z., ii, p. 184. C. terreum, Hdbk. N.Z. Flora, p. 613.

Examination of the type specimen at Kew shows this to be an immature specimen of some Thelephora.

55. Stereum, Pers.

Pileus with a central stem, or dimidiate and imbricated; pileus silky or strigose; hymenium smooth; spores colourless (rarely tinged with colour).

Stereum, Persoon, Obs. Myc., p. 35.

The leading features of the present genus are the smooth or glabrous hymenium, and the velvety or strigose pileus. Some species are destructive wound-parasites, attacking forest trees.

*Pileus supported on a central stem.

Stereum sowerbeii, Berk., Fl. N.Z., ii, p. 182; Hdbk. N.Z. Flora, p. 612; Mass., Brit. Fung. Fl., i, p. 129. Syn., Elvella pannosa, Sowerby, Fung., tab. 155.

White; pileus funnel-shaped 2–2.5 cm. across, rough with projecting points, but not velvety, margin variously incised; stem up to 1 cm. long, central; spores elliptical, 5 × 3 μ, hyaline.

On the ground. Northern Island, New Zealand, Tasmania, Australia, Britain, United States.

A very beautiful species, snow-white, tinged with pale-buff when old, and of a waxy appearance when fresh; sometimes with a distinct round stem ½ in. or more in height, at others several plants grow close together, having their stems more or less confluent at the base. It has no relationship with Cladoderris, as suggested by Fries in Sum. Veg. Scand., p. 332.

Pileus attached laterally.

Stereum lobatum, Fries, Epicr., p. 547; Sacc., Syll. vi, no. 7311; Hdbk. N.Z. Flora, p. 612; Austr. Fung., p. 184 (all in part). Syn., Stereum luteo-badium, Fr.; Stereum boryanum, Fr.; Stereum ostrea, Nees; Stereum sprucei, Berk.; Stereum perlatum, Berk.

Sessile, often imbricated or running on horizontally, pilei sessile sometimes fan-shaped and fixed by the narrow portion, horizontal, margin entire or variously lobed, thin, rigid, upper sterile surface tomentose or minutely velvety, orange or brownish, with darker concentric bands of colour, becoming glabrous towards the margin; hymenium usually bright ochraceous, sometimes duller, and verging on a greyish tint; 8–15 cm. across; spores subglobose, 5–6 μ diameter.

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On trunks, fallen timber, &c. Middle and Northern Islands, New Zealand. Widely distributed, especially in tropical and subtropical countries both in the Old and New World.

A variable species both in size, colour, and amount of rugosity of the pileus, but readily distinguished by the thin rigid substance; velvety zoned pileus, and smooth ochraceous or greyish hymenium.

Stereum lobatum, Fries, Epicr., p. 547; Hdbk. N.Z. Flora, p. 612; Sacc., Syll. vi, no. 7311; Austr. Fung., p. 184. Syn., Stereum perlatum, Berk., in Hook. Journ. iv, 1842, p. 153; Stereum sprucei, Berk., Journ. Linn. Soc., x, p. 331; Stereum luteo-badium, Fr., Epicr., p. 547; Stereum boryanum, Fr., Epicr., p. 547; Stereum ostrea, Nees, Nov. Act. Nat. Cur., xiii, p. 13, pl. 2.

Pileus thin, rigid, umbonato-sessile, coriaceous, tomentose, usually ochraceous and often zoned with bay, margin almost glabrous, 10–40 cm. across; hymenium smooth, even, pallid; spores subglobose, 5–6 μ.

On dead wood. Northern and Middle Islands, New Zealand. Tasmania, Victoria, New South Wales, Queensland, New Guinea, Philippines, India, Ceylon, Bourbon, Malay Peninsula, Java, Malacca, Surinam, Seychelles, southern United States, Mexico, Cuba, Brazil, Venezuela, Madagascar, Peru, Mauritius.

A widely distributed and variable species, distinguished by its large size, thin substance, and pale dull-yellow hymenium. The margin is often variously lobed.

Stereum cinereo-badium, Klotzsch, Nov. Act., 19, tab. v, fig. 3; Hook., Fl. N.Z., ii, p. 182; Sacc., Syll. vi, no. 7337.

Pileus dimidiate, sessile, robust, coriaceous, tomentose, margined, chestnut-brown, zones smooth, black; hymenium smooth, glaucous, flesh-coloured.

On dead wood. Northern Island, New Zealand. South America.

An imperfectly known species. There is no specimen in the Kew Herbarium from New Zealand or elsewhere.

Entirely resupinate, or with the margin only more or less free.

Stereum illudens, Berk., in Hook. Journ., iv, p. 59; Sacc., Syll. vi, no, 7329; Austr. Fung., p. 185.

Effused on the matrix, the upper part free, horizontal, margin usually crisped and wavy, substance very thin, coriaceous, rigid when dry; upper surface of pileus coarsely velvety, brown, often with paler zones, radially plicate, 2–6 cm. long, 2–3 cm.

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wide; hymenium even, smooth, rufous or brown; spores elliptical, 6–7 × 4 μ.

On dead logs, branches, &c. New Zealand. Tasmania, Victoria, New South Wales, Queensland, Western and South Australia, Venezuela.

A characteristic Australasian species, distinguished by the dark, hairy, waved pileus and dark-coloured hymenium.

Stereum pannosum, Cooke, Grev., viii, p. 56.

Pileus coriaceous, becoming rigid when dry, effused, the upper portion free and reflexed, 3–6 cm. across, sterile surface dingy - grey, indistinctly zoned, hirsute; hymenium glabrous, dingy - grey, becoming primrose, cracked when dry; spores elliptical, 5–6 × 3–4 μ.

On dead bark. New Zealand. At present only known from New Zealand.

Effused, free margin often torn or split. Distinguished by the dingy-grey or dull-lead colouring of every part. Stereum illudens differs in brown tinge of every part, and S. lugubre in the blackish papillose hymenium.

Stereum lugubre, Cooke, Grev., xii, p. 85.

Coriaceous, rigid; pileus effused and reflexed, about 2.5 cm. deep, tomentose, zoned, cinereous, becoming pallid, zones darker; margin rather acute, pallid; hymenium somewhat papillose, smooth, naked, black.

On logs. New Zealand.

A very distinct species by its black obtusely papillate hymenium. Pileus about 1 in. deep, often densely imbricated and extending laterally several inches. There is no specimen of this species present in the Kew Herbarium, hence I am unable to supplement Cooke's original diagnosis, reproduced above.

Stereum rugosum, Fries, Epicr., p. 552; Fl. N.Z., ii, p. 183; Hdbk. N.Z. Flora, p. 612; Sacc., Syll. vi, no. 7336; Cooke, Austr. Fung., p. 187.

Broadly effused, sometimes shortly reflexed, coriaceous, becoming thickish and rigid, sterile surface at length smooth, brownish; hymenium pale greyish-yellow, primrose, becoming red when cut or bruised; spores cylindrico-elliptical, straight, 11–12 × 4–6 μ.

On bark and dead wood; sometimes growing on living trees and proving to be a very destructive parasite. Northern Inland, New Zealand. Victoria, New South Wales, Western Australia, Europe, North and South America.

Very variable in form, wholly adnate, partly reflexed, or

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sometimes almost saucer-shaped and attached by a central point when young. Agrees with Stereum sanguinolentum in becoming red when bruised, but distinguished by the thicker rigid substance and in the larger straight spores. The hymenium is sometimes pale-yellow, at others greyish or livid

Stereum vellereum, Berk., in Hook. Fl. N.Z., p. 183; Hdbk. N.Z. Flora, p. 612; Cooke, Austr. Fung., p. 184; Sacc., Syll. vi, no. 7367.

Resupinate with the margin free, or fan-shaped, attached by a narrow base, and imbricated, thin, 2–5 cm. across, greyish, velvety, margin zoned and often lobed; hymenium ochraceous, even, glabrous; spores subglobose, 4–5 μ diameter.

On branches and twigs. Northern and Middle Islands, New Zealand. Victoria, Singapore, north-west Himalaya.

Usually growing on small branches, and then resembling Hymenochœte tabacina in habit, with broad, free, more or less lobed wings; substance thin; when growing on thick branches or logs, often imbricated and narrowed at the base. Resembling Stereum hirsutum in colour and habit, hut thinner and with different spores.

Stereum hirsutum, Fries, Epicr., p. 549; Hdbk. N.Z. Flora, p. 612; Sacc., Syll. vi, no. 7288; Austr Fung., p. 185

Entirely resupinate, or more frequently with a free margin which is often more or less lobed or wavy; pileus coarsely strigose, 3–8 cm. broad, dingy-ochraceous, becoming pale and greyish, indistinctly zoned, thin and coriaceous; hymenium even, glabrous, naked, ochraceous or tan-colour; spores elliptical, 6 μ long.

On trunks and branches. Northern and Middle Islands, New Zealand. Tasmania, Victoria, Queensland, New South Wales, Western Australia, South Australia, Java, India, North and South America.

Very variable in form. When growing on a broad surface often wholly resupinate or with a very narrow free margin. On smaller branches there is often a broad, free, reflexed portion, or several such overlapping. Pileus coarsely velvety or strigose; hymenium usually bright ochraceous, often with varying shades of pink or grey.

Stereum ochroleucum, Fries, Hym. Eur., p. 639; Sacc., Syll. vi, no. 7283; Austr. Fung., p. 186. Syn., Corticium ochroleucum, Fries, Epicr., p. 557; Berk., Hdbk. N.Z. Flora, p. 613.

Coriaceous, thickish, separable from the matrix, sometimes entirely resupinate, at others more or less free round the margin,

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or almost entirely free and fixed by a broad effused base, flaccid, silky, dingy - ochraceous, 8–15 cm. broad; hymenium even, glabrous, pale - ochraceous, cracked when dry; spores broadly elliptical, 8 × 6–7 μ.

On dead wood and bark. Middle Island, New Zealand. Queensland, Tasmania, India, Europe, North America, Cuba, Venezuela.

Ochraceous, villose or strigose, often becoming bald when old; sometimes broadly effused and entirety adnate, in others the margin only free and upturned, in others again quite free and fixed by a narrow base. It is not unusual to meet with all transitions from entirely adnate to the flabelliform condition on the same trunk. Hymenium pale-ochre, smooth, cracked, especially when dry; the latter character separates it from Stereum hirsutum, and also from Corticium, which the adnate form resembles superficially.

Stereum latissimum, Berk., Fl. N.Z., ii, p. 183: Hdbk. N.Z. Flora, p. 613; Sacc., Syll. vi, no. 7419.

Forming broad, very thin, chalk-white patches, minutely subtomentose, margin abrupt.

On bark. Northern Island, New Zealand.

Forming patches many inches in length and breadth, very thin, following all the inequalities of the matrix, chalk-white; under the lens very minutely subtomentose; margin abrupt, by no means byssoid.

An imperfectly described species of which no type specimen is known to exist.

56. Lachnocladium, Lév.

Stipitate, much branched, branches narrow, one side tomentose and sterile, the other covered with the smooth hymenium; spores colourless.

Lachnocladium, Lév., in Orb. Dict., viii, p. 487.

Closely allied to Stereum, differing mainly in the much-divided pileus. Superficially resembling some species of much-branched Clavaria; differing in the tougher texture and in the hymenium being confined to one side of the branches.

Lachnodadium flagelliforme, Cooke, Austr. Fung., p. 179, fig. 79. Syn., Clavaria flagelliformis, Berk., Fl. N.Z., ii, p. 186; Hdbk. N.Z. Flora, p. 614; Sacc., Syll. vi, no. 8018.

Very much branched, divided to the base or nearly so, branches tufted, cylindrical, fastigiate, forked, tips acute, and

– 36 –

undivided; spores hyaline, broadly elliptical, 5 × 3.5 μ. Entire fungus dingy-white or pale-brown, 4–5 cm. high.

On the ground, probably springing from buried twigs. Bay of Islands, Northern Island, New Zealand.

57. Craterellus, Fries.

Terrestrial. Plant altogether more or less funnel-shaped, hymenium covering the outside of the funnel, glabrous, smooth or rugulose.

Craterellus, Fries, Epicr., p. 531

Resembling superficially some species of Cantharellus; the latter, however, are distinguished by the presence of narrow, thick, irregularly forked gills running down the outside of the pileus.

Craterellus insignis, Cooke, Grev., xix, p. 2; Sacc., Syll. ix, no. 880.

Erect, more or less tufted, sometimes grown together, 3–4 cm. long, about 2.5 cm. broad; pileus fan - shaped, tan - colour, irregularly striate, margin lobed and wavy, flesh thin; hymenium waxy, rugulose, darker than the pileus; stem slender, expanding upwards into the pileus, tan-colour; spores elliptical, tinged brown, 2–5-3 × 1.5 μ.

On dead trunks. New Zealand.

Resembling in general appearance some of the central-stemmed species of Stereum, but differing in the soft fleshy consistency

58. Cyphella, Fries.

Minute; cup-shaped, mouth not contracted, often narrowed into a stem-like base; hymenium internal; outside velvety or downy.

Cyphella, Fries, Syst. Myc., ii, p. 201.

Minute, often clustered; resembling in habit a small Peziza.

Cyphella densa, Berk., Fl. N.Z., ii, p. 184; Hdbk. N.Z. Flora, p. 614; Sacc., Syll. vi, no. 7837.

Gregarious, obliquely funnel-shaped, fawn-coloured, pendulous, attached by a short narrow stem-like base, very minutely pilose, flexible; hymenium lining the inside of the funnel, smooth, even; spores broadly elliptical, hyaline, 7 × 5 μ.

On living bark of Corynocarpus. Cape Kidnappers, Northern Island, New Zealand.

The pilei are obliquely funnel-shaped or more exactly resemble the head of any ordinary clay pipe, suspended by a very short stem. A very fine endemic species.

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Cyphella filicicola, Berk. and Curt., Grev., ii, p. 5 (1873); Sacc., Syll. vi, no. 7898.

Scattered, sessile but attached by a narrowed base, more or less pendulous, obliquely funnel-shaped, often irregular in form, umber or brownish, externally minutely downy under a lens, 2–3 mm. long.

On dead fern-stems. New Zealand. Carolina, United States.

Shaped like the bowl of a smoking-pipe, and attached by a very short stem, or, rather, the narrowed base of the head of the pipe, the cavity pointing downwards.

Cyphella albo-violascens, Karst., Fung. Fenn. Exs., no. 715; Sacc., Syll. vi, no. 7817; Austr. Fung., p. 196.

Gregarious, sessile, 1–3 mm. diameter, globose and closed when young, then hemispherical, externally snow-white and densely downy; hymenium even, more or less tinged with violet; spores colourless, elliptical, usually slightly inœquilateral, 12–15 × 8–9 μ.

On wood, bark, twigs, &c. New Zealand. Australia, Cape of Good Hope, Europe, South America, United States.

Resembling a minute downy Peziza, for which it was mistaken by early authors. Often proliferous; hymenium and margin becoming blackish.

ClavarieÆ.

An entire absence of differentiation into a sterile (pileus) and fertile (hymenium) surface respectively, and the even hymenium, are the characteristic features of the present group.

In the simpler forms the hymenophore is club-shaped, every portion of the club being fertile or covered with the hymenium. In other species the club becomes more or less divided, whereas in numerous species the fertile portion is broken up into numerous branches, the whole resembling a much-branched tree or coral in miniature.

The species are usually small, often brightly coloured, and with few exceptions grow on the ground. All the species are edible. Some of the minute species spring from sclerotia, and amongst these are parasites on various cultivated plants; but the injury caused by members of the Clavarieœ is practically a negligible quantity.

Analysis of the Genera.

59.Clavaria. Soft and fleshy, simple or much branched, branches terete, axils usually rounded.

60.Pistillaria. Minute, club-shaped, simple, rigid and horny when dry.

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59. Clavaria, Fries.

Sporophore erect, club-shaped or fusiform, or variously and often excessively branched, axils of branches often rounded; spores colourless or coloured. Basidia two- or four-spored.

Clavaria, Fries, Syst. Myc., i, p. 465.

Usually terrestrial, rarely growing on wood. Calocera superficially resembles the branched species of Clavaria, but differs in the partly gelatinous consistency and different structure of the basidia. Lachnocladium differs in tough consistency and in the branches being flattened and having the hymenium on one side only.

*Spores ochraceous or yellow.

Clavaria flaccida, Fries, Syst. Myc., i, p. 471; Hdbk. N.Z. Flora, p. 615; Sacc., Syll. vi, no. 7972.

Slender, very much branched, entirely ochraceous, stem very short, branches crowded, repeatedly forked, upper axils rounded and the acute terminal branchlets converging; spores broadly elliptical, ochraceous, 4–5 × 3 μ.

On the ground in woods among moss, &c. Sometimes growing on masses of dead leaves, &c. Northern Island, New Zealand. Europe.

Varying from 2–5 cm. high; colour clear ochraceous without any tinge of brown; does not become green when bruised; terminal branchlets converging like callipers; mycelium whitish, creeping over leaves, &c.; stem sometimes 2 cm. long, at others almost obsolete.

Clavaria crispula, Fries, Syst. Myc., i, p. 470; Hdbk, N.Z. Flora, p. 615; Sacc., Syll. vi, no. 7991; Cooke, Austr. Fung., p. 201.

Pale yellow-brown or tan-colour, becoming ochraceous; stem rather slender, with downy rooting strands of mycelium, 4–7 cm. high; branches numerous, wavy, spreading, repeatedly dividing, terminal branchlets acute, spreading; spores pale-yellow, elliptical, 5 × 3 μ.

On the ground, at base of trunks, &c. Northern Island, New Zealand. Western Australia, Europe, United States, Brazil.

Stem thin; branches numerous, lax, rather wavy or flexuous.

Spores hyaline.

Clavaria flava, Schæffer, Fung. Bavar., tab. 175; Sacc., Syll. vi, no. 7929; Cooke, Austr. Fung., p. 198. Syn., Clavaria, lutea, Hook., Fl. N.Z., ii, p. 185; Hdbk. N.Z. Flora, p. 614.

Stem stout, short, white, breaking up into numerous rounded, tapering, crowded, even-topped, yellow branches,

– 39 –

8–14 cm. high; spores elliptical, hyaline or with a slight tinge of yellow, 8–10 × 4–5 μ.

On the ground, in woods. New Zealand. Victoria, Queensland, Europe, United States.

Edible, as are all known species of Clavaria. Brittle; stem often 2–3 cm. thick; forming dense tufts of crowded branches; yellow colour usually most pronounced at the tips of the branches.

Clavaria arborescens, Berk., Fl. N.Z., ii, p. 186; Hdbk. N.Z. Flora, 614; Sacc., Syll. vi, no. 8022.

Amethyst-colour; stem slightly wavy, 2–3 cm. high, slightly thickened upwards, slender, dividing at the apex into a few main branches that bear short fastigiate branchlets at their tips; spores hyaline, elliptical, 6 × 4 μ.

On the ground. Bay of Islands, Northern Island, New Zealand.

Berkeley considers this species as showing affinity with Clavaria macropus. To me it appears to resemble a slender form of C. cinerea.

Clavaria colensoi, Berk., Fl. N.Z., ii, p. 186; Hdbk. N.Z. Flora, p. 615; Sacc., Syll. vi, no. 8039; Cooke, Austr. Fung., p. 201.

Stem compressed, short, breaking up into several primary branches, which in turn become inflated at the apex and bear several slender secondary branchlets divided at the acute tips, 2–3 cm. high; spores elliptical, 5x 3 μ.

On dead wood and on the ground. Northern Island, New Zealand. Queensland.

All the branches have a tendency to become flattened, axils of branches rounded. The swollen apices of the branches are sometimes more or less excavated and the branchlets originate from the margin of the cup.

No account is given in the original description as to the colour of the plant. The following is Berkeley's description of this species: About 1 in. high, attached to the soft decayed wood by a few short towy fibres, which, like the whole plant, are brown when dry. Stem mostly compressed, branched from the base or a little above it, repeatedly forked; branches sub-fastigiate, delicate; apices forked, very acute. Closely allied to C. delicata, but the brown fibres by which it is attached, and other points, forbid its association with that species, of which I have authentic specimens from Fries.” (Fl. N.Z., ii, p. 186.)

– 40 –

Clavaria mucida, Pers, Comm., tab. 2, fig. 3; Sacc., Syll. vi, no. 8125; Cooke, Austr. Fung., p. 203.

Gregarious but not usually tufted, simple or sparingly branched, branches linear, tip sometimes cristate or divided into fine short branchlets, white or with a tinge of yellow or rose, surface even, 1–2 cm. high, slender; spores hyaline, averaging 6 × 3 μ.

On wet rotten wood. New Zealand. New South Wales, Europe, United States.

Clavaria contorta, Holmsk, Ot. i, p. 29.

Erumpent; in clusters of 2–5 specimens, simple, stuffed, variously twisted, contorted, and wrinkled, primrose, yellowish, often with a red or brown tinge, about 1 in. high; spores white, subglobose, about 4–5 μ diameter.

On fallen branches. New Zealand. Europe, United States.

Easily known by growing on wood, and in being erumpent, or bursting through the bark

Clavaria pusio, Berk., Fl. N.Z., ii, p. 185; Hdbk. N.Z. Flora, p. 614; Sacc., Syll. vi, no. 8017.

Stem slender, thickened upwards, where it divides into a few cylindrical acute branches equal in length to the stem and spreading at an acute angle, rarely divided, 1.5–3 cm. high; spores elliptical, hyaline.

On the ground. Northern Island, New Zealand.

The colour is brownish when dry, but it is probably paler or whitish when fresh.

Clavaria inœqualis, Flor. Dan., p. 74, fig. 4; Hdbk. N.Z. Flora, p. 615; Sacc., Syll. vi, no. 8069; Austr. Fung., p. 202.

Yellow, gregarious or fasciculate, fragile, stuffed, clavate, apex obtuse, simple or sometimes forked, 4–7 cm. high; spores elliptical, colourless, 9–10 × 5 μ.

Among grass and moss. Bay of Islands, Northern Island, New Zealand. Tasmania, New South Wales, Victoria, Europe, United States, Ceylon.

Scattered or in small loose tufts, clubs clavate or cylindrical; apex obtuse, sometimes forked or variously cut and divided, sometimes compressed, but not distinctly apiculate, and brown.

Clavaria misella, Berk. and Curt., Journ. Linn. Soc. (Bot.), x, p. 339; Sacc., Syll. vi, no. 8139.

Entirely white, simple or rarely with a single branch springing from near the base, slightly clavate, quite slender, 1–2 cm. high, base somewhat spongy; spores hyaline, subglobose, about 4 μ diameter.

– 41 –

Growing on living moss. Middle Island, New Zealand. Cuba.

Becoming opaque and remaining even when dry, which, in addition to the different spores, distinguish it from Clavaria paupercula, Berk. and Curt., a small species also growing on moss.

60. Pistillaria, Fries.

Minute, club-shaped, simple, becoming cartilaginous when dry.

Pistillaria, Fries, Hym. Eur., p. 686.

Very closely allied to Clavaria, if distinct as a genus. Differing mainly in minute size, and in becoming cartilaginous and rigid when dry.

Pistillaria ovata, Fries, Syst. Myc., i, p. 497; Hdbk. N.Z. Flora, p. 615; Sacc., Syll. vi, no. 8259.

Club obovate or ellipsoid, often more or less compressed, sometimes slightly lobed, white, hollow; stem short, glabrous, pellucid; entire plant 3–7 mm. high; spores elliptical, 7–8 × 3–5 μ.

On dead leaves, herbaceous stems, &c. Northern Island, New Zealand. Europe.

Variable in size and form, but always minute; distinguished by the short, polished, hyaline stem.

TremellineÆ.

The members of this group are characterized by the more or less gelatinous nature of the entire fungus when growing. During the process of drying the plant shrinks very much, and becomes hard and horny, expanding and becoming gelatinous again when soaked in water.

As representing the simplest structure presented by the Basidiomycetes, the basidia depart from what may be termed the normal or typical form in various genera. In Dacryomyces the basidium is cylindrical, with two much elongated and stout sterigmata at its apex. In this genus the spores are septate. In Tremella the basidium is stout and broadly clavate or obpyriform, with four stout sterigmata at its apex. When the sterigmata are just commencing growth the basidium, viewed from above, shows two apparent lines crossing at right angles, and the basidium has been spoken of as cruciate. This appearance is caused by the bases of the four incipient sterigmata. In the genera Auricularia, Hirneola and Septobasidium the basidia are transversely septate, each cell bearing a single spore. This form of basidium agrees morphologically with the sporophores in the Uredineœ, with which, according to Brefeld, the Basidiomycetes are allied.

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Analysis of the Genera.

*Basidia subglobose, longitudinally quadripartite in a cruciate manner at maturity, and producing at the apex four elongated sterigmata.

61.Exidia. Cup-shaped or variously lobed; spores sausage-shaped curved; sterile surface minutely velvety.

62.Tremella. Brain-like or variously lobed and contorted; spores globose or ovoid.

63.Næmatelia. Firm, convex, with a central firm nucleus.

**Basidia cylindrical or clavate, divided at the apex into two long sterigmata.

64.Dacryomyces. Small, pulvinate, gyrose.

65.Guepinia. Irregularly cup-shaped or flabellate; hymenium on one surface only, the other sterile and silky.

66.Calocera. Erect, simple or branched, subcylindrical.

***Basidia elongated or fusoid, transversely septate, each cell producing one spore.

67.Auricularia. Broadly attached, effuso-reflexed, upper surface sterile and strigose.

68.Hirneola. Cartilaginous, human-ear shaped, attached by a narrow point, sterile surface minutely velvety.

69.Septobasidium. Resupinate, not gelatinous but leathery.

61. Exidia, Fries.

Inflated, tremelloid, marginate or vaguely effused, often minutely papillose, black or dusky.

Exidia, Flies, Syst. Myc., ii, p. 224.

Forming irregular subgelatinous masses on dead wood and branches. The spores become 1—many septate on germination, each cell of the spore giving origin to a very short promycelium, bearing a cluster of strongly curved sporidiola.

Exidia albida, Bref., Unters. Mykol., vii., p. 94, pl. v, fig. 14; Sacc., Syll. vi. no. 8352; Austr. Fung., p. 207. Syn., Tremella albida, Huds.

Gelatinous, bursting through cracks in bark and wood, under the form of wavy or contorted heaps 1–3 cm. across; white at first, soon becoming dingy-yellow or brown; when mature primrose, with the dense mass of spores resting on the surface; spores oblong, slightly curved, 16–20 × 5–7 μ.

On dead branches, &c. New Zealand. Australia, Tasmania, Europe, United States.

Very soft and gelatinous, form irregularly lobed, soon deliquescing.

62. Tremella, Dill.

Gelatinous, tremelloid, immarginate, generally smooth (not papillose nor rugulose), variously lobed and contorted, often bright-coloured.

Tremella, Dill, Hist. Musc., p. 41.

Forming foliaceous variously contorted gelatinous masses

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oozing out of dead wood, branches, &c.; spores subglobose, continuous, on germination the germ-tube bears numerous broadly elliptical sporidiola. In some species dense racemes of conidia are produced in the substance of the gelatinous sporophore previous to or contemporaneous with the formation of basidiospores.

Tremella lutescens, Pers., Syn., p. 622; Sacc., Syll. vi, no. 8377; Cooke, Austr. Fung., p. 208.

Very soft and gelatinous, lobes crowded, entire, wavy, pallid then yellowish, 2–8 cm. diameter; spores subglobose, 12–16 μ. diameter; conidia globose, 1.5–2 μ diameter.

On fallen and rotten trunks and branches. New Zealand. Victoria, New South Wales, Queensland, South Australia, Tasmania, South Africa, Europe, North America, Brazil.

The conidia are produced at the tips of densely corymbose branches buried in the gelatinous substance of the fungus.

63. Nœmatelia, Fries.

Subgelatinous, convex, pallid, with a firm central white nucleus.

Nœmatella, Fries, Syst. Myc., ii, p. 227.

Readily distinguished by the presence of a firm, solid, white central nucleus, around which the subgelatinous sporiferous portion is spread, forming a convex body when extended. In some species the external gelatinous layer contracts and disappears when dry, the white nucleus alone being visible. The original structure, however, returns when the specimen is immersed in water.

Næmatelia nucleata, Fries, Syst. Myc., ii, p. 227; Sacc., Syll. vi, no, 8450.

Sessile, depressed, gelatinous, more or less contorted, almost translucent, with a central, white, opaque, hard mass, 0.5–1 cm. across; spores broadly elliptical, hyaline, 7 μ long.

On damp rotten wood. New Zealand. Britain, southern United States.

Quite glairy and soft when moist; when dry the outermost gelatinous portion contracts, the white nucleus alone being visible.

64. Dacryomyces, Nees.

Gelatinous, homogeneous, more or less contorted, often bright-coloured; spores simple or septate.

Dacryomyces, Nees, Syst., p. 89.

Minute gelatinous fungi occurring on dead wood, often yellow or orange; spores cylindric-oblong, curved, often one or more septate at maturity or during germination, sometimes even

– 44 –

becoming muriform. Chains of conidia resembling the basidiospores in form often occur in immense numbers in the substance of the fungus.

Dacryomyces deliquescens, Duby, Bot. Gall., p. 729; Sacc., Syll. vi, no. 8472; Cooke, Austr. Fung., p. 209.

Gelatinous, roundish or irregular and variously gyrose, yellow, almost translucent and subdeliquescent, basal portion root-like, emerging from the matrix, patches 1–4 mm. broad; spores hyaline, cylindrical, obtuse, slightly curved, 3-septate, 15–17 × 6–7 μ.

On decaying wood. Northern Island, New Zealand. Tasmania, Europe, Siberia, United States.

Forming little gelatinous pale-yellow pustules on dead wood, often very abundant in rainy weather.

65. Guepinia, Fries.

Cartilaginous or subgelatinous, erect, substipitate, spathulate or expanded, one surface fertile, the other sterile and minutely velvety.

Guepinia, Fries, Elench., ii, p. 30.

Small, thin, and flexible; distinguished from others with similar basidia by the differentiation into a sterile and fertile surface respectively; spores curved. Chains of conidia sometimes produced on the sterile side of the pileus.

Guepinia spathularia, Fries, Epicr., ii, p. 32; Fl. N.Z., ii, p. 185; Hdbk. N.Z. Flora, p. 614; Cooke, Austr. Fung., p. 210; Sacc., Syll. vi, no. 8520.

Cæspitose; pileus erect, spathulate or irregularly flattened, thin, soft and tough, sterile side pale, pubescent, as is also the short rooting stem; hymenium orange, wrinkled; up to 5 cm. high, usually smaller; spores elliptical, becoming septate, 10 × 6 μ.

On dead wood. Northern Island, New Zealand. Victoria, Queensland, New South Wales, Ceylon, Java, India, United States, Cuba, Brazil.

Very tough when growing, often springing out of cracks in wood, extending for some inches and growing more or less into each other, hence becoming very irregular in form. General form more or less battledore or fan shaped, with a stem 1–2 cm. long, especially when springing from cracks in the wood, a very common place of growth, and under such circumstances the plants are often closely crowded into rows following the crack for several inches. Altogether larger and differing from G. pezizœformis in the crowded habit.

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Guepinia pezizœformis, Berk., in Hook. Journ., 1845, p. 60; Hdbk. N.Z. Flora, p. 614; Sacc., Syll. vi, no. 8518; Cooke, Austr. Fung., p. 210, fig. 96.

Bright orange-red, cartilaginous and elastic when moist, pileus obliquely saucer-shaped or almost flat, stem very short, minutely velvety, 3–4 mm. broad; hymenium slightly corrugated or wrinkled; spores elliptical, hyaline.

On dead wood. Bay of Islands, Northern Island, New Zealand. Tasmania, Queensland, Western Australia.

Usually more or less fan-shaped; rigid and contracted when dry. Growing solitary or scattered.

66. Calocera, Fries.

Cartilaginous, viscid, rigid and horny when dry, vertical, simple or branched; branches terete, often forked at the tips.

Calocera, Fries, Syst. Myc., i, p. 485.

Resembling the branched forms of Clavaria in habit and general appearance; differing in the cartilaginous structure and form of the basidia. Growing on wood. The spores are sausage-shaped and curved, becoming septate on germination and producing clusters of elliptical sporidiola.

Calocera viscosa, Fries., Syst. Myc., i, p. 486; Sacc., Syll. vi, no. 8147.

Irregularly branched, all branches of about uniform diameter, 2–3 mm., 3–7 cm. long, deep-orange, viscid, smooth and polished, very tough when growing, rigid and horny when dry, rooting base 3–5 cm. long, tough; spores cylindric-oblong, apiculate, slightly curved, 9–10 × 4–5 μ, hyaline.

On decaying stumps of pines, &c. New Zealand. Malacca, Europe, United States.

Superficially resembling a branched Clavaria, but differing in the tough consistency and different basidia.

Calocera stricta, Fries, Epicr., p. 581; Sacc., Syll. vi, no. 8163; Austr. Fung., p. 204

Simple, solitary or gregarious but not crowded and confluent, linear, erect, apex subacute, 1–2 cm. high, 2 mm. thick, orange-yellow, tough when moist, rigid and remaining even when dry, orange or yellow, base with white down; spores elliptical, 7–8 × 5–6 μ.

On dead wood, especially of conifers. New Zealand. Victoria, Ceylon, Europe, United States, Cuba.

Distinguished from the closely allied Calocera striata in being firmer in structure, and hence not shrinking and becoming striate or wrinkled when dry.

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Calocera furcata, Fries, Syst. Myc., i, p. 486 (1821); Sacc., Syll. vi, no. 8150.

Tufted, slender, 1–2 cm. long, often once or twice forked, yellow, viscid, branchlets acute, downy, and rooting at the base; spores hyaline, elliptic-oblong, slightly curved, 8–10 × 4–5 μ.

On rotten branches, &c. New Zealand. Europe.

Subgelatinous, slender, not becoming horny when dry, but adhering to the drying-paper. Stem simple slightly pubescent, rooting, sometimes solitary.

Calocera cornea, Fries, Syst. Myc., i, p. 486; Sacc., Syll. vi, no. 8158; Austr. Fung., p. 204.

Clubs tufted, rooting, club-shaped, smooth, viscid, subulate, simple or rarely with a short branchlet, orange-yellow or sometimes pale-yellow, about 1 cm. high, 1–2 mm. thick; spores cylindric-oblong, 7–8 × 5 μ.

On naked wood. New Zealand. Australia, Europe, South America, United States.

Very rigid when dry. Frequently grows in rows, springing from cracks in dry hard wood.

67. Auricularia, Bull.

Effused or the upper portion reflexed; hymenium inferior, with raised ribs or folds, inflated and gelatinous when moist, collapsing when dry; sterile surface velvety or strigose.

Auricularia, Bull., Champ. Fr., p. 277.

Resembling species of Stereum in habit and appearance; differing in consistency and in structure of basidia. Spores oblong, curved, producing on germination a branched promycelium bearing several strongly curved sporidiola.

Auricularia mesenterica. Fries, Epicr., p. 555; Austr. Fung., p. 205; Sacc., Syll. vi, no. 8294.

Resupinate, upper margin free and reflexed, often very broadly effused, several specimens running into each other, subgelatinous and swollen when moist, rigid when dry, 3–10 cm. across; hymenium wrinkled and folded, smooth, brownish-purple; pileus coarsely velvety, zoned different colours, margin not deeply lobed; spores sausage-shaped, hyaline, smooth, continuous, slightly curved, 18–20 × 7 μ.

On trunks. New Zealand. Australia, Tasmania, Europe, South America, United States.

Often broadly effused and with numerous partly free imbricated pilei. Distinguished at sight from the species of Hirneola by the strigose zoned pileus.

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68. Hirneola, Fries.

Substance thin, cartilagineo-gelatinous, soft and tremelloid when moist, rigid when dry; sporophore cup-shaped or human-ear shaped, fertile surface polished, sterile surface velvety.

Hirneola, Fries, Fung. Natal, p. 24.

Differs from Auricularia, its nearest ally, in not becoming bullately inflated when moist, the substance being no thicker when moist than when dry. Basidia rod-shaped or fusoid, transversely septate, each cell bearing a single sterigma which in turn bears an oblong curved spore.

Hirneola polytricha, Montag., Bel. Voy. Ind. Or., Crypt., p. 154; Austr. Fung., p. 206, fig. 90; Sacc., Syll. vi, no. 8311.

Hemispherical, then expanded, sessile but narrowed to a more or less central or oblique point of attachment, thin and elastic., rigid when dry; hymenium even, dark-brown with a purple tinge; externally even, minutely but densely velvety, greyish, becoming a rich yellowish-brown when dry; size variable, 4–12 cm. across; spores hyaline, colourless, smooth, sausage-shaped, slightly curved, 14–15 × 6–7 μ.

On trunks, branches, &c. New Zealand. Australia, Tasmania, Java, Ceylon, Tahiti, Madagascar, South Africa, South America, Cuba, Mexico, Chatham Islands, Lord Howe Island, Torres Straits.

Differs from Hirneola auricula-judœ in the absence of raised wrinkles on the pileus. A very widely distributed species, and one of the few fungi used on a large scale, and over a widely extended area, for food. Mr. T. Kirk gives the following account in the “Transactions and Proceedings of the New Zealand Institute,” vol. xi, p. 454 (1878). “An Edible Fungus: Hirneola polytricha is collected and sent to China, where it is highly prized for food and medicine. In 1887 220 tons, valued at μ11,318, were collected in New Zealand and exported.”

Hirneola auricula-judœ, Berk., Outl., p. 289, pl. 18, fig. 7; Cooke, Austr. Hdbk., p. 206; Sacc., Syll. vi, no. 8312.

Hemispherical, then expanded and more or less resembling a human ear in shape, sessile, thin, soft and flexible when moist, rigid when dry, 3–8 cm. diameter; hymenium glabrous, uneven with anastomosing ridges and folds, dingy flesh-colour, then blackish or dark-brown; externally showing irregular wrinkles, minutely but densely tomentose, greyish-olive, often brownish when dry; spores smooth, hyaline, continuous, sausage-shaped, slightly bent; 20–25 × 7–9 μ.

On dead branches; in Europe most abundant on Sambucus.

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New Zealand. Australia, Tasmania, Madagascar, South America, Cuba, Mexico, United States.

Distinguished from H. polytricha in having the pileus and hymenium wrinkled and veined. Auricularia mesenterica differs in the shaggy zoned pileus. Edible.

Hirneola hispidula, Berk., Exot. Fung., p. 396; Austr. Fung., p. 206; Sacc., Syll. vi, no. 8323.

Campanulate, then expanded, sessile, oblique, thin and flexible when moist, rigid when dry, 8–12 cm. across; hymenium dark-brown, even, or more or less veined; externally covered with a dense velvety pile, yellowish-brown or with an olive tinge, even or slightly veined; spores sausage-shaped and slightly bent, hyaline, continuous, smooth, 19–24 × 7–8 μ.

On dead wood. New Zealand. Australia, Mauritius, Ceylon, South America, Java, Hong Kong.

A large and fine species, somewhat variable in form and colour, sometimes narrowed at the point of attachment into a stem-like base. Hymenium sometimes with a purple tinge. Often growing in clusters of 2–6. Known more especially by the hairy pileus being almost hirsute, and the hairs longer than in other species.

69. Septobasidium, Pat.

Effused and resupinate, coriaceous, not moist or gelatinous; hymenium separating from the lower stratum; basidia transversely septate, curved, sterigmata borne on the convex side of the basidium; spores hyaline, continuous.

Septobasidium, Patouillard, Journ. de Bot., 1892, p. 63.

Superficially resembling Thelephora, but readily distinguished by the transversely septate basidia.

Septobasidium pedicellatum, Pat., Journ. de Bot., 1892, p. 63: Sacc., Syll. xi, no. 743. Syn., Thelephora pedicellata, Schweinitz, Syn. Carol., no. 108; Sacc., Syll. vi, no. 7188; Cooke, Austr. Fung., p. 180; Hdbk. N.Z. Flora, p. 611.

Resupinate; rather soft and elastic, densely fibrous, thick, basal layer composed of fascicles of hyphÆ, tawny-cinnamon, margin whitish, radiating; hymenium paler, forming a thin separable pellicle which is often cracked irregularly; basidia curved, transversely septate, springing from a broadly pyriform basal cell; spores oblong, hyaline.

On branches of living and dead trees of various species. New Zealand. United States, Cuba, Brazil, Ceylon, India, Victoria, Queensland, Western Australia.

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Forming effused patches often 8–12 cm. long, encircling branches; more or less felty and soft, hymenium often not developed. Very destructive when it attacks fruit-trees.

Explanation of Plates I and II.
Plate I.

  • 1. Tremella lutescens, Fries; natural size.

  • 2. Basidia of same, in different stages of development; X 500.

  • 3. Polyporus arcularius, Fries; natural size.

  • 4. Fomes australis, Fries; a very small specimen; natural size.

  • 5. Auricularia mesenterica, Fries; natural size.

  • 6. Transversely septate basidium of No. 5; each cell of the basidium bears a single sterigma, with a sausage-shaped spore at its apex; X 400.

Plate II.

  • 1. Cyphella densa, Berk.; natural size.

  • 2. Surface view and section of same; slightly X.

  • 3. Spores of same; X 400.

  • 4. Guepinia spathularia, Fries; a group of plants; natural size.

  • 5. Clavaria aborescens, Berk.; natural size.

  • 6. Pistillaria ovata, Fries; natural size.

  • 7. Poria hyalina, Berk.; natural size.

  • 8. Hirneola polytricha, Mont.; natural size.

  • 9. Craterella insignis, Cooke; natural size.

  • 10. Stereum illudens, Berk.; natural size.

  • 11. Section of No. 10; natural size.

Art. II.—Transpacific Longitudes.

[Read before the Wellington Philosophical Society, 5th April, 1906.]

On the 31st December, 1900, articles of contract were made by Her Majesty's Government, Canada, New South Wales, Victoria, New Zealand, and Queensland on the one part, and the Telegraph Construction and Maintenance Company on the other, for the construction and laying of the Pacific cable. The contract called for the completion of the whole cable on or before the 31st December, 1902. The cable was finished two months earlier, and, after undergoing the required test of a month, entered upon its commercial career on the 8th December, 1902. Thus was the project that had been advocated with persistence from some quarters for a quarter of a century an accomplished fact; the missing link of about eight thousand miles across the Pacific between Canada and Australia, in the world's metallic girdle, was now supplied.

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Before laying a cable a survey is always made along the proposed route in order to select the most favourable ground, just as the railway engineer runs lines of levels before the final location of the railway. The cable engineer determines his levels by means of the sounding-line (piano wire), and at the same time obtains samples of the ocean-bed. It may be stated here that the direct route of the Pacific cable between the stations was departed from in order to avoid hills, craters, and hard or undesirable ground for the cable to rest upon.

From the survey the number of miles (nautical) required for the different flections was as follows: From Vancouver Island to Fanning Island, 3,654; from Fanning to Suva, Fiji, 2,181; from Suva to Norfolk Island, 1,019; from Norfolk to Southport, Queensland, 906; Norfolk to Doubtless Bay, New Zealand, 513.

The first section of the cable is about a thousand miles longer than any that had been laid before. This necessitated a considerable increase in copper for the conductor and in guttapercha for the dielectric. The working-speed of a submarine-telegraph cable depends on, and is inversely proportional to, the product of the total resistance of the conductor multiplied by the total electro-static capacity of the core, so that, other things being equal, the speed varies inversely as the square of the length of the cable. In the long section there were used 600 lb. of copper and 340 lb. of guttapercha per nautical mile; on the Suva-Fanning section 220 lb. of copper and 180 lb. of guttapercha; and on the remaining three sections the copper and dielectric were in equal proportions of 130 lb. each.

In the neighbourhood of Fiji, at a depth of 2,500 fathoms a temperature of 34.1μ Fahr. was noted, being the lowest temperature taken during the survey. There is very little difference in the temperature of the ocean at great depths, say below 3,000 fathoms, over a great extent of the earth's surface, the temperature being only a few degrees above the freezing-point, or 32μ Fahr.

The greatest depth, 3,070 fathoms (about three miles and a half), was found on the Fanning-Fiji section, where the bottom specimens consisted principally of radiolarian ooze. This ooze is found at the greatest depths, and was obtained by the “Challenger's” deepest sounding in 4,475 fathoms. The United States steamer “Nero” sounded in 5,269 fathoms (six miles), this being the deepest sounding recorded in the ocean, and the-material brought from the bottom was radiolarian ooze.

Of the 597 samples of sea-bottom obtained on the Pacific-cable survey, 497 were such that they could be divided into distinct types of deposits. It was found that 294 samples referred to globigerina ooze, sixty-five to red clay, forty-three to radio-

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larian ooze, forty-five to coral mud or sand, twenty-seven to pteropod ooze, twelve to blue or green muds, and eleven to organic mud or clay.*

The pressure at a depth of 3,000 fathoms, in which a considerable portion of the Pacific cable is laid, is about 4 tons to the square inch. When the cable is being laid at such depths, it will be approximately twenty miles astern of the ship before it touches bottom.

Deep-sea cables last longer in the tropics than in the northern oceans. The reason is to be found in the fact that in the tropics marine life, from which globigerina ooze is derived, is more abundant than in the more northerly or southerly waters. It is the sun and the warmed surface-water that call into life these countless globigerina, which live for a short space, then die and fall to the bottom like dust, making such a good bed for the cable to rest in. In the arctic currents where the surface is cold the water does not teem with life in the same way as it does in the tropics, and consequently there is less deposit on the bottom of the ocean.

A submarine cable consists first of a core, which comprises the conductor, made of a strand of copper wires, or of a central heavy wire surrounded by copper strips as in the Pacific cable, and the insulating covering, generally made of guttapercha, occasionally of indiarubber, to prevent the escape of electricity. As far as cabling is concerned, this is really all that is necessary—an insulated conductor. This, however, would not, in the first place, be sufficiently heavy to lie in the ocean, and, secondly, would be too easily injured and destroyed by the many vicissitudes to which it would be subjected. For this reason a protection in the form of a sheathing of iron or steel wires surrounds the core, the nature, size, and weight of the sheathing being dependent upon the depth of the water and kind of ground over which it has to be laid. The deep-sea section, being the best-protected from all disturbing influences outside of displacement of the earth's crust by earthquakes or volcanic action, is naturally the one of the smallest dimensions; and for the shore end, which is exposed to the action of the waves; to driftwood, to the grinding of ice in the more northerly latitudes, and to the danger of anchorage, especially of fishing-boats, the sheathing must be very heavy. So that, while the deep-sea cable is somewhat leas than 1 in. in diameter, that for the shore end is nearly 2 ½ in. in diameter. The action of the waves is limited to a depth of only about 13 fathoms, so that their influence on the cable, manifested by wear and chafing, is confined to the shore end.

[Footnote] * Report of Sir John Murray.

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The Pacific cable is equipped with the most modern apparatus at the various stations, and the cable is worked duplex—that is, messages are sent and received on the same cable at the same time.

Canada had carried longitude work from Greenwich across the Atlantic and thence to Vancouver. The completion of the British Pacific cable offered an opportunity for continuing the work across the Pacific in the interests of navigation and geography, besides tying for the first time longitudes brought eastward from Greenwich with those brought westward, making the first longitude girdle round the world.

In October, 1902, the Hon. Clifford Sifton, then Minister of the Interior, authorised the carrying-out of the transpacific longitudes, and the Governors of the South Seas, Australia, and New Zealand were respectively officially notified thereof. In preparing the programme for carrying out the work the climatic conditions of the various stations to be occupied were studied so that the most favourable times and seasons might be chosen. It was found that Suva, Fiji, was the governing factor, as it was by far the rainiest place of the series. The work was placed in my charge, and Mr. F. W. O. Werry, B.A., was associated with me as the other observer.

The instrumental outfit of the two observers was practically the same. Each observer was provided with a Cooke and Son astronomic portable transit, each of 3 in. clear aperture, the one of 34 in. the other of 36 in. focal length. Each transit was provided with reversing-apparatus. The transits of stars were observed over eleven threads in groups of three, five, and three respectively. The eye-piece attachment carried a micrometer (one revolution about a minute of arc with thread parallel to the transit threads) for latitude work; and the whole attachment was necessarily movable through 90μ, so that the movable or micrometer thread becomes horizontal. The recording of transits was made, by means of a key, on a Fauth barrel chronograph. Each observer was provided with two sidereal box chronometers, one being a spare instrument in case of accident. There were, besides, dry cells, switchboards, and minor accessories to complete the outfit. I carried, too, a half-seconds pendulum apparatus and a Tesdorpf magnetic instrument, the latter similar to the ones furnished Drygalski, of the “Gauss,” on his Antarctic expedition.

At each station—that is, at Fanning, Suva, Norfolk, South-port, and Doubtless Bay—a brick or cement pier was built, and an observing-hut covering the same. At Vancouver, which is used as a longitude reference point for the whole of British Columbia, we have a permanent transit-house.

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Bamfield, on the west shore of Vancouver Island, is the eastern end of the Pacific cable, and was not occupied as an astronomic station, but simply as an exchange station—that is, for the comparison of the Fanning and Vancouver chronometers, to be described more fully later.

Longitude work consists in simply determining the accurate sidereal time for each of two places, the longitude of one of them being known, at an absolute instant, and then comparing such times: the difference between them will be the difference in longitude. The operation may be briefly stated: Each observer determines the error of his sidereal chronometer at a particular instant; then by means of the telegraph line or cable the two chronometers are compared, to be explained later; this comparison may be likened to an instantaneous photograph of both chronometers. Applying the respective chronometer corrections for the instant of comparison to the times thus shown by the two chronometers, we obtain the absolute local sidereal time for each place for the same instant; and, as before, the difference between these times is the difference of longitude.

Now, suppose we have a transit instrument with a single vertical thread, and that thread situate in the axis of collimation; furthermore, the axis of the telescope horizontal, no inequality nor ellipticity of pivots, and the pointing of the telescope truly in the meridian; then, if we record the transit of a star across the thread, and the time noted is free from personal equation, we obtain immediately the clock-corrections by comparing the observed time with the right ascension of the star for that time and day. The many conditions imposed in the last sentence show the many sources of error, the effect of which must be evaluated ere we obtain the desired quantity—the clock-correction; in other words, the true local sidereal time at a given instant.

We must therefore devise means for determining the instrumental errors, some of which are practically constant—inequality and ellipticity of pivots; while the others—level, azimuth, and collimation—are more or less variable from day to day. Careful readings, at the beginning and end of a season, of the former will evaluate them. For the latter we will speak of the level-corrections first. This quantity is determined directly by means of the striding-level placed upon the axis of the instrument. Readings should be taken as frequently as the intervals between stars admit. With sensitive levels, reading about a second of arc for divisions, great care must be exercised in allowing the level to come to rest. My own practice is not to take a reading until fully a minute has elapsed after placing the level, and as a light is necessary for reading at night, the reading should be taken quickly, for even a short exposure of the level to light

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will cause a change in the reading. I consider a six-minute interval between stars the minimum during which a deliberate reading (including reversal of level) for inclinations of the axis can be made. How to treat the various level-readings for one position of the instrument will depend upon circumstances. The readings may show a decided and unquestionable gradual change of level; in such a case the readings may be plotted and the level-reading for each star interpolated therefrom. If, on the other hand, the level - readings are confined within the errors of reading and small fluctuations, we may then take the mean of the various readings as the reading for that position of the instrument. The angular value of the level - reading expresses the angle between the vertical plane (in the case under consideration the meridian) and that described by the transit; the two great circles intersect each other in the horizon, where the level-correction is nil. The level factor, usually designated by B, is expressed by cos (φ—δ) sec δ. This factor computed for each star, multiplied by the inclination of the axis, expressed in time, gives then the level-correction to be applied to the respective transits. Errors of level are measured directly, while those of azimuth and collimation with portable astronomic instruments are not directly measured, as is the case with the large transits in observatories. This leaves then the determination of three unknowns—the azimuth, collimation, and clock corrections; the minimum number of stars to determine which is three. With only three stars, however, there would he no measure of the accuracy of the observations, for one, and only one, value for each of the unknowns would satisfy the three observation equations; there would be no probable error. If the instrument is not in the meridian it is evident that the times of transit of stars north of the zenith will suffer a correction of opposite sign from those to the south. If the telescope is pointing west of north, north stars transit too late, and south stars too soon; and vice versa if pointing east of north. As polar stars move slowly they are well adapted for obtaining the azimuth-correction, and hence one polar star is included in each time set for each position of the instrument, and the general azimuth-factor is sin (φ—δ) sec μ.

With the collimation-error, however, the correction for north and south stars is of the same sign for one position of the instrument; but when the instrument is reversed, then the error is of opposite sign, and the transits of stars are similarly affected. The effect of the collimation - error becomes therefore more apparent and is more accurately deduced when some stars are observed in one position of the transit, and others with the telescope or axis reversed.

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The effect of the collimation-error on the times of transit varies directly as the secant of the declination of the star, hence the collimation factor is sec μ.

In order, therefore, to obtain a satisfactory time-determination—which is really the quantity sought—we observe more than the absolutely necessary three stars, and find the most probable value by the method of least squares.

In the programme of the transpacific longitudes it was arranged that (barring cloudy nights) on each night there should be two independent time - determinations; each determination to be derived from fourteen stars, divided into two groups of seven each, of which one was a polar. Furthermore, one group was observed clamp east, and the other clamp west. The six other stars of each group were “time” stars, and selected near the zenith and south (in the Northern Hemisphere) thereof. Instead of three we now have fourteen observation equations from which to deduce the three unknowns, already mentioned, by the usual method of forming the three normal equations. It is desirable to reduce the effect of azimuth and collimation on the derived clock-correction; we attain this by making the algebraic sum of the azimuth-factor as small as possible, and similarly with the algebraic sum of the collimation-factors.

In deducing the time - correction it evidently must signify the correction at some particular epoch, for every clock and chronometer has a rate. The epoch chosen is generally the mean of the various transits constituting a set, and the transit of each star is corrected for rate, as if all stars had been observed at that mean time. If, after having obtained the azimuth and collimation errors, we apply them with their respective factors to each transit and compare this corrected transit with the apparent right ascension corrected for aberration, we obtain the clock-correction of that transit or star, and the difference between this and the clock-correction of the normal equation gives us a residual. Each star thus furnishes a residual, and from them is found the probable error of a single observation as well as of the deduced clock-correction from all the stars. The average probable error of the latter is about 0.01 s. for good work.

A word about rate. Rate is one of the most difficult problems with which we have to deal in field longitude work. It is not the magnitude of the rate, although a small rate is very desirable, but the constancy: this is the crux. A chronometer may have an apparently constant daily rate, yet the hourly rate for the twenty-four hours may and does vary. Again, the rate is not the same when the current is on as when it is off; the former obtaining when observing and the latter the rest of

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the day. The rate deduced from two independent time-determinations of the same night, when the temperature is practically constant during the time of observation and the clock is in circuit with the battery (one cell) only during that time, is seldom, if ever, the same as that obtained from day-to-day observations.

In our programme we have two independent time-determinations for each night. Each set of transits is reduced to the epoch of the mean of the times of transit of the stars comprising that set. The rate which is applied for each transit to the mean epoch, and for which some magnitude must be assumed, is practically a vanishing quantity in the resulting clock-correction. The ideal time of exchange would be at that epoch when the effect of rate is eliminated. But, for various reasons, this is found to be impracticable. In the programme, then, of two independent time-determinations, for obvious reasons the exchange was arranged to take place about midway between the two epochs.

An interpolation between the two epochs gives the clock-correction at the instant required—that of the signals. This assumes that the rate is constant during the interval and is represented by a straight line. If extrapolation is necessary, as sometimes occurs, the rate-value has less weight. It is highly desirable that the temperature of the chronometer be kept as uniform as possible, and, if necessary, special provision made to attain this end.

We are supposed now to have made a complete time-determination, and are ready for exchange of signals—that is, of a comparison between the two clocks of the two stations.

As some of the exchanges were over land lines, I shall explain this method of exchange first, taking the case of Vancouver and Bamfield. Each of these stations was supplied with a switchboard. The portable switchboard has been in use many years and has given every satisfaction. On it are mounted a talking relay, a signal relay, and a pony or clock relay; the last is never on any circuit but that of the chronometer with one dry cell. Besides, there is an ordinary talking-key and a signal-key, the latter breaking circuit when depressed while the ordinary telegraph-key makes circuit. Along one edge of the board there is a row of binding-posts for connecting with the clock, chronograph, main line, and batteries, of which there are three dry cells for the chronograph, and, as stated, one for the chronometer. And, lastly, there is a three-point switch, by means of which the main line can be thrown on or off the points of the clock relay, and plugs to cut in or off any relay. While observing, the chronograph-circuit passes over the points of the clock

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relay, and, as the clock or chronometer breaks circuit every two seconds (omitting the 58th second so as to identify the minute), the points of the clock relay separate every two seconds, and hence record the clock-beats on the chronograph. In the chronograph-circuit is the break-circuit observing-key too, by means of which the transit of each star over the eleven threads is recorded.

It is customary when beginning the exchange to put the telegraph-line for a minute at each station over the points of the clock relay, whereby the circuit of the main line is broken by each chronometer every two seconds—that is, we let the clocks (chronometers) record simultaneously over the line, each chronograph thus obtaining the record of both clocks. From this record we immediately see the relative position of the respective minutes—in fact, of the seconds too—enabling one readily to identify corresponding arbitrary signals, by means of which the more accurate chronometer-comparison is made. Theoretically, the comparison by the chronometers recording directly over the line, as above, is as good as by arbitrary signals. The trouble lies in scaling or measuring the former. As, for an interval of a minute, the relative position of the two-second breaks of the two chronometers is the same, after having measured one such interval on the chronograph sheet the mind is involuntarily biassed; we know that all the others should be the same, and, consequently, we cannot measure, say, thirty, our minimum number, with that freedom of mind which would be the case if we did not know what measure to expect: hence the device of the arbitrary signals. In this case each chronometer records only on its own chronograph. One observer now sends by means of the signal (break-circuit) key twenty arbitrary signals; the chronograph - circuit, which always passes over the points of the clock relay, is now made to pass too over the points of the signal relay, which is on the main-line circuit. Hence a signal sent will be recorded on each chronograph, and each chronograph has its own chronometer-record for interpreting any signal, just as it interprets the transits while observing.

As the word implies, these arbitrary signals are intentionally made irregular, and will average about two seconds apart. The other observer now sends, similarly, forty signals, and again the former twenty more, so that the mean of the times of sending of the two observers about coincides, thereby eliminating differential rate of the two chronometers. It is customary when sending signals to give a rattle with the key at the beginning and end of each set. If there is no trouble on the line the whole exchange is over in five minutes. A few minutes are required for conversation about the condition of the sky. If the prospects

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are hopeless for the night for one, the other desists from further observations. The accuracy with which these comparisons are made is far beyond the accuracy that is possible in a time-determination: while the probable error of the latter is, say, 0.01 s., that of the former is generally less than 0.002 s.

The exchange on the cable is similar to that just described of arbitrary signals. The chronograph here is replaced by the paper fillet of the cable service. It is scarcely necessary to observe that nowadays signals (messages) on the cable are not read by means of deflections of a small mirror, interpreted on an opal glass scale by means of a reflected beam of light, but are read from the fillet of paper on which a siphon records in ink the deflections. As the current is very weak the siphon is not in direct contact with the paper, but, by an ingenious vibrating device, it deposits a tiny drop of ink at very brief intervals. A cable message looks like a profile of the Rocky Mountains, the ups and downs having an interpretation like the dots and dashes in the Morse system of telegraphy. From experience it is found impracticable to have the clock recording directly on the cable for interpreting signals sent or received. However, it is necessary to have a time-measuring scale on the fillet. We accomplish this by attaching another siphon to the frame of the cable instrument. This one is quite independent of the cable. It is actuated by a long vertical rod attached to the horizontal arm of an ordinary sounder, and connected to the siphon by a silk fibre. This latter siphon drags an ink-line on the fillet. The sounder is put in circuit with the clock, and hence every time the clock or chronometer breaks circuit the sounder makes a sharp break in the line on the fillet, and a time-scale is obtained close to and parallel to the zero-line of the cable-siphon. By projecting vertically these recorded clock-breaks on to the cable-siphon record, we can interpret in time the arrival or departure of a signal. We must know, however, the relative position of the two siphons. The signals are sent with one of the two cable-keys (on cables there are always two keys, one for sending positive and the other for sending negative currents). To the lever of the cable is adjusted another lever which is in the clock-circuit. It is so adjusted that the moment the cable-key makes contact—that is, sends a current into the cables—at the same moment the clock-circuit is broken, thereby both siphons record the event simultaneously, and the parallax between the two siphons is obtained. As a check on the value thus obtained for the parallax, a slight tap is given to the frame carrying both siphons, thereby disturbing both, and the parallax obtained. By the above arrangement, when sending signals we have two records on the fillet, one by the clock-siphon, the other by the

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cable-siphon. In receiving signals there is, of course, only the record of the cable-siphon, the other siphon recording only the chronometer-beats, which, on the fillet, measure about 1 in. for the two seconds. The speed of the fillet may be varied to any degree. It will be seen that a comparison of clocks by this means is simply a matter of careful linear measurement. Were the records at the two stations instantaneous, then the two records would be identical; but such is not the case. Each signal arrives late at the distant station, and therefore the two records will differ by twice the time of transmission, assuming that the time of transmission is the same in each direction, an assumption which we cannot avoid. On the long section of the cable between Bamfield and Fanning, about four thousand two hundred statute miles, the time of transmission was a third of a second, equivalent to about twelve thousand statute miles per second.

In the first longitude-work by cable before the introduction of the recording-siphon, instead of arbitrary signals, the clock-beats were sent by hand at intervals generally of ten seconds, and the time of arrival of the signal, as indicated by the reflecting-galvanometer, was noted by the “eye and ear” method. The uncertainties and “personal equation” in this method of exchange and comparison of clocks are apparent.

We have now explained briefly how the clock-correction is obtained for a given instant, and how the comparison of the two clocks is made. The application of the clock-corrections respectively to the times of exchange gives apparently the local sidereal time for each place at the same instant. Each value is, however, affected by a small correction—the personal equation of each observer. As the quantity sought is the difference between the local sidereal times, the absolute personal equation of each observer is unimportant; it is the difference between the two personal equations that affects the difference of longitude. On land lines, where the ready means of transportation is good, it has been customary (up to the present, when, by the introduction of the registering-micrometer, the personal equation is eliminated) for the observers to exchange stations, the mean result of the two differences of longitude being free from personal equation: this is on the assumption that the personal equation of the observers remains constant during the longitude campaign. On this assumption, if there is a series of stations odd in number, and the observers occupy alternate stations, it will be seen that the odd-numbered stations will be free from personal equation, and the even-numbered ones affected by it. Now, between British Columbia and Australia, and also between British Columbia and New Zealand, the number of stations is odd—i.e., there are three intermediate stations, Fanning, Suva, and

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Norfolk; hence Southport (Queensland), Doubtless Bay (New Zealand) and Suva (Fiji) are free from personal equation.

Personal - equation observations were, however, made at Ottawa by the two observers using the same clock and determining its correction at the same time on the same stars with the two transit instruments, and the resulting difference of personal equation, 0.124 s., applied to Fanning and Norfolk.

Southport was connected with the observatories at Sydney and at Brisbane, and similarly Doubtless Bay with the observatory at Wellington. Personal-equation observations were made between the respective observers.

It was on the 29th September, 1903, that the first satisfactory clock exchange was had with Sydney, and so this night may be considered as the one when for the first time longitude from the west clasped hands with longitude from the east, and the first astronomic girdle of the world was completed. The immediate reasons for the first telegraphic connection in longitude between Australia and the prime meridian, Greenwich, were (1) with a view of confirming the position of the eastern boundary of the Colony (now State) of South Australia, 141° E.; (2) for obtaining the longitude of stations to be occupied for observing the transit of Venus in 1882. To attain this end connection was made astronomically between Sydney, Melbourne, Adelaide, Port Darwin, and Singapore. A connection was made, too, between Sydney and Wellington. All Australian and New Zealand longitudes at present rest on the position of Singapore as accepted in 1883, which then, quoting from the Government report for 1886 of South Australia, “had twice been telegraphically determined—first in 1871 by Dr. Oudeman, of Batavia, and Mr. Pogson, of Madras, and more recently by Commander Green, United States Hydrographic Department.” The determinations of the latter were accepted. It may be remarked that at this time the Thomson (Lord Kelvin) recording-siphon had not yet been introduced, and that the clock exchanges between Port Darwin and Singapore over the cable were made by use of the deflecting mirror or reflecting galvanometer, already spoken of, a method involving more or less uncertainty in noting by “eye and ear” the movement of the mirror and the instant of time of its occurrence.

Singapore was dependent in position upon Madras, the initial meridian for the great trigonometrical survey of India.

For over a century observations have been taken from time to time to determine the longitude of Madras. The early ones, before the advent of cables and telegraphs, were dependent mostly on lunar observations, some on Jupiter's satellites. In 1891 the Survey of India had not adopted the then best value, so that at the International Geographic Congress held at Berne

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in that year the question arose, why the known error in longitude of 2′ 30″ was not corrected on the Indian maps and charts. This gave rise to a discussion in India, and the whole longitude work was reviewed, with the result that a determination de novo was decided upon, carrying the work directly from Greenwich via Potsdam, Teheran, Bushire, and Karachi, where connection was made with the three arcs of the great trigonometrical survey between Karachi and Madras. This work was carried out by Captain (now Major) S. G. Burrard, R.E., and Lieutenant Lenox Conyngham, R.E., in 1894-6. The resulting longitude of Madras was 5 h. 20 m. 59.137 s. ± 0.022 s.

In 1903 a redetermination of Greenwich-Potsdam was carried out by Professor Dr. Albrecht and Mr. Wanach. Stations were exchanged and observations made with a Repsold registering-micrometer. The exchange of stations was made to test the elimination of personal equation by means of the registering-micrometer, and the result was highly satisfactory, the weighted mean of the one result agreeing with the weighted mean of the other to the third place of decimal of a second of time. It may be stated here that the introduction of the registering-micrometer in longitude-work marks a distinct epoch in that class of work, not only in assuring greater accuracy in the results, but also in very materially reducing the cost of longitude-work of the first order by saving of time and money in doing away with the necessity of exchange of stations. Since the completion of the transpacific longitude-work, the two Cooke transits used in that campaign have been provided with the registering-micrometer made by Saegmueller, of Washington, and the longitude work of 1905 was carried out with that attachment.

From the 1903 determination by Albrecht we have for the longitude of Potsdam 0 h. 52 m. 16.051 s. ± 0.003 s. This value is 0.098 s. greater than that of Burrard obtained in the series of 1894-6 referred to above.

In the reduction (1885) of the Australian longitudes, the longitude of Madras was accepted as 5 h. 20 m. 59.42 s., and the derived value of Sydney was 10 h. 4 m. 49.54 s.

In making the comparison between the longitude of Sydney as brought from Greenwich eastward with that brought westward, the best and most recent available data are utilised for the longitude of Madras.

Taking, then, Albrecht's value for the arc Greenwich-Potsdam, and the values of Burrard for the arcs Potsdam-Madras, we obtain for the longitude of Madras 5 h. 20 m. 59.235 s. ± 0.021 s.

As there have been no new determinations of the various arcs from Madras to Sydney, the values given in the report of May, 1885, by Ellery, Todd, and Russell, on Australian longitude, are used. Adding the latter to the above-accepted value,

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we obtain for Sydney 10 h. 4m. 49.355 s. ± 0.088 s. The Canadian value is 10 h. 4 m. 49.287 s. ± 0.058 s. Difference, 0.068 a. = 1.02″ = 84 ft. for the latitude of Sydney—that is, the first girdle of the world closed within 84 ft.

New Zealand.

The longitude of Wellington is discussed in the report of 8th August, 1884, of the Surveyor-General of New Zealand, and in Appendix No. 1 of that report Mr. C. W. Adams fully describes and tabulates the result of the determination for the difference of longitude between Sydney (Australia), and Wellington. The time-determinations of this series are of a high order, and deserve every confidence. At the time (1883) the siphon recorder had not been introduced on the cable, so that the clock-beats were sent by hand and the deflections of the reflecting-galvanometer were noted by eye. As Mr. Adams says in the above appendix, “I received them at Wellington by reflecting-galvanometer, but, instead of noting each signal by ‘eye and ear,’ I simply tapped the key and recorded each signal on my chronograph”—that is, as soon as the motion of the light-spot had impressed itself on the brain, the key was tapped to record the event. Although many trials were made for “the loss of time in receiving signals,” and the results are inconsistent amongst themselves, yet the lack of self-recording cable-apparatus is the weak point in the 1883 determination for difference of longitude. The range for difference of longitude for the 1883 determinations is satisfactory, but of course may involve constant errors without affecting the range. As we shall presently see, the inter-agreement between the difference of longitude between Sydney and Wellington obtained in 1883, and that in 1903, when the siphon recorder was used, is remarkably close.

As Mr. T. King, observer at Wellington, has so fully given the evolution of “the longitude of the Colonial Observatory, Wellington,” in the “Transactions of the New Zealand Institute,” vol. xxxv, 1902, pp. 436-47, it is not necessary here to cover the same ground.

As has already been explained, the determination of the longitude of Southport (Australia) and Doubtless Bay (New Zealand) is free from personal equation, and, so far as the Canadian work is concerned, these two places are necessarily better determined than places dependent upon them. In other words, Doubtless Bay is better determined in longitude in the Canadian arcs than is Wellington; for Wellington to be as well determined as Doubtless Bay would mean perfect observations and perfect exchange of time-signals between the two places, which is of course impossible, no matter how good the work.

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Upon my arrival in Wellington in November, 1903, I was very cordially received by Sir James Hector and Mr. T. King. Sir James, by the way, we Canadians claim as a kinsman, for we have not forgotten the very valuable work he did nearly half a century ago in the Rocky Mountains in connection with the Palliser expedition. The Hon. Mr. Richard J. Seddon, Premier, who had been officially notified of my coming, offered every facility for the successful issue of the work, and Sir Joseph Ward, Postmaster-General, kindly placed the telegraph-lines at my disposal. Mr. King and I discussed the work in hand—the connection of Doubtless Bay and Wellington. The star programme, the routine of observing and exchange of time-signals were followed as already explained. Mr. King in his time work always observes by “eye and ear,” and this method he followed too in the longitude work, including the personal-equation observations, while I, as usual, recorded my observations on the chronograph.

The main consideration was the installation of electric apparatus to enable the exchange of time-signals between the two stations. After explaining to Mr. J. K. Logan, Superintendent of Government Telegraphs, what was required, the electrician, Mr. Buckley, and Mr. Chisholm installed the necessary batteries and relays at the observatory, a description of which, furnished me by Mr. Logan, follows later. A brief résumé of the apparatus at the Wellington Observatory may be given.

The observatory was established in 1869, and is used for time service only. It is situate on the summit of the hill within the old cemetery, and overlooks the city, harbour, and surrounding country. The building has two rooms—a clock-room and a transit-room.

Clocks.—In the former are three mean-time clocks, and one sidereal—Dent No. 39720—having electrical attachment making contact or circuit every second except the 60th in order to identify the minute. The clocks are all mounted on brick and cement bases, and are fastened to substantial braced frames.

Transit.—The transit is by Troughton and Simms, and is mounted on a rather high stone pillar. It has an aperture of 2 ¾ in., and a focal length of 32 in. The reticule has seven threads at equal equatorial intervals of about 17 seconds of time. There is a sensitive striding-level, and one oil-lamp for illuminating the field. The single small setting-circle reads to minutes, and the reversing of the telescope is done directly by hand.

Meridian Mark.—The meridian mark, placed thirty-five years ago, which also serves for testing collimation in the daytime, is a 3 in. iron bar set in cement, and shows well above the skyline of the Tinakori Range to the north.

Chronograph.—The chronograph is of the Morse pattern and

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records on a tape. It is provided with two styles, side by side. The one records, embossing by make-circuit, the second-beats of the sidereal clock, while the other similarly records the signals by the transit-key, also the clock or arbitrary signals received (from Doubtless Bay) when making a comparison of the clocks for the determination of the difference of longitude. The transit and arbitrary signals on the tape are readily interpolated and expressed in time from the embossed dots or records indicating the seconds of the local sidereal clock.

Electric Apparatus.—Mr. J. K. Logan, Superintendent of Government Telegraphs, has furnished the following description and diagram of the arrangement especially installed at the Wellington Observatory for the differential longitude work with Doubtless Bay, as this was the first time that an automatic exchange of clock-signals had been made with the observatory. (The Wellington clock made contact (circuit) every second, while the chronometer at Doubtless Bay was arranged to “break” circuit.)

Two British Post Office polarised relays, the coils of each of which were joined in parallel, giving a resistance of 150 ohms. for each relay, were connected in multiple through three Leclanché cells to the terminals of the clock. 120 Leclanché cells, with the copper earthed, were joined to one of the local terminals of one of these relays, and, by adjustment, the tongue of this relay was made to bear against the stop connected to that terminal. The terminal connected with the tongue was then joined to the copper terminal of a Siemens relay of 500 ohms resistance. The line was connected to the Z (zinc) terminal of the Siemens relay through a switch arranged to disconnect it from the time-recording instruments and connect it to the speaking (Morse) instruments when required. The local terminals of the second British Post Office polar relay were connected through eight Leclanché cells to the terminals of the magnet-coils of the back style of the chronograph. The local terminals of the Siemens relay were conducted through eight Leclanché cells to the terminals of the magnet-coils of the front style of the chronograph. At every make of the clock the tongue of the Post Office relay that was connected to the back style of coils made contact and caused the style to emboss, thus registering every clock-beat. The other Post Office relay at every beat of the clock broke contact at its tongue; the line-current was thus broken and a signal recorded at Doubtless Bay. As this line-current passed through the Siemens relay at the observatory, and while passing held the tongue of that relay open against the bias given to it, at every break of the current the tongue, by reason of that bias, moved across and closed the local circuit, thereby recording marks on the front style. When signals were to be received

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from Doubtless Bay, the observatory battery of 120 cells was cut off, battery being applied to the sending end. At every break of the current at Doubtless Bay the Siemens relay tongue moved to close the circuit and the breaks were recorded by the front style, marks being made at the same time by the observatory clock with the other style. Arbitraries were received from Doubtless Bay in the same way. When arbitraries were being sent from the observatory it was arranged, by means of a two-way switch, to cut off the clock from one Post Office relay—i.e., the one the tongue of which was in the main-line circuit. This relay was then worked by the closing of a key, the line current being broken at the tongue of the relay in the same way as when the clock was operating the relay. This break was recorded at Doubtless Bay and also on the front style at Wellington, by the movement of the tongue of the Siemens relay, at the same time the clock was recording on the back style. It is desired to indicate that for received signals the tongue of the Siemens relay had to move to close the circuit, and the front style then to move to mark the tape. The signals of the observatory clock had to cause the Post Office relay tongue to move to close the circuit, and the back style then to move to mark the tape. The record of the outgoing signals either from the clock or by arbitraries was got after the clock or the key had caused the Post Office polar relay tongue to break the circuit, which in turn caused the Siemens relay tongue to move to close the circuit of the front style, and which style had then to move to impress the tape. The line was 704 miles long, Wellington to Doubtless Bay, and was of 11 ½ copper throughout, 200 lb. to the mile.”

No repeaters were used.

As it was impracticable for the observers to exchange stations it was decided to observe for personal equation at Wellington, and this was done.

I took train to New Plymouth, thence by steamer to One-hunga, and Auckland, and thence by the “Clansman”—the connecting-link between the world and Ultima Thule—to Doubtless Bay. Here, close to the cable-station, the pier and observatory were built. The foundation—a cubic yard—of the brick pier (22 in. by 27 in.) was in compact sand, and hence very satisfactory. The telegraph-line was led directly into the observatory and there connected with the switchboard. It may be remarked that another pier was built in another building where gravity observations were made with the Mendenhall half-seconds pendulum; magnetic observations were also taken.

A triangulation has been carried over the North Island by the Survey Department of New Zealand, and by instructions

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of the Surveyor-General, Mr. J. W. A. Marchant, the District Surveyor, Mr. V. J. Blake, made a connection of the triangulation from Station 20 on the west side of the entrance to Mangonui Harbour to the observatory, and by that means obtained the position of the observatory in terms of the New Zealand initial station and meridian. At Doubtless Bay the superintendent of the cable-station, Mr. C. L. Hertslet, rendered invaluable services both in the cable and land-line exchange of signals. His thorough knowledge of the circuits quickly overcame any difficulties or mishaps that arose.

Six independent determinations of difference of longitude between Doubtless Bay and Norfolk were obtained, and the same number between Doubtless Bay and Wellington.

We will now deduce the longitude of Doubtless Bay, giving the various transpacific arcs determined in 1903.

Longitude.
Stations. Difference of Longitude Probable Error. Time. Probable Error. Arc. Probable Error.
H. M. S. S. H. M. S. S. " " "
Vancouver 8 12 28.368 W. ±0.050 123 7 5.520 ±0.75
Vancouver-Fanning 2 25 5.406 ±0.021
Fanning 10 37 33.774 W. ±0.054 159 23 26.610 ±0.81
Fanning-Suva 1 28 43 837 ±0.008
Suva 11 53 42.389 E. ±0.055 178 25 35.835 ±0.82
Suva-Norfolk 0 42 1.243 ±0.011
Norfolk 11 11 41.146 ±0.055 167 55 17.190 ±0.82
Norfolk-Doubtless Bay 0 22 15.000 ±0.021
Doubtless Bay 11 33 56.146 ±0.060 173 29 2.190 ±0.90
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The geographic position of Station 20 was furnished by Mr. Marchant, Surveyor-General.

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

We have then—
Longitude, Station 20 173 31 37.1
Station 20 to Station A (by Mr. Blake) 0 2 24.1
Station A 173 29 13.0
Station A to observatory (by Mr. Blake) 0 0 3.66
Observatory 173 29 9.34

or, 11h. 33 m. 56.623 s. The Canadian value is 11 h. 33 m. 56.146 s. Difference, 0.477 s., or 7.15”, or 595 ft. for the latitude of Doubtless Bay.

It may be remarked that the position of Station 20 is dependent upon the initial station, Mount Cook, at Wellington, through a chain of triangles about seven hundred miles long. From the roughness of the country it was expedient to carry on a network of triangulation for land survey and settlement purposes, and the refinements of a primary triangulation were not aimed at. In the closing for Wellington it will be found that the difference is 0.038 s., or 0.57”, and of the same sign as the above, making thereby the difference between the telegraphic determination, Wellington-Doubtless Bay, and the one obtained by triangulation 0.439 s., equivalent to 549 ft. at the latitude of the latter.

In the following table is given the deduction of each difference of longitude, Doubtless Bay-Wellington. Column 1 gives the date; column 2 the direction in which the arbitrary signals were sent; columns 3 and 4 the respective sidereal times at the two stations of the mean of the times of the signals sent; column 5 is the comparison of the scalings of the same signal—that is, each signal is measured on the two chronographs and expressed in time to the hundredth of a second of the respective clock; there would be at least thirty such signals, and each signal would show the “difference” between the two clocks at that instant, plus or minus the “transmission-time,” according to the direction sent, westward or eastward; the “difference” given in column 5 is the mean of the thirty individual differences. If the two clocks had no rate, or the same rate, then the difference between the comparison of signals sent in the two directions would give twice the transmission-time; when, however, the clocks had different rates, we must introduce the correction

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“relative rate” of column 6 to one of the comparisons (the later one is taken to correspond to the sign of the rate) in order to make it comparable with the other. Column 7 has been explained. The difficulties encountered with rate have already been adverted to. They become very apparent when deducing the transmission-time, especially when that quantity is very small, as in the present instance. When we are dealing with transmission-time of a third of a second, as is the case between Fanning and Bamfield, one or two hundredths of a second variation affects but little the various transmission-times; but it is very different when the transmission-time falls near the limit of certainty of the rate. The relative rate of the 18th December was deduced as the others from the best available data, and the result shows a negative value of a hundredth of a second for transmission-time. This, however, does not affect the difference of longitude. Columns 8 and 9 give the deduced clock or chronometer corrections respectively at the two stations for the same instant, the mean of the times of the two exchanges. With the three data then—the difference between the two clocks at an absolute instant, and the respective clock - corrections for that instant—we obtain column 10, the difference of longitude, each with its respective probable error deduced from the probable error of the respective clock-corrections.

We have then the following values:— H. m. s s.
Dec. 6—Difference of longitude 0 5 9.231 0.027
" 7 " 0 5 9.200 0.018
" 11 " 0 5 9.225 0.032
" 12 " 0 5 9.156 0.021
" 17 " 0 5 9.210 0.032
" 18 " 0 5 9.199 0.020
Weighted mean 0 5 9.198 0.007
Personal equation 0 0 0.257 0.045
Difference of longitude 0 5 8.941 0.045
Doubtless Bay longitude 11 33 56.146 0.060
Wellington longitude 11 39 5.087 0.075
Sidereal Time Chronometer Correction
Date. Direction. Wellington. Doubtless Bay. Difference. Relative Rate. Transmission Time. Wellington. Doubtless Bay. Difference of Longitude.
1903. H. m. H. m. H. m. s. S. S. S. M. s. M. s.
Dec. 6 Wellington to Doubtless Bay 3 20.67 3 15.96 0 4 42.655 +0.347 0.019
Doubtless Bay to Wellington 2 44.47 2 39.74 0 4 43.039
Mean 3 2.57 2 57.85 0 4 42.847 -5.189 -0 31.573 5 9.231
" 7 Wellington to Doubtless Bay 3 27.24 3 22.77 +0 4 28.059 +0.096 0.018
Doubtless Bay to Wellington 3 16.17 3 11.70 0 4 28.191
Mean 3 21.70 3 17.23 0 4 28.125 -3.180 -0 44.255 5 9.200
" 11 Wellington to Doubtless Bay 3 31.66 3 28.2 0 3 38.167 +0.076 0.026
Doubtless Bay to Wellington 3 23.39 3 19.76 0 3 38.296
Mean 3 27.53 3 23.89 0 3 38.232 +1.119 -1 29.874 5 9.225
" 12 Wellington to Doubtless Bay 3 41.72 3 38.32 0 3 24.142
Doubtless Bay Wellington 4 1.58 3 58.18 0 3 24.018 +0.171 0.024
Mean 3 51.65 3 48.25 0 3 24.080 +3.530 -1 41.546 5 9.156
" 17 Wellington to Doubtless Bay 4 5.70 4 3.27 0 2 25.720
Doubtless Bay to Wellington 4 26.16 4 23.73 0 2 25.659 +0.144 0.041
Mean 4 15.93 4 13.50 0 2 25.690 +3.702 -2 39.818 5 9.210
" 18 Wellington to Doubtless Bay 3 52.85 3 50.61 0 2 14.365 +0.083 -0.010
Doubtless Bay to Wellington 3 42.20 3 39.95 0 2 14.428
Mean 3 47.53 3 45.28 0 2 14.397 +4.582 -2 50.220 5 9.199
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Based on the present value of Sydney, 10 h. 4 m. 49.54 s., and the 1883 value, Sydney-Wellington, 1 h. 34 m. 15.77 s., gives the longitude of Wellington (N.Z.I. report, 1902, p. 442) 11 h. 39 m. 5.31 s. This requires the correction of -0.185 s., the same as applied to Sydney for the adopted value of Madras, dependent upon the work of Professor Albrecht and Major Burrard. We have then for the value of Wellington via Madras-Sydney, 11 h. 39 m. 5.125 s.

The Canadian value is 11 h. 39 in. 5.087 s. Difference, 0.038 s., or 0.57″, or 43 ft.

We have at Wellington, then, another closing of the girdle of the world, as we had the first at Sydney.

The weakest link, yet good, in the longitude of Wellington is the personal equation of the two observers; all the other links are very strong. The observations therefor were made with the same (Wellington) instrument, as it was impracticable to build another pier and mount my Cooke transit there

A final word in closing. Gratifying as the above closing-error is, it is questionable whether one would be justified at the present time, with our improved methods, in making a circuit of the globe in longitude, with the number of stations necessary therefor, in expecting à priori a closing-error of less than one-tenth of a second.

The task that Canada set out to perform, to bind Australia, New Zealand, and the Pacific islands to Canada by the “all-red line,” thereby completing the first astronomic girdle of the world, has been successfully accomplished.

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Art. III. — Notes on the Flesh-eating Propensity of the Kea
(Nestor notabilis).

[Read before the Otago Institute, 12th June, 1906.]

Plate IV.

At a meeting of the Philosophical Institute of Wellington, last session, statements were made to the effect that the mutton-eating habit attributed to the kea (Nestor notabilis) was a myth: at any rate, this was the inference drawn from the reports of the meeting published in the New Zealand Press; and a paragraph founded on this report appeared in Nature for the 28th December, 1905.*

Since this alleged change of habit is of very great interest to biologists, and has received world-wide currency from the account given by Wallace in his “Darwinism” (p. 75), from which it has been copied into many books discussing evolution, it was very startling to be informed that the flesh-eating habit was non-existent in fact, and had only existed in the imagination of certain sheepowners and their shepherds. Biologists have for years been using this alleged change of habit as an illustration of the fact that variation in habit, as well as in structure, occurs in nature; and to be told now that the change of habit is quite mythical is extremely disconcerting.

But what evidence did those who deny the existence of the habit bring forward? And on what evidence is the allegation of the habit founded?

Although I am unable to answer the first question, this is of little consequence, since I am able to give the evidence of some of the many witnesses to the existence of the habit.

As a matter of fact my attention had been called to the subject in conversation with an Australian colleague during the meeting of the A.A.A.S. in Dunedin in 1904; and as a result of that conversation I proceeded to make inquiries of various people in the South Island who were reputed to have had experience of the attacks of the kea on their sheep. And although I made these inquiries to satisfy my own doubts, yet, in view of the importance of ascertaining and establishing the truth (or otherwise) of the matter, I have come to the conclusion to put on record the letters containing the personal

[Footnote] * Immediately on reading this I forwarded a brief note to Nature (12) summarising the facts of the present communication.

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experience of a few persons living in Otago at the time the attacks were first noticed.

These letters were in answer to a series of questions which I put to my correspondents, and these answers entirely support the generally accepted opinion that in certain parts of the South Island certain keas have acquired the habit of attacking sheep and of devouring their flesh. These letters, in short, merely confirm the statements contained in Buller's “Birds of New Zealand,” which were founded on the statement of men living in the same localities as some of these same folk to whom I have written. A good account of the matter has already been given by Mr. Huddlestone in 1891 (6).

The history of the matter is, briefly, as follows:—

In or about the year 1867 it was observed that on certain sheep-runs in Otago, in the neighbourhood of Lake Wanaka, sheep were wounded in a rather mysterious manner. It was noticed in the case of sheep killed for food that a healed wound occurred sometimes in the loin or sides; when shearing, too, similar healed and even open wounds were found in or about the region of the loins; also, when mustering, sheep were seen with more or less pronounced wounds, raw and bleeding, and even with entrails hanging out of large holes in the side of the abdomen.

It was on Mr. Henry Campbell's station at Lake Wanaka that the first efforts seem to have been made to trace the origin of these injuries, but similar facts had been noticed on other stations. Mr. Campbell gave instructions to his shepherds to keep a good look-out for the animal that caused the wounds:* so poisoned mutton-fat was laid out in suitable and likely places, and men were set to watch. It was found that keas were attracted and devoured the mutton more or less greedily, and were poisoned thereby; and the suspicion that they were the enemy was soon turned to certainty by the observation of Mr. James MacDonald, at that time (1868) head shepherd, and now a sheep-farmer at Dipton in Southland. He saw a kea

[Footnote] * It was by some supposed to be a disease. It has even been suggested that the damage attributed to the kea has, in part at least, been caused by gulls (Larus dominicanus). These birds are known to peck at the eyes of lambs, but I am not aware whether they have ever been detected eating sheep or attacking them while alive.

[Footnote] † Mr. John Campbell writes that he believes he was the first to detect the true cause—he states that this was in 1870; but as MacDonald had already reported the occurrence in 1868, there seems either a confusion of dates or Mr. Campbell's memory has played him false as to his being the discoverer, for he states, “The kea was not suspected of attacking sheep in any way in 1868, and not for two years afterwards.” But MacDonald's name is referred to by two or three of my correspondents as being the first to discover the cause of the wounds.

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at work on the back of a sheep. I will quote from his letter to me:—

“I do not know whether I was the first to see the kea attack sheep, but I was the first to report it to Mr. Henry Campbell of Wanaka Station.

“In 1868 my orders were to go all over the run after the snowfall and see that the sheep were evenly [distributed] over the ground, that no hill or spur had more sheep on it than it could well carry. While I was at this work, the snow [being] about 2 ft. deep, I went out to the tops: in a small basin under the top, on the west side, facing a rocky country that we called ‘skay,’ there was a mob of sheep snowed in and unable to get out. There I saw the kea at work. He would come down from the rocks, settle on a sheep's loin, and peck into the sheep, which would run through the mob; but [the bird] stuck to the sheep all the time till he got a piece out of the sheep, then he would fly to the rocks. I watched the bird at this work and did not disturb him till I was fully satisfied. … Then I went down to the station and reported to Mr. Campbell. He could not credit me, and all hands on the station [refused to] believe that the birds would do it; so I was ordered to go to another hill, called the Black Hill, and Mr. Campbell came with me, and some more men, and at the first mob we came to Mr. Campbell and the rest saw [the keas] at work with their own eyes.”

The announcement, first published in the Dunstan Times, was received with incredulity and ridicule by Mr. H. Campbell's fellow-sheepowners. But not for long; soon other sufferers noticed the kea at work, and those who had laughed laughed no more.

So serious was the trouble that on Mr. H. Campbell's station men were engaged as “kea-shooters,” one of whom—Mr. John King, of Pembroke—has been good enough to give me a good deal of information, and to send me the names of several of the gentlemen whose letters are printed below; and on their replies to my questions the present communication is founded.

There can be no doubt, it seems to me, that sheepowners have suffered very considerable loss from the attacks of keas; but whether the numbers of sheep killed by them as given by various persons are the correct ones or are exaggerated I have no means of ascertaining. Sir Walter Buller (13) gives instances, so that I need not repeat them. But it will be noticed that Mr. Dougald Bell believes that the kea kills 5 per cent. of the sheep in the region of Lake Hawea, and records the loss of twenty-five during last season. But, seeing that several people attribute their heavy financial losses to the kea, it seems improbable

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that the exaggeration would be very great. Surely we may credit these men, directly concerned as they were, with sufficient intelligence to investigate the facts with enough care to convince themselves that the losses were due to keas.

It has been suggested that it was in the interests of shepherds and so forth to exaggerate the loss, so that the bonus on kea-beaks might be kept high. I should imagine that ‘cute men of business, and Scotchmen to boot, would scarcely be so befooled by their men.

The Kea seen at Work.

I have been asked by incredulous folk, “Has any one ever seen a kea at work on the sheep?” I was at one time unable to answer this question, till I investigated the subject, but thanks to my correspondents I am now in a position to say, Most certainly; on plenty of occasions have the birds been seen on the backs of sheep, and shot while at work pecking away at the wool and flesh. (See letters at end of this article.)

Several people—viz., Reischek(5), Fraser, Bell—have examined the crop of the bird and found wool and mutton therein.

I will quote from some of my correspondents' letters in answer to my questions:—

J. Campbell writes, “I was coming down the Matatapu when I saw a mob of sheep rushing about as if a dog was disturbing them. When I got nearer I saw a flock of birds hovering over the sheep. These were keas. I stopped and watched for a few minutes, and presently I saw a sheep singled out from the others and make towards some rocks with a kea planted on its back. By running under rocks and rubbing against them the sheep got the bird off its back; but the same [bird] or another was soon on again. This went on for some time till the sheep became exhausted.”

A. Fraser writes, “I have seen the kea attacking the sheep and also eating into a sheep when the latter was stuck in deep snow. I have opened scores of keas' crops and found wool and meat therein.”

J. H. King says, “I have often seen the birds at work on a sheep, and have shot them while on the sheep's back. I have seen a flock of twenty or thirty birds attack a mob of sheep in high precipitous country. The kea would harass them until one bird would suddenly alight on a sheep's back holding on to the wool of the rump. The sheep so attacked, would separate from the mob and rush frantically about: it would either go over a bluff or drop down from exhaustion.

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The kea, which had still held on, was joined by several others, and they soon destroyed [? devoured]* the sheep.”

Again, Mr. Cameron writes, “When 'snow-raking'—that is, taking sheep from high country—keas would gather round the sheep in great numbers, attack one or more quite close to us shepherds. A sheep would get frightened, run out of the flock with one or half a dozen [!] keas on its back, kill or wound it severely before they would let go.”

The majority of my correspondents give a similar account of the facts, and several of them—Messrs. Bell, Cameron, King, McGregor, McKenzie—state that they have often seen keas on the sheep's back; and most of them give details as to occasions on which they have shot the birds while at work.

It must be borne in mind that some of my informants are and always have been managers, and so are less likely to have had the opportunity of witnessing the attacks than the shepherds; and, further, as Mr. Ford states, the attacks are frequently, if not usually, made during the night, and therefore, unless specially watched, their attacks are rarely witnessed. Nevertheless, all the men to whom I wrote have actually seen the bird at work. My correspondents are all, I believe, trustworthy witnesses, and I see no reason to doubt the truth and accuracy of their statements.

Area of Country affected.

The kea is confined to the South Island, and occurs only in the high mountainous parts—i.e., along the Southern Alps and other high ranges. But it is not throughout this country that the kea has been a nuisance to sheepowners. In travelling through Marlborough or in North Canterbury my inquiries met with a negative reply—indeed, some of the men to whom I put the question scoffed at the idea of the bird doing serious damage to sheep. From further inquiries, however, I find that it has been known for some years that along the eastern flanks of the Alps in Mackenzie country, Canterbury down to Earns-law, near Lake Wakatipu in Otago, various sheep-stations have suffered loss from the attacks of the bird. Thus, I have records from Mount Cook Station and Ben Ohau Station, in the first-named district; while the Wanaka, Hawea, and Wakatipu districts yield abundant evidence of the existence of carnivorous keas. And one of my correspondents (Mr. King) gives the Takitimo Mountains as the southern limit. Further, during the present year the farmers of Amuri County, in North

[Footnote] * Throughout this article words or phrases in square brackets are interpretations or paraphrases of mine.

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Canterbury, met in conference at Culverden and decided to petition the Government to increase the bonus on kea-beaks.

In the Otago Daily Times for the 16th February, 1906, we read that “the keas have been very numerous in the mountainous parts of Amuri County during the last two years. They have descended on the Amuri highlands from the mountains, and several landowners stated that the losses of sheep attributed to these birds had increased from about 7 ½ to 8 per cent. last year to 15 per cent. this year. Musterers brought reports of having seen sheep killed by keas,” &c., &c.—a repetition of the course of events well known in Otago. While from the same newspaper for the 22nd March, 1906, one learns that on the West Coast, too, the kea is at work: “At Mahitahi, Bruce Bay, in South Westland, Mr. T. Condon lost last year 150 sheep, which he believed to have been killed by keas. He had seen the bird at work on the sheep. … There has been no complaint of the birds attacking the settlers' flocks in the more northern sections of the Coast—as, for instance, round the Franz Josef Glacier.”

But even within the limit of distribution of the kea many stations seem to be free from its attacks. Thus, Buller(13) quotes H. Campbell as stating that, in 1868, when he was suffering from the attacks at Wanaka, the keas were not attacking the sheep at another station owned by him, some thirty miles away, “at the same altitude (4,000 ft. to 5,000 ft.), in the same district, and where the birds are plentiful”; and my own inquiries quite bear out this evidence of the sporadic distribution of the habit.

Is every Kea carnivorous?

I put this question to my correspondents, and the replies render it impossible to give a definite answer: for whereas Messrs. Bell, Cameron, Ford, and Holmes believe that every kea in a district indulges in the habit—as “the old birds teach the younger directly they are fledged,” writes Ford—there are some of my informants who take the opposite view, that “only the more daring bird's” attack sheep (McGregor); that “the habit is peculiar to individual keas in a flock” (McKenzie); and this view is held by Mr. Green of the “Hermitage,” Southern Alps; and I am inclined to share the latter opinion, for if every kea in the district were to become carnivorous the loss to sheep-owners would be much greater than it is.

There are many stations, as I have stated, in the area occupied by the kea whose owners have apparently no fault to find with the bird, and even in those districts where they are most troublesome it is probably only certain birds that are

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the victims of this craving for flesh. It is—as some of my correspondents put it—only the more daring and older birds that attack the sheep. Just as, in India, there are only certain tigers that, unprovoked, indulge in human flesh—the “man-eaters”—so, amongst the keas, we must recognise certain “sheep-eaters,” though whether their number is on the increase seems unascertained.

The Kea is still at Work.

It will be seen from the letters that the kea is still at work in, at any rate, certain districts, whereas in some of those stations at which they were formerly a pest the birds at present appear to be less addicted to the habit. This may be owing to the fact that on certain stations it is no longer profitable to keep sheep (see King's letter), or because possibly the “sheep-eaters” have migrated to other regions. At any rate, at Hawea, D. Bell “shot three keas attacking living sheep on the 12th January, 1906—shot one on the sheep's back. I shot eight keas during the term we were mustering in January, and their crops were full of mutton.” And he says that “they kill sheep now as much as ever.” Other evidence from Makarora is to the same effect: “The sheep still come in more or less wounded, and even with the entrails hanging out, and when killed for food, as many as one in four present healed wounds in the back” (Ford). But one thing seems certain, that the birds now go more thoroughly to work, and make a more complete “job” of it than in old days. Then the sheep were usually merely wounded. True, the wounds may have been serious, and some of the sheep died of their wounds. But nowadays the carcase is devoured, the bones are left picked clean. Mr. D. Bell sent me the arm-bones (humerus) of sheep from which the keas had actually extracted the marrow. In each of the four bones sent, a more or less triangular hole had been neatly made by the kea, just below the “head.” (Plate IV.)

Mr. King writes, “One thing has been noticed lately by those who were among the sheep in high country—viz., that the keas when they kill a sheep now pick the carcase clean; leaving nothing but the skeleton, the skin almost turned inside out.”

Mr. Bell also refers to this increase in the damage done to individual sheep. It has been suggested by Taylor White(8) that originally the bird merely sucked the blood, but that it soon discovered that the flesh itself is good to eat, and, as we learn, now recognises the “importance of being thoroughly in earnest” by devouring the entire carcase.

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It will be remembered that Wallace states that the bird showed a marked preference for the “kidneys,” and it is very popularly supposed that it makes a special effort to obtain these organs; but, as the majority of my correspondents point out, this is an error. It is a mere accident that the bird, in certain cases, makes its first attack in the region of the kidney, for it is on the rump that the bird finds it most convenient and safest to settle, and naturally pecks at one side or the other of the backbone, working through to the underlying kidney. But, as-a matter of fact, the bird will commence the attack at that part of the sheep which is most prominent: if a sheep be lying on its side in a gully or elsewhere the kea commences at the side of the abdomen.

The eclecticism of the kea is in that exaggerated, as has already been pointed out years ago by Huddlestone and by Taylor White; but such a legend dies very hard indeed.

The Method of Attack.

The method of attack is varied. It may be (a) on the moving sheep; (b) on a fallen sheep; (c) on a snowed-up sheep.

Perhaps the best account of the first method is that contained in Mr. R. McKenzie's letter: “I have seen a kea attack and hang on to a living sheep. The bird flew on to a sheep's back, and commenced driving its beak through the wool into the flesh—not necessarily just over the kidney, as is supposed to be the custom. The frenzied sheep jumped and ran about in any direction for dear life, then, separating itself from the mob, made a direct line down a steep slope, and in its mad career finally dropped over a precipice, until which moment the bird held on with its claws, its wings slightly extended as if to steady itself or to be ready to fly off at any moment. The instant the victim left terra firma the bird relaxed its hold, but was observed to fly almost straight down as if bent on securing the sheep. Both were then lost to view.”

Similar statements occur in other letters (Bell, Cameron, McGregor), and have been made to me by shepherds and others orally.

(b.) A sheep will sometimes, in coming down hill, roll over and perhaps lodge in a gully or elsewhere, and be unable to rise. The kea will then attack. I quote from a letter to the Otago Witness, November 22, 1905, written by Mr. J. A. Wraytt, of Gareton: “When travelling along the bridle-track down the east side of Lake Wanaka I saw a sheep some 50 yards below, kneeling down with its head poked under a shelf of rock. There was a kea on its back, and about half a dozen sitting

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on the rocks close by. On going down to the spot I found the sheep, which was or had been a strong, fat one, with most of the wool stripped off its back and lying around in small tufts. There was a large hole through the sheep's loins, into which the kea kept diving and filling its mouth. The others had apparently had their fill, as their heads were drenched with blood.”

(c.) Frequently when the sheep are snowed up, and weakened no doubt by want of food, they will fall victims to the birds. Thus Mr. E. Cameron writes, “A snowslide carried a sheep with it. I happened to be on the hill about the time it happened. It was all covered with snow but its nose and one hind leg, but still alive. The uncovered leg was eaten to the bone—not a scrap left on it—with half a dozen keas fighting over it.”

The kea is known to be a very fearless bird, and we have a record in Mr. King's letter: “One man, who has had a good deal of experience in the high country, tells me that on one occasion, at Lake Ohau, in the Mackenzie country, he saw two keas attack a mob of sheep standing in the yards at the wool-sheds of that station. Shearing was going on at the time, and this was seen by all the shearers present. The birds were shot.”

The sheep do not necessarily die from the attack, for, as mentioned, the animals are frequently observed to present healed wounds when being shorn; and cases are known in which part of the intestine has been actually torn away, and the broken end has adhered to the edges of the hole in the body-wall, so as to form a new exit for the dung. Such a case is recorded in Buller's work, and one of my correspondents—Bell—states that he has met with two instances of it.

The sheep that are selected by the keas as victims are apparently always those in good condition, and with long wool, especially such as have missed a shearing; whereas a newly shorn sheep, or one in poor condition, is never attacked. One readily understands the choice of a well-covered sheep, as it affords a firm foothold for the bird.

Period of the Attacks.

The attacks are most frequent in the spring and winter—when presumably the natural food of the bird is scarce, or covered with snow—but by no means are they confined to these seasons. Further, it may be noted that the night-time is the period during which much of the damage is done, though the bird is not absolutely or entirely nocturnal.

Any one who has spent a few days in the region of the Southern Alps knows that the birds may frequently be seen

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during the day; and many people who have spent a night in the Ball Hut on the side of the Tasman Glacier will recall their noisy habits in the early morning, before the dawn.

The Normal Food.

The kea, like other parrots, is normally a vegetarian, though it includes insects in its diet. At any rate, so it appears from the few reliable statements that exist.

According to that accurate observer, the late Mr. Thomas H. Potts(1), the kea gathers “its subsistence from the nectar of hardy flowers, from the drupes and berries of dwarfed shrubs that contend with a rigorous climate and press upwards almost to the snow-line of our alpine giants. To these food-resources may be added insects found in the crevices of rocks, beneath the bark of trees,” &c.

Mr. Potts also states(2) (“Out in the Open,” quoted by Buller in “Birds of New Zealand,” i, p. 168, footnote) that when the snow covers these subalpine shrubs, and insect-life is dormant, the kea is forced to go lower and lower down the mountains to take shelter in gullies, where it feeds on the hard, bitter seeds of the kowhai [Sophora tetraptera], small hard seeds in the fruit of Pittosporum, the black berries of Aristotelia fruticosa (the “native currant”), as well as on the fruit of the pitch-pine and the totara.

The most detailed bill of fare is that given by Huddlestone(6): it includes the grub of such insects as the weta (Deinacrida) and cicada, which are to be found in the ground. “Besides grubs, they fed on the berries of various alpine shrubs and trees, such as the snowberry, Gaultheria, Coprosma, Panax [= Nothopanax], the little black seed in a white skin of Phyllocladus alpinus, the Pittosporum with its hard seed in a glutinous mass like birdlime, and the red berry of the Podocarpus [nivalis], also on roots of various herbaceous plants—Aciphylla squarrosa and A. Colensoi, Ranunculus lyallii, celmisias, &c.”

My own observation, while in the Southern Alps this year, adds to the list the orange berries of the low-growing heath Leucopogon fraseri. Two birds were feeding on these berries within two yards of where I was sitting: they eat the juicy part of the berry, putting out the skin and usually the “seed” also, which I found afterwards on the ground, though now and then I heard the bird crack the seed, so that occasionally, at any rate, it swallows this.

Both Potts and Huddlestone refer to it eating the “hard seeds” of Pittosporum and Podocarpus; but I wonder whether this is habitual; why should they neglect the juicy covering?

Mr. Cameron writes, “The food of the kea is berries, roots of

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spear-grass (Aciphylla) and cabbage-tree [Cordyline], and snow-grass [Danthonia sp.]”.

Several of my correspondents note the fact that keas eat “roots,” though they are not in agreement as to what the roots are. Mr. Taylor White (8) suggests—though, on what evidence does not appear—that the bird feeds on “lichens which cover the rocks in high mountainous regions.” This, it seems to me, would be a very innutritious diet, and it is little likely that, in the presence of a fairly abundant choice of juicy berries and other fruits, the bird would touch so poor a food. But it appears from his article that there were no berry-bearing subalpine shrubs in the locality he was acquainted with. Certainly around the “Hermitage,” in the Mount Cook district, the subalpine scrub is abundant, and lichens would, one imagines, be the last resort of the bird.

Mr. McGregor writes, “I have watched a kea picking grubs out of a dead tree, and frequently noticed them picking into the earth for roots, with their beaks.” But none of the above observers, or, so far as I can ascertain, any one else, seems to have examined the crops of any of the birds in an untroubled district—where, that is, the carnivorous habit has not shown itself—so that it is difficult to determine with absolute certainty the whole range of the normal diet of the bird.

The Origin of the Carnivorous Habit.

The above being its normal food, how has it come about that the bird has taken to eating mutton? Various suppositions have been put forward. One of these may at once be disposed of. It has been suggested that the kea mistook a sheep lying down for the plants termed by settlers the “vegetable sheep” (Raoulia mammillaris and R. eximia)! Thus Mr. (now Judge) F. R. Chapman wrote some years ago (7), “It is said that the keas tear them (the plants) up with their powerful beaks, and that these birds learnt to eat mutton through mistaking dead sheep for masses of Raoulia.”

Now, as a matter of fact, these large species of Raoulia do not occur in the Wanaka district, nor on the Southern Alps in the neighbourhood of Mount Cook. None of my correspondents, all of whom know the country round Wanaka well, mention the plant as providing any sort of food for the kea. I think that Mr. Chapman must have been misled. Further, it is extremely doubtful (see later) whether the keas devour “dead” sheep—i.e., such as are found lying on the hills, that may have died of “natural causes”—one, in short, that the birds have not killed themselves.

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This “vegetable sheep” business is another myth that ought to be eliminated from the history of the habit.

There can be no doubt that the origin of the habit is traceable to the kea's natural curiosity: its bump of inquisitiveness is very highly developed, and it will investigate any unusual object—turning it over, peeking at it, and so forth.

It is very easy to imagine a kea coming across a fallen still-living sheep or one partially covered by snow, proceeding to pluck at the wool, and so coming down to the skin; then its beak would nibble at the flesh, and the bird would soon find that blood is a good sort of juice to swallow.

Or, as some have suggested (e.g., Mr. McGregor), the bird inquired into the fresh skin of a sheep hanging on a fence, or was attracted by a carcase, newly killed, suspended from a gallows: either of these possibilities seems probable, though only one or two of my correspondents refer to this matter, in reply to my query. It does not seem certain, however, that a kea will feed on a sheep that has died from “natural causes” on the hills. Mr. T. White and several of my correspondents state that they have never seen them thus feeding. At the same time, one of my correspondents refers to the bird devouring the carcase of deer that have been shot and left on the hills; and Buller refers to a dead foal being similarly eaten; and it is also stated that the corpse of a man who fell over a precipice was found torn to pieces by keas. Reischek(5) made the ingenious suggestion that the birds acquired the habit from finding and feeding on maggots which had appeared on the carcase of a sheep which had died on the hills, and “having thus acquired a taste for fat, became emboldened to attack live sheep.”

But these are all suppositions, and if it be true that the bird does not in fact touch a dead sheep, these suppositions cannot be true.

Whether the habit is related in any way, originally, to a scarcity of food does not seem at all clear; the answers are all rather vague, and there is a diversity of opinion on the matter. Thus Mr. King writes, “I do not think that scarceness of the natural food had much to do with the attacks. I have been told that the kea attacks sheep in the West Coast in the low country close to the bush, where food would be plentiful.” But that this scarcity has occurred is certain. The burning-off of the alpine scrub and the bush in gullies, as has occurred in many districts, would deprive the bird of its normal food during the whole year; while elsewhere, when the normal food is covered by snow in winter-time, the bird would be compelled to seek a new diet, or migrate, or die. The kea has met the problem by adopting the first of these three alternatives; and as, at the same time as

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the burning occurred, the sheep were introduced into the district, a new source of food was at once open to it. And the suggestion that the kea, on the disappearance, temporary or permanent, of its normal food, would proceed to investigate a fallen sheep, or one snowbound—weakened perhaps from cold or absence of food—seems quite in accordance with the bird's general habit of inquiry and its catholicity of diet.

The change of diet is not so abrupt as at first appears, since part of the bird's food consists of flesh in the form of insects, and possibly there is not a great amount of difference in taste between a good, fat, juicy weta or beetle grub and a piece of raw sheep, while the case with which a whole flock of birds can obtain a full meal must be a distinct improvement upon the more wearisome task of digging in the earth and probing rotten logs for a small mouthful as a reward.

The observations of birds kept in captivity are few, and not entirely in agreement. Dr. DeLautour (as quoted by Buller from the Field) noted that his bird—which was the first living specimen to be exhibited in the Zoological Gardens in London—ate only the flesh and would not touch the fat, preferred mutton to beef, and was not averse to pork. Reischek states that a kea he had in captivity preferred meat to vegetables.

I had a caged kea under observation for a week, through the kindness of Mr. Harry Buckland of Waikouaiti. We fed it normally on various vegetables, such as carrots and parsnips, and on fruits, such as apples and bananas, all of which it seemed to like. We tried it with mutton flesh and fat, and so long as we watched it the bird neglected the meat, though it ate it during the night. On two occasions we presented it with a saucer containing cut-up vegetables, mutton—lean, fat, and kidney. It went to work at once on the vegetables. On one occasion it did not eat the mutton so long as daylight lasted, but during the evening we found that it had devoured the flesh, later on the fat, and next morning the pieces of kidney had disappeared. On the second occasion—on which it had not been fed during the earlier part of the day—it again attacked the vegetables first, later the flesh, then the fat, and lastly, during the night, the kidney. Of course, this bird may not have developed a carnivorous habit before its capture, some three months previously, for, as I have pointed out, only some keas exhibit the taste for mutton. Moreover, it must not be supposed, I think, that even a bird that has once developed the habit eschews vegetable diet thereafter; it is likely that its diet will be a “mixed” one.

Nor is the kea alone in evincing a liking for flesh; its near ally the kaka (Nestor meridionalis), normally a honey and

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grub eater, has been observed to devour mutton. Travers (3) states that “They are fond of raw flesh, and I have seen them hovering in front of a sheep's pluck hung on a tree… eating fragments which they tear off, giving preference to the lungs.”

At the time that Wallace referred to this new food-habit of the kea it was the only case of the sort known. But a closely analogous instance has been described for an African starling, the “rhinoceros-bird” (Buphaga), which formerly fed upon ticks and insects infesting the skin of herbivora. But a few years' ago the cattle-plague decimated the herds of wild and domestic cattle, antelopes, &c., and it is now found that the bird, thus deprived of its natural food, has become carnivorous. It pecks holes in the skin of healthy beasts, and even eats their ears off, causing wounds from which the animals frequently die—at any rate, causing considerable damage (9).

Correspondence and Inquiries.

I wrote letters containing a series of questions to some fifteen persons, whose names had been given to me by various people as likely to have first-hand acquaintance with the bird and its depredations. These fifteen persons are, or have been, actively engaged in connection with sheep-runs in the neighbourhood of Lakes Wanaka, Hawea, and Wakatipu, and elsewhere. I have received replies from ten, which replies are embodied in the present article.

To these ten gentlemen I am extremely obliged for the courtesy and readiness with which they gave me every information I asked for:—

Dougald Bell, now owner of Hawea Lake Station, has had thirty-three years experience of sheep-farming in the district round Lakes Wakatipu, Hawea, and Wanaka. His first observation was in the Hunter Valley (Hawea) in 1874; his last, in 1906, at Lake Hawea.

Ewan Cameron, of Pembroke, was shepherding on the Crown Range in 1868; and has had experience of kea-attacks on the Matukituki, and Matatapu, and right branch of the Shotover.

John Campbell, now of Cromwell, at one time shepherd at Wanaka West.

William Ford, at one time shepherd on Mr. H. Campbell's station, has ever since been sheep-farming in the Wanaka district.

Alexander Fraser, now Stock Inspector in Nelson, was at one time sheep-farming in the Wanaka and Hawea districts (1871-83), and suffered severe losses of sheep from kea-attacks.

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M. Stuart Holmes, now of Kakanui, has been sheep-farming since 1874, and was at one time manager of Lake Wanaka Station (1881-82).

John King, of Pembroke, was at one time engaged as “kea-shooter” on H. Campbell's station in the “seventies,” and has had a long experience in that district.

James MacDonald, now a farmer at Dipton, Southland, at one time head shepherd on Henry Campbell's station at Wanaka.

Robert McGregor, of Hawea Flat, speaks of the attacks at Triplet Peaks, Lake Hawea, from 1877 to 1883.

Roderick McKenzie, now of Birchwood, Southland, was part owner in 1889 to 1891 of Hawea Lake Station.

In addition to these correspondents, have received information of kea-attacks from Mr. James W. R. Green, now porter at the Hermitage Inn, Southern Alps. He had been employed as rabbiter at Mount Cook Station, and has shot keas while attacking sheep. And from some other sources I have obtained information.

In addition to answering a series of questions which I sent them, several of the above gentlemen wrote letters containing their personal experiences, which appear to be of Sufficient interest to entitle them to be placed on record.

1. Mr. Dougald Bell.
“Hawea Lake Station, 20th February, 1906.

“In reply to yours of the 20th November re keas killing sheep … I can safely say that they kill 5 per cent. of the flocks in all the country that is infested with keas. As a matter of fact, this season in my small flock [number not given] during the time we were mustering I personally counted twenty-five sheep killed by keas; I also shot three keas killing a live sheep. I shot one of them on the sheep's back, tearing away at the kidney-fat and meat. [The majority of my correspondents do not lay stress on this, and no doubt Mr. Bell merely means that the bird was tearing away at this region.] After I shot the keas, the sheep, a big, strong, half-bred wether, which was holed all along the back and ribs, managed to get up and walk away, but he would be sure to die, as he was too much holed and worried to live. I shot eight keas during the time we were mustering in January last, and they were all full of mutton in their stomachs [crops].

“There is another matter I should like to point out to you about keas: when they have eaten all the flesh off the bone, then they tackle the shoulder [i.e., humerus] and leg bones, and take all the marrow out of the bone by chipping the bones

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with their beaks until they obtain an entrance. I am sending you four shoulder-bones, some old, and some fresh ones killed last winter. [One of these is here figured.]

“There is also another point: Two sheep came in during mustering at Hawea Lake—a ewe and a wether—which had been attacked by keas some time during the season. Each had holes in their backs, and the main gut had been cut through and pulled up through the backs; the gut had grown to the [skin of the] back and [a new anus had been formed] which caused a black streak down each flank where the droppings fell out. One of these cases happened in the year 1887 and the other in 1899. It was in the month of January that they mustered in each case, and brought into the station for shearing when [the above facts were] noticed.

“I have had thirty-three years' of experience with keas, and so know a little about them. I first came up here in 1873. When the keas first started to attack sheep they used to pull a tuft of wool off the back of the sheep over the kidneys for a while before killing [I presume he means that they merely tasted the flesh and left the sheep], but now it is very different, as nearly every sheep they tackle they kill outright.

“There is a lot of country in this locality that could be stocked with sheep if it were not for the keas: but it is not safe to put sheep on it, as the birds would kill half of them.”

2. Mr. Ewan Cameron.
“Pembroke, 28th September, 1905.

“The first notice of keas killing sheep in Wanaka was in 1868 by James McDonald, head shepherd for Mr. Henry Campbell, owner of Wanaka Station at that time … Mr. Campbell wrote a letter to the Dunstan Times describing the destruction that keas were causing among his sheep, with the result that all the people in that part of the country laughed at him.

“In that year I was shepherding in the Crown Range, and after reading Mr. Campbell's letter I saw at once what was killing my sheep. The place of attack is nearly always on the loin, behind the last rib; they [keas] tear down to the kidney, pull out the entrails, and sometimes leave the sheep without killing it. It is a common thing for sheep to come into the yards with their entrails hanging over the side; also with new wounds, or old ones healed up.

“[As an instance of the ferocity of the keas, I may mention that] One season, at the head of the Matukituki, I had four hundred sheep that did not come in at the proper time for shearing. I put them in a safe place after snowfall at the beginning

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of winter: when I went to shift them on the 1st September [I found that] the keas had killed two hundred of them: a good many were devoured, and some not touched but with the usual wound above the kidneys.

“Rose Bros. had a run on the Matatapu, a continuation of the same range I was on. They mustered their sheep (about three thousand) in the beginning of winter, left them in a large mountain paddock at night, and next morning found thirty-five killed.

“They [the keas] do most of the damage at night. On another occasion, on my own run, a snowslide carried a sheep with it. I happened to be on the hill about the time it happened, and saw the sheep still alive but covered with snow except its nose and one hind leg: the uncovered leg was eaten to the bone, not a scrap [of flesh] left on it, and half a dozen keas fighting over it.”

3. Mr. John Campbell.
“Cromwell, 13th August, 1906.

“I have much pleasure in telling you what I know of this wonderful bird. I first saw the kea in the Wanaka Lake district in '68…. I was shepherding at this time on what was then called ‘Campbell Station’ (Wanaka West), and often killed numbers of keas when out on the mountains towards nightfall…. As regards their attacks on sheep, I think I was the first to see the kea on a sheep's back. The shearers often drew our attention to a scar on the sheep's hack opposite the kidneys, and being on the same place on each sheep, some thought it was a disease (for some were found dead wounded in the same way). The kea was not at all suspected until some time afterwards.

“I was coming down the Matatapu (boundary between Wanaka and Wanaka West Stations)* when I saw a mob of sheep rushing about as if a dog was disturbing them. When I got nearer I saw a nook of birds hovering over the sheep. These were keas. I stopped and watched for a few minutes, and presently I saw a sheep get singled out from the others and make towards some rocks with a kea planted on its back. By running under rocks and rubbing against them the sheep got the bird off its back, but the same one or a fresh one was soon on again. This went on for some time till the sheep was exhausted. I

[Footnote] * In a second letter he writes, “It was late in the year '70 when myself and mate, David Kinnard, were coming home from Glencoe to Glendhu on Wanaka Station. We saw sheep on the hill-top disturbed as by dogs. I went to the top, but Dave would not come up.”

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then went down and drove away the birds and examined the sheep, which had the wool and flesh torn by the birds in the same spot as that previously noticed by the shearers. This solved the mystery, and when I related the story to the station-manager and fellow-workers they were greatly surprised to learn that the kea was the culprit….

“I am convinced that the chief reason why the bird attacks the sheep in that particular spot is because it is the only place that it can perch for any length of time without the sheep putting it off.” [He does not believe in the keas seeking after the kidney-fat.]

4. Mr. Alexander Fraser.
“Nelson, 2nd August, 1905.

“In reply to yours of the 22nd ultimo, I was engaged sheep-farming in the Hawea and Wanaka Lake district between 1871 and 1883. Suspicion arose in the first-named period that keas were attacking sheep, suggestion being that they learnt it from picking at sheep-skins and carcases hung on gallows. I lost some thousands of sheep from keas. I have seen the kea attacking the sheep, and also eating into a sheep when the latter was stuck in deep snow. I have opened scores of keas' crops and found wool and meat therein. I have laid poison in dead sheep in snow, gone back later and found dead keas; also have often-poisoned keas with mutton suet [poisoned, I presume]. The natural food originally of the kea was berries, and grubs and insects it dug out of the ground. The burning of the alpine country probably diminished its natural food. They breed inside broken rock. Never found a nest. Keas are very numerous between the above dates, though shot and poisoned by the thousand.”

5. Mr. John King.
“Pembroke, 22nd July, 1905.

[Part of this letter has already been quoted above, and many of the facts contained in it are incorporated in the general account. It continues:] “I have seen sheep snowed up on the ranges, and on one occasion I counted twelve dead out of about fifty, and numbers of keas sitting about on the rocks gorged, very much after the manner of vultures. On another occasion I was with Mr. Campbell and some of his shepherds who had a large mob of sheep in front of them taking them across some hilly country, when we noticed two keas hovering over the sheep. Presently one swooped down on a sheep not 30 yards from where we were. This sheep immediately left the others and rushed past me. I was carrying a gun at the time, and shot the bird

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on the sheep's back. I could recount many instances where I have seen the keas at work, but probably what I have written may suffice.”

6. Mr. R. McKenzie.
“Birchwood, Southland, 23rd October, 1905.

[I have quoted portion of Mr. McKenzie's letter on page 78, describing one method of attack, as being a very clear account.]

“The habit of attacking sheep I firmly believe to be peculiar to individuals in a flock. I also believe the reputed slaughter by keas of hundreds or even scores of sheep in a single night to be gross exaggerations. On one occasion, during a snowstorm, when two or three hundred sheep had been hemmed in for a few days, I found three or four sheep killed and mostly eaten up by the birds. …

“With regard to the bird digging in over the kidney, I believe this to be accidental rather than by design or instinct, for as the maddened sheep tries to escape by running away—always down-hill—the bird hangs on to the highest part (that is then the rump just over the kidney) of the sheep, and begins operations there.”

Literature.

(1.)

Potts, 1871: Nature, iv, 489.

(2.)

Potts, “Out in the Open.”

(3.)

Travers, 1872: Trans. N.Z. Inst., iv, 210.

(4.)

Menzies, 1878: Trans. N.Z. Inst., xi, 376.

(5.)

Reischek, 1885: Trans. N.Z. Inst., xviii, 98.

(6.)

Huddlestone, 1891: N.Z. Journ. Sci. (n.s.), 198.

(7.)

Chapman, 1891: N.Z. Journ. Sci. (n.s.), 203.

(8.)

Taylor White, 1895: Trans. N.Z. Inst., xxvii, 273.

(9.)

Lankester, Nature, lxii, 366.

(10.)

Otago Daily Times, March 22, 1906.

(11.)

Otago Daily Times, Feb. 16, 1906.

(12.)

Benham, 1906: Nature, lxxiii, 559.

(13.)

Buller, “History of New Zealand Birds,” i.

Explanation of Plate IV.

Outline of humerus of sheep which has been opened by a kea.—a, a. The surface of this bone has been picked away, exposing the canceller structure. b. The cavity of the shaft exposed by the removal of a somewhat triangular piece of bone.

All the four humeri had been opened by the kea at this spot, in order, no doubt, to get at the marrow.

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Art. IV.—Notes on the Distribution of Ores in Horizontal Zones in Vertical Depth.

[Read before the Otago Institute, 8th August, 1905.]

It has been clearly demonstrated by mining operations in the United States, England, and elsewhere that while in many veins the metallic sulphides are intimately mixed without any definite arrangement, in other veins, particularly those of lead, zinc, and iron, there is a more or less orderly distribution in horizontal zones in a vertical direction: that is, there are certain horizontal zones each of which is characterized by a dominant sulphide.

This arrangement of the metallic contents of a vein in more or less horizontal zones was noticed in Cornwall many years ago; and no better example could be found than that presented by the celebrated Dalcoath Mine, which commenced as a tin-mine, at a lower depth yielded nothing but copper, and again below that, tin.

In the great lead and zinc-mining region of Ozark, in the lower Mississippi Valley, the vertical distribution of the ores, according to Bain,* is as follows:—

(1.)

Oxidized zinc and lead ores, with galena.

(2.)

Blende, with a little galena.

(3.)

Iron-sulphide predominates, and increases with depth.

Spurr, in his report on Monte Cristo mining district, in Washington, states that the quartz, pyrite, chalcopyrite, pyrrhotite, blende, galena, realgar, stibnite, and calcite show a marked tendency to aggregate themselves in horizontal zones in the order named above.

Rickard mentions the orderly distribution of ores in Colorado. Weed states that in the Castle Mountain district, in Montana, the order appears to be galena on top, passing into highly zinciferous ores below, and these into low-grade pyrite.§

[Footnote] * H. F. Bain, U.S. Geol. Survey, Twenty-second Annual Report, Part II, p. 161.

[Footnote] † J. H. Spurr, loc. Cit., p. 841.

[Footnote] ‡ J. A. Rickard, Trans. Inst. Min. and Met. London, Vol. vi, 1809, p. 196.

[Footnote] § Weed and Pirsson, Bull. 139, U.S. Geol. Survey, 1896.

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At the Broken Hill mines in New South Wales the general distribution of ores in vertical depth has been as follows:—

(a.)

Oxidized ores of lead and silver.

(b.)

Galena with blende.

(c.)

Blende with galena.

Weed, in his paper on “Ore-deposition and Vein-enrichment by Ascending Hot Waters,”* appears to support the hypothesis which assumes that the distribution of ores in horizontal zones is the result of primary concentration by ascending hot solutions.

The eruption of igneous magmas is often succeeded by intense solfataric action of which notable examples are found in the Yellowstone Park in the United States, and in the volcanic region of the North Island of New Zealand. The ascending waters slowly circulating in contact with the heated rocks below become superheated, and in their upward course dissolve various, substances, which they carry with them along the line of least resistance—that is, towards the hot-spring pipe or vent. Many substances insoluble in normal conditions are rendered easily soluble in the presence of heat and pressure. The underground water will therefore possess its greatest solvent power where the greatest heat and pressure are attained, which will naturally be at the greatest depth. With loss of heat and pressure the less soluble substances held in solution will be precipitated—that is, those substances whose dissolution was effected under extreme heat and pressure. As the waters ascend they will continue to lose heat and be relieved of pressure, with the result that the dissolved. minerals will be precipitated in the inverse order of their solubility. When the hot waters reach the surface the only substances in solution, in most cases, will be the extremely soluble alkaline sulphates, carbonates, and silicates. An obvious result of this process of vein-filling will be an impoverishment of the veins at great depths, due to the migration of the valuable minerals from below to the zones of precipitation above. It is a notorious fact that hot springs seldom deposit metallic sulphides at the surface. The great mushroom-capped veins of the Hauraki region and Great Barrier Island, in New Zealand, are composed of siliceous sinters, chalcedonic and crystalline quartz, manifestly the result of long-continued solfataric activity. The overhanging mushrooms of quartz are almost devoid of gold and metallic sulphides; but the necks in all cases contain gold, and sulphides of silver and iron.

[Footnote] * W. H. Weed, Amer. Inst. Mining Eng., Vol. xxxiii, 1903.

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Had denudation removed the mushroom caps, the sulphide-bearing necks would now be exposed at the surface.

The well-known Martha Lode, at Waihi, consists of chalcedonic and crystalline quartz, apparently the result of hydro-thermal activity, which at one time probably manifested itself at the surface. There is no overhanging cap.

At the outcrop the quartz is almost pure silica, containing no sulphides excepting a trace of argentite associated with free gold containing about one-third its weight of silver. Above water-level the ore is clean, and free from oxidized products.

In many places both above and below water-level the joints in the Veinstone axe discoloured with films of manganese and iron oxides, which appear to owe their origin to the infiltration of meteoric water from the wall-rock, and not to the oxidation of contained sulphides.

Between the adit level and No. 1 level there began to appear small limonite-crusted cavities in the thin veins of crystalline quartz which traverse the main lode. At No. 1 level there are detached branches of iron-pyrites in the quartz, and at No. 2 level the sulphide ore forms a rib two or three feet thick.

The lode is being worked to a depth of 750 ft. below adit level; and although there has been an increase in the proportion of iron-pyrites, there has been no decrease in the gold and silver values.

A greater measure of denudation than the lode has already suffered would have exposed the sulphide ore at the surface.

In the study of vein-filling it is always well to bear in mind that veins which outcrop at the surface may have been truncated to a greater or less degree by denudation.

After their formation, some veins, through movement of the walls, have been brecciated and recemented by circulating mineralised waters. Such waters, ascending through the crushed vein-matter, would deposit their metallic contents as sulphides through the reaction of primary sulphides contained in the ore.

In this way a secondary concentration of sulphide ore may be effected by ascending waters. The common belief, however, is that secondary enrichment is in the majority of cases the result of the transference of material from the oxidized portions of a vein to a lower level through the agency of descending waters, from which the metallic contents are precipitated by the reducing action of organic matter or primary sulphides.

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Art. V.—Notes on the Formation of Zones of Secondary Enrichment in certain Metalliferous Lodes.

[Read before the Otago Institute, 8th August, 1905.]

It has been noted in many parts of the globe that masses of ore of exceptional richness often occur in the oxidized portion of the ore-body—commonly in that portion lying at the boundary of the oxidized and unaltered sulphides.

Microscopic investigation has proved that these rich masses are not of primary but of secondary origin. Their genesis is supposed to be due to the migration pf the valuable contents of the upper part of the vein to, and their concentration in the lower part of the oxidized zone.

In some cases the processes of dissolution, migration, and redisposition may have taken place over and over again, each cycle resulting in an increasing degree of concentration.

The veins in which secondary enrichment are most often seen are those of gold, silver, copper, lead, and zinc.

Gold-ores, in the zone of weathering, are often augmented in value by the long-continued disintegration of the vein and the enclosing rock, whereby the gold set free from its matrix is permitted to concentrate at the outcrop.

Extensive areas of the Australian Continent have been subject to subaerial denudation since the close of the Palæozoic period; and it is doubtless due to this cause that so many notable examples of mechanical enrichment of gold-bearing veins have been found in Victoria and Western Australia.

The migration of gold, copper, silver, lead, and zinc from the upper to the lower parts of the veins is effected by descending surface waters in the zone of vadose circulation. The processes involved in the migration are chemical dissolution and electrochemical deposition.

Chemical processes may operate in various ways to cause secondary enrichment, as follows:—

(a.)

By the removal of worthless metals, thereby leaving the valuable contents in a purer form.

(b.)

By removal of worthless metals, and their replacement by valuable metals removed from a higher part of the vein.

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(c.)

By deposition of valuable metals on primary sulphides in those portions of the vein subject to the influence of circulating surface waters. In this case the primary sulphides may form the nuclei for the deposit of the secondary sulphides.

Manifestly the first operation in the process of secondary enrichment is the chemical weathering and oxidation of the metallic contents of the vein.

The oxidation of base sulphides can be seen in operation every day. In a mass of mixed sulphides of iron, copper, zinc, and galena, the iron will be the first to be affected, from its affinity for oxygen. Iron-pyrites is decomposed and forms ferrous-sulphate, which is changed into Fe(OH)3, Fe2(SO4)3, and H2SO4. The H2SO4 attacks fresh iron-sulphide and forms ferrous sulphate, liberating H2S, which at once combines with free oxygen to form H2SO4. The ferrous sulphate changes to the ferric sulphate, which attacks gold and sulphides of copper, lead, zinc, and silver. The process of dissolution is necessarily slow, on account of the extreme dilution of the solutions.

For many years it was believed that the only secondary enrichment that could take place was the formation of rich bonanzas of carbonate ores and chloride of silver, in the zone above water-level. But careful investigation has shown that primary sulphides have been enriched by the deposition of secondary sulphides even in places below the present water-level.

It was proved experimentally by Skey* in 1870 and Liversidge in 1893 that gold is more readily precipitated from its. solutions by metallic sulphides than by organic matter. Furthermore, Skey showed that sulphides of the base metals were readily precipitated from their alkaline sulphide solutions in a solid coherent form in the presence of iron-pyrites, galena, blende, stibnite, &c.

The descending acid solutions formed in the zone of oxidation will attack the constituents of the rocks through which they percolate, producing alkaline silicates and sulphides.

Gold dissolved by ferric sulphate would be also carried down and deposited as leaf, scale, or wire gold in cracks in sulphide ore, thereby causing local enrichment.

It is maintained by some writers that secondary sulphides have been found below water-level. The evidence on this question is not quite conclusive. Changes of water-level may have taken place since the secondary sulphides were deposited.

[Footnote] * Skey, Trans. N.Z. Inst, Vol. iii, 1879, p. 226.

[Footnote] † Liversidge, Proc. Roy. Soc. N.S.W., Vol. xxvii, 1893, p. 287.

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The property possessed by silica, clay, and porous substances of absorbing metals from dilute aqueous solutions may be an important factor in the formation of zones of secondary enrichment in the oxidized portions of metalliferous lodes. In this we may have the explanation of the rich kaolin ores of silver at Broken Hill, of the concentration of gold in the talcose clays of the lode-formations of Kalgoorlie, and of the copper-bearing shales of Europe and America.

The researches of Emmons,* Weed, and others have thrown much light on the secondary enrichment of vein deposits, and much still remains to be done before safe generalisations can be formed.

Impoverishment of Veins in Depth.

T. A. Rickard, when discussing Professor Posepny's paper on “The Genesis of Ore-deposits,” states that the general non-persistence of ore in depth is a fact capable of proof. He contends that since heat and pressure are the two factors which increase the solubility of mineral substances, the deep region will favour solution the most, while the shallow zone will favour precipitation owing to the decrease of heat and pressure.

There is much in favour of this contention, and many examples could be adduced in its support in all parts of the globe.

Moreover, progressive poverty in depth below a certain point must be the natural corollary of the general law governing the orderly distribution of ores in horizontal zones in vertical distance through the agency of ascending waters.

In some cases impoverishment in depth is determined by the prevailing geological conditions. Ore-veins which are confined to a single overlying formation often die but or become exhausted on reaching the underlying rock.

A notable example of this is afforded by the hydrothermal veins of the Thames, Tapu, Coromandel, and Kimotunu mining districts in the Hauraki mining region of New Zealand, where the gold-bearing veins are contained in altered andesites which rest on a highly eroded surface of Lower Mesozoic slaty shales and sandstones.

Mining operations have in all cases shown that when the veins which occur near the borders of the andesite-flows reach the basement rock they die out completely, or end in small strings which soon disappear in depth.

[Footnote] * S. J. Emmons, Trans. Am. Inst. M.E, Vol xx, 1900.

[Footnote] † W. H. Weed, Bull. Geol. Soc. Am., Vol. xi, 1900, p. 179; and Trans. Am. Inst. M.E., Vol. xx, 1900, p.

[Footnote] ‡ T. A. Rickard, “Genesis of Ore-deposits”: Discussion, p. 190.

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The principle of secondary enrichment precludes the continuance of the enriched portion of the vein downward in vertical distance.

When the values of secondary enrichment are added to ore already of a payable quality, the result is a rich shoot or bonanza; but when, as often happens, the secondary values are added to lean ore, then the net result is to render the lean ore just profitable. Hence below the zone of enrichment the ore will be lean and unprofitable.

Absorption of Metals by Clays in relation to Secondary Enrichment.

It has been noted in many mines that the ore in the zone of secondary enrichment is commonly associated with or contained in, a matrix consisting of clay or other finely divided mineral matter. Of this there are no better examples than the kaolin silver-ores of Broken Hill and the talcose gold-ores of Kalgoorlie.

Clays and clayey matter are the natural products of the alteration of rocks and ores in the zone of oxidation, hence their presence calls for no comment. But the frequent occurrence of rich ores in a clayey matrix in the zone of oxidation in certain lodes has led to much speculation as to the relation existing between the clay and its metallic contents.

It has been suggested by some writers that this association is not accidental, nor the result of paragenesis, but due to some quality of the clay. That clay and finely divided matter possess the property of absorbing or extracting metals from their aqueous solutions has long been known; and with this knowledge in mind it has been contended that the clayey matter acting as a porous filter in the lower part of the zone of oxidation has absorbed the metals from the descending solutions, thereby effecting a concentration of the valuable contents.

There is much to be said in favour of this view, but it has still to be determined whether clayey matter is a primary or merely a contributing cause in the formation of zones of secondary enrichment.

Walter Harvey Weed* early in 1905 described some. experiments made by himself and others in the laboratory of the United States Geological Survey on the absorptive property of clays, &c. The results obtained confirmed the researches of W. Skey

[Footnote] * W. H. Weed, “Absorption in Ore-deposition,” Engineering and Mining Journal, Feb. 23, 1905.

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in 1869, and of E. Kohler* in 1903, who found that clays and porous substances such as gelatinous silica, carbonaceous and colloidal matter, possess the power of extracting metals from their dilute aqueous solutions. As the subject is one having an important bearing on the concentration of metals in the zone of oxidation, and perhaps also in bed-impregnation and vein-filling, it may be of interest to notice more fully the experiments made by Skey in the New Zealand Government Laboratory in 1869, 1871, and 1874. Skey, it should be noted, was one of the pioneers in this department of research, and it is a tribute to the marvellous ability and skill he displayed in his research-work to find his determinations so fully verified by investigators of later date.

In 1869 he proved experimentally that finely pulverised massive quartz, rook-crystal, and silica possess the power of absorbing or extracting the oxide of iron from its acetate solution. He also found that prepared silica especially manifests this property if ignited at a low temperature; and, besides, takes oxides of copper and chromium from their acetate solutions. The more finely divided the silica the more apparent is the absorption.

In 1871 Skey found that when a weak ammoniacal solution of copper containing a little caustic potash is poured upon a filter of Swedish paper (cellulose), the liquid which passes through the paper is quite or nearly colourless, and the filter is found to have retained all, or nearly all, the copper of such solution.

In 1874 he showed that clay possesses the property of absorbing and fixing natural petroleum in such a way as to form a substance resembling natural oil-shale, the oil being chemically combined with the clay.§ He does not appear to have tried to ascertain the absorptive power of clay upon solutions of the metals, but his discovery that silica and porous substances, such as cellulose, possess the property of absorbing metals from their solutions has an important bearing upon the chemistry of ore-deposition.

[Footnote] * E. Kohler, Zeitschrift fur Praktische Geologie, 1903, p. 49 et seg.

[Footnote] † W. Skey, “On the Absorptive Properties of Silica, and its Direct Hydration in Contact with Water,” Trans. N.Z. Inst., Vol. ii, p. 151: Wellington, N.Z., 1869.

[Footnote] ‡ W. Skey, “Absorption of Copper from its Ammoniacal Solution by Cellulose in Presence of Caustic Potash,” Trans. N.Z. Inst., Vol. iv, 1871, p. 332.

[Footnote] § W. Skey, “Notes on the Formation and Constitution of Torbanite and similar Minerals,” Trans. N.Z. Inst., Vol. vii, 1874, p. 387.

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Art. VI.—Notes on the Origin of the Metal-bearing Solutions concerned in the Formation of Ore-deposits.

[Read before the Otago Institute, 8th August, 1905.]

Ore-deposits may be divided into two genetic classes—namely, those deposited by metal-bearing solutions, and those formed by direct magmatic segregation in an igneous magma. The former include the majority of ore-deposits of economic value; the latter are comparatively rare and unimportant.

It is manifest that the waters concerned in the formation of ore-deposits of the first class must be either (a) meteoric and descending, or (b) magmatic and ascending.

Descending waters are believed to gather their mineral contents from the rocks through which they percolate. They are said by Van Hise and other writers to descend to great depths by gravitative stress, and to move laterally towards open channels and fissures, where they deposit mineral matter. In this we find the basis of the theory of lateral secretion.

Hot ascending solutions are held to be genetically connected with igneous masses—that is, they are either directly magmatic, or they are liberated from sedimentaries in the form of gases and aqueous vapour. In this conception we have the fundamental basis of the theory of ascension of solutions.

Theory of Lateral Secretion.

According to this theory it is assumed that meteoric waters percolating through the country rock, by the aid of carbondioride and alkalies dissolve out certain constituents, which are afterwards deposited in fissures and cavities.

The origin of the theory is unknown, but it is certain that Delius, in 1770, Gerharde, in 1781, and Lasius, in 1789, were supporters of it, the latter basing his argument upon a careful examination of the veins of the Hartz Mountains.*

In 1847 Professor Bischof, of Bonn, a distinguished geologist and chemist, in his fascinating “Text-book of Chemical and Physical Geology,” discusses the chemical processes which take place when meteoric waters and different kinds of aqueous

[Footnote] * Georg. Lasius, “Observations on the Hartz Mountains,” Hanover, 1789: “Ores and Minerals,” Vol. ii.

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solutions come in contact with rocks. His work created a new scientific basis of research in this branch of economic geology. He contended that ores were obtained by leaching from the rocks traversed by the veins, and suggested the possibility of the vein-constituents being found in the adjacent rocks.

In 1855 Forchhammer, the famous chemist of Copenhagen, found traces of lead, copper, and zinc in the roofing-slates of North Wales, a discovery which was held to afford conclusive proof of the origin of ore-veins by processes of lateral secretion.

In 1873 Professor F. Sandberger, of Wurzburg, dissatisfied with the meagre results obtained from the examination of sedimentary rocks, directed his attention to a systematic chemical investigation of the rocks traversed by ore-veins, and of the vein-stuff itself, in different mining centres in the Black Forest. In clay-slate he discovered copper, zinc, lead, arsenic, antimony, tin, cobalt, and nickel; in sandstone, lead and copper. Titanic and phosphoric acids were found to be present everywhere in small quantity.

Sandberger's results showed that a close relationship existed between vein-contents and the country rock; but he was by no means satisfied as to the origin of the heavy metals. He accordingly extended his investigation to an examination of the constituents of igneous rocks. He crushed large samples of rock and separated the constituent minerals by solutions of different densities. Large samples of the individual crystallized silicates thus isolated were subjected to careful analysis. In this way he found all the usual elements formed in metalliferous veins. Thus, in olivine he found iron, nickel, copper, and cobalt; in augite, copper, cobalt, iron, nickel, lead, tin, and zinc; and in the micas, many base metals. Gold, mercury, and tellurium were not sought for.

In 1880 Sandberger announced his belief that the mineral contents of veins were derived not from some unknown depth, but from the immediate wall-rock.

Gold-bearing veins are common in slates and sandstones of marine origin; and, as sea-water, according to the announcement of Sonstadt* in 1872, contains somewhat under a grain of gold to the ton, it is held by the exponents of lateral secretion that the sea is therefore the source of the gold in veins traversing marine sedimentaries. It is maintained that when sediments are deposited on the floor of the sea they must necessarily entangle a certain proportion of sea-water, and that when these sediments become consolidated the gold must remain in them.

[Footnote] * Confirmed by Professor Liversidge in 1893 and Sir William Ramsay, F.R.S., in 1905.

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The theory of lateral secretion received a new impulse from the researches of Sandberger. It seemed competent to explain the origin of many ore-veins, and, although strongly opposed by Professor Stelzner of Freiburg and Professor Posepny of Przibram, it found much support in America, in a more or less modified form.

Thus, Emmons,* discussing the manner in which he considers the Leadville ore-deposits were produced, summarises his views on ore-formation in general as follows:—

(1.)

Ore-deposits have been deposited from solution, rarely in open cavities, most frequently by metasomatic interchange.

(2.)

Solutions do not necessarily come directly upwards, but simply follow the easiest channels of approach.

(3.)

The material was derived from sources within limited and conceivable distance, very often the older intrusive rocks.

Emmons, while supporting the principle of lateral secretion, disclaims the narrow views of Sandberger, who limits the source of the vein-contents to the wall-rock in immediate contact with the vein.

In the critical discussion which followed the publication of Professor Posepny's paper on the genesis of ore-deposits, in 1893, Blake and Winslow reaffirmed their belief that the zinc and lead ores of Wisconsin were framed by lateral secretion.

Becker,§ while strongly dissenting from Posepny's view that metasomatic replacement was incapable of producing Such pronounced ore-bodies as those at Leadville, makes a clear statement of the supposed operation of metasomatic processes. He says, “Replacement, like solution, must occur along fissures or channels, and metasomatic ore-bodies will present analogies in form to the open spaces of caves of solution.”

Rickard discusses the problem of ore-formation from a wide standpoint, and is not a dogmatic supporter of the extreme doctrines of either ascension or lateral secretion. He affirms that there is no ground for the belief in the existence of a reservoir of water at great depth, and maintains that all ascending water must at one time have been descending water.

This last can only be true in regard to meteoric waters. So far as the existence of deep-seated water is concerned, his view

[Footnote] * S. F. Emmons, “The Genesis of certain Ore-deposits,” Trans. Am. Inst. M.E., Vol. xx, p. 125.

[Footnote] †“The Genesis of Ore-deposits,” New York, 1901, p. 199.

[Footnote] ‡ Loc. cit., p. 188.

[Footnote] § Loc. cit., p. 200.

[Footnote] ∥ Loc. cit., pp. 190 and 211.

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is not in accord with the hydro-fusion theory of modern petrologists.

Rickard, who possesses a personal knowledge of the goldfields of Australia and New Zealand, discusses the probable origin of the veins of the Thames Goldfield, in the latter country, and in the main agrees with Captain Hutton that they were formed by processes of lateral secretion by thermal waters.

Professor J. Le Conte* in a carefully prepared thesis combats the extreme views of both Posepny and Sandberger. He makes an earnest attempt to combine what is true in each, and reconcile their differences. It is manifest, however, that he leans favourably to the side of lateral-secretion processes, although not defined as such. He considers both sides right and both wrong. Ascensionists, he thinks, are right in deriving metals mainly by ascending solutions from great depths, but wrong in imagining these depths to be an exceptionally metalliferous barysphere, and wrong in not allowing subordinate contributions by lateral currents from the wall-rock higher up. The lateral-secretionists, on the other hand, are right, he thinks, in deriving metals by leaching from the wall-rock, but wrong in not making the thermosphere the main source.

Le Conte succinctly summarises his views in the following terms:—

(1.)

Ore-deposits, using the term in its widest sense, may take place from many kinds of waters, but especially from alkaline solutions; for these are the natural solvents of metallic sulphides, and metallic sulphides are usually the original form of such deposits.

(2.)

They may take place from waters at any temperature and pressure, but mainly from those at high temperature and under heavy pressure, because, on account of their great solvent power, such waters are heavily freighted with metals.

(3.)

The depositing waters may be moving in any direction—up-coming, horizontally moving, or even sometimes down-going—but mainly up-coming, because by losing heat and pressure at every step such waters are sure to deposit their contents abundantly.

(4.)

Deposits may take place in any kind of water-way—in open fissures, in incipient fissures, joints, cracks, and even in porous sandstones, but especially in great open fissures, because these are the main highways of ascending waters from the greatest depths.

[Footnote] * J. Le Conte, “The Genesis of Ore-deposits,” p. 270.

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(5.)

Deposits may be found in many regions and in many kinds of rocks, but mainly in mountain regions and in metamorphic and igneous rocks, because the thermosphere is nearer the surface, and ready access thereto through great fissures is found mostly in these regions and in these rocks.

Professor C. R. van Hise, in a classic paper on “Some Principles controlling Deposition of Ores,”* defines his views in the following sentences:—

(1.)

That the greater number of ore-deposits is the result of the work of underground water.

(2.)

That the material of ore-deposits is derived from rocks within the zone of fracture.

(3.)

That by far the major part of the water depositing ores is meteoric.

(4.)

That the flowage of water underground is caused chiefly by gravitative stress.

(5.)

That the waters which perform the first work in the genesis of ore-deposits are descending waters.

(6.)

Lateral secretion is therefore an essential step in the first concentration of ore-deposits.

(7.)

That sulphide ores are generally deposited by ascending waters in trunk channels.

(8.)

That the majority of ore-deposits, if not all, are partly deposited in pre-existing openings, and are partly replacements of wall-rocks.

It is manifest that he attaches too little importance to the genetic connection existing between ore-deposits and eruptive processes, and places too much dependence upon the formative power of meteoric waters.

Professor Kemp contends that mining operations in America show conclusively that mines become drier with increasing depth; and deep mining in South Africa, Australia, and New Zealand adds confirmation to this view. The dryness of mines in depth seems to destroy the foundations of Van Hise's main contention respecting the underground circulation of meteoric water.

“Van Hise admits that there are ore-deposits which have a direct igneous origin, but thinks they are or limited extent. In his rejoinder to Kemp he somewhat modifies his former concep-

[Footnote] * “The Genesis of Ore-deposits,” p. 282; also Trans. Amer. Inst. M E., Vol. xxx, 1901, p. 27.

[Footnote] † J. F. Kemp, “The Rôle of Igneous Rocks in the Formation of Veins,” “The Genesis of Ore-deposits,” 1901, p. 681.

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tion with respect to the action of meteoric waters, and admits that the rôle of igneous intrusions may be very considerable.*

It has been suggested by the opponents of lateral secretion that the metals contained in the silicate minerals of eruptive rocks are not primary but secondary constituents. According to their view, lateral secretion is only a process of concentration.

Ascension of Solutions.

According to this theory it is assumed that the material which fills a lode has been brought in solution from great depths, and not derived from the rocks in the immediate vicinity of the lode.

In his classic memoir on “The Genesis of Ore-deposits,” the late Professor Posepny, an ardent supporter of the ascension hypothesis, laid great stress upon the occurrence of sulphur and cinnabar at Sulphur Bank, California, impregnating a decomposed basalt, and still mildly in process of formation from gaseous emanations and hot mineral waters.

Similar conditions exist at Steamboat Springs, in Western Nevada, where we have an example of a mineral vein in process of formation. The matrix is banded siliceous veinstone, containing iron and copper sulphides, sulphur, and metallic gold.

Sandberger, who was an equally strenuous supporter of lateral secretion, objected to this view on the ground that he knew of no spring which deposited mineral incrustations on the walls of their channels. He regarded the Sulphur Bank and Steamboat Springs phenomena as exceptional.

Becker, who made a special examination of the deposits at Sulphur Bank and Steamboat Springs, strongly opposed the views of the extreme ascensionists. And with regard to the origin of the deposits he expressed the following views: “The evidence is overwhelming that the cinnabar, pyrite, and gold of the quicksilver-mines of the Pacific Slope reached their present positions in hot solutions of double sulphides which were leached out from masses underlying the granite, and the granite itself.” Further on he says, “I regard many of the gold-veins of California as having an origin entirely similar to that of the quicksilver-deposits.”

Becker's views postulate a new hypothesis lying midway between the ascension and lateral-secretion theories, and now

[Footnote] * C. R. van Hise, “The Genesis of Ore-deposits,” New York, 1901: Discussion, p. 763.

[Footnote] † G. F. Becker, “The Geology of the Quicksilver-deposits of the Pacific Slope,” U.S. Geol. Survey, Vol. xiii, 1888, p. 449.

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receive more general support than the extreme views of Posepny and Sandberger.

According to the definition of lateral secretion by the latter, the descending waters became charged with mineral matter by leaching the rocks in the region of vadose circulation. On the other hand, Posepny assumed that the ascending waters became charged at great depths by coming in contact with a deep-seated repository of metalliferous matter, the existence of which is necessarily hypothetical.

The modification suggested by Becker leans towards the ascension theory, and differs only from lateral secretion in assuming a deeper source for the mineral contents of the vein-matter.

Summary.

From the data recorded in the preceding papers we may deduce the following conclusions respecting the genesis of ore-deposits:—

(1.)

That the majority of ore-deposits are genetically connected with igneous rocks.

(2.)

That circulating underground water and gases are the principal agents concerned in the dissolution, primary concentration, and deposition of vein-matter.

(3.)

That ore-deposits do not necessarily occupy pre-existing fissures and cavities.

(4.)

That ore-deposits were in many cases formed by meta-somatic replacement.

(5.)

That vein-filling waters are ascending waters, in most cases probably of magmatic origin.

(6.)

That the mineral contents are derived from rocks contiguous to the zone of fracture or zone of metamorphism.

(7.)

That the principal agents of dissolution are water and gases aided by heat and pressure.

(8.)

That precipitation from the ascending waters takes place in orderly horizontal zones in accordance with the natural laws governing solution and precipitation.

(9.)

That secondary enrichment is in the majority of cases due to the migration of mineral contents from a higher to a lower level, through the agency of descending meteoric waters.

The theories of lateral secretion and ascension of solutions are based on the fundamental assumption that the mineral matter filling cavities was deposited from circulating waters. Their differences lie principally in the different conceptions as to the direction and operation of the circulating liquids.

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1.

The lateral-secretion theory supposes—

(a.)

That the filling of cavities was the work of descending meteoric waters;

(b.)

That the filling-matter was principally obtained from the adjacent rocks by a process of leaching by thermal waters.

2.

The ascension theory assumes—

(a.)

That the filling of veins was effected by deep-circulating waters, ascending through open or partially open fissures;

(b.)

That the mineral and metallic contents were derived from a deep-seated mineralised zone.

Many writers assume that ascending and descending waters are merely units in what may be termed a hydraulic circuit, the interchange being caused by gravitation assisted by capillary, and the difference of temperature of the ascending and descending waters. Much of the movement, it is claimed, is necessarily lateral and towards channels filled with ascending waters.

It was maintained by Professor Posepny and Dr. Raymond that descending waters were merely oxidizing, and incapable of depositing sulphides. This contention has, however, been successfully disproved by Emmons, Becker, Van Hise, and other American geologists, who have shown the existence of secondary sulphides both above and below water-level, or in what may be termed the zone of vadose circulation.

Professor Daubree always maintained that metallic sulphides could not be deposited without the agency of organic matter; but Skey, as far back as 1870, proved experimentally that from its solution of carbonate of soda, potash, and ammonia, gold is reduced by sulphides, but not from it a solutions in alkaline sulphides.* He found that 1 grain of iron-pyrites was able to reduce 8.5 grains of gold.

[Footnote] * W. Skey, Trans. N.Z. Inst., Vol. iii, 1870, p. 226.

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Art. VII.—Notes and Descriptions of Lepidoptera.

[Read before the Wellington Philosophical Society, 3rd October, 1906.]

The following notes on new and interesting forms are based mainly on material received through the kindness of Messrs. G. V. Hudson and A. Philpott, principally collected in the southern parts of the South Island. It is evident that the fauna is still far from being exhausted. I think that good results might be especially obtained by (1) searching for, and particularly breeding, the more minute forms, which require close observation, and (2) collecting at high altitudes earlier, and perhaps also later, in the year.

CaradrinidÆ.

Leucania temenaula, n. sp.

♂. 34-36 mm. Head and thorax greyish-ochreous mixed with brownish. AntennÆ serrate, fasciculated. Forewings elongate-triangular, costa almost straight or subsinuate, termen bowed, waved, oblique; greyish-ochreous, mixed with whitish-grey and brownish; subbasal, first, second, and subterminal lines obscurely pale or whitish, inconspicuous, more or less-partially dark-edged interiorly, sometimes with some black irroration; orbicular and reniform more or less distinctly whitish-edged and laterally dark-margined, orbicular round, reniform trapezoidal; claviform small, whitish, dark-edged, touching first line, sometimes obsolete; median line obsolete; three pale dots on costa posteriorly; a terminal series of dark-fuscous crescentic marks. Hindwings fuscous, with dark-fuscous terminal line; cilia whitish, base fuscous-tinged.

Rakaia (Fereday), Dunedin (Hudson); three specimens. A distinct though inconspicuous species, allied to moderata, bu [ unclear: ] lighter and less uniform in colouring, spots more distinct, termen rather more oblique, hindwings much less dark posteriorly.

Leucania pachyscia, n. sp.

♀. 33–40 mm. Head and thorax grey-whitish mixed with dark fuscous, thorax sometimes with blackish anterior angulated bar. Forewings elongate-triangular, costa almost straight, termen bowed, waved, oblique; whitish, sprinkled with brownish and black scales; subbasal line partially margined with dark

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fuscous; first and second lines indicated by dark-fuscous interior margins; spots indicated by partial dark-fuscous margins, orbicular roundish, reniform trapezoidal, claviform suboval; median shade more or less marked, formed by dark - fuscous suffusion; subterminal line indicated by well-marked thick dark-fuscous anterior margin; a terminal series of dark-fuscous crescentic marks: cilia whitish, with distinct bars of blackish irroration. Hindwings whitish-fuscous, with suffused fuscous terminal fascia; cilia whitish, base more or less infuscated.

Mount Arthur (4,700 ft.) and Lake Wakatipu, in January; two specimens. Easily known by the whitish ground-colour and strong dark præsubterminal shade.

HydriomenidÆ.

Chloroclystis halianthes, n. sp.

♂. 26–27 mm. Head, palpi, and thorax fuscous mixed with whitish and dark fuscous, shoulders partially suffused with red; palpi 2¼. AntennÆ biciliated with long fascicles. Forewings triangular, costa posteriorly arched, termen little bowed, oblique, subsinuate above tornus; pale fuscous mixed with white and dark fuscous, sometimes partially suffused with pale red, appearing purplish-tinged; normal fasciÆ formed of blackish irroration; median band broad, anterior edge hardly curved, below middle sometimes largely suffused with white and partially with pale red, posterior edge running from before ¾ of costa to tornus, very obtusely angulated below middle, followed by well-marked pale double second line; fifth and sixth fasciÆ little marked, below middle obsolete through contraction of area: cilia fuscous mixed with whitish, distinctly barred with dark fuscous. Hindwings with termen tolerably evenly rounded, hardly prominent on vein 3; pale-grey, towards dorsum sprinkled with black and white.

Lake Wakatipu (Hudson); two specimens. Of the New Zealand species of this genus that have fasciculate antennÆ in ♂, muscosata, antarctica, and bilineolata are green-tinged species, dryas a clear brown species, plinthina and aristias are red-tinged species: halianthes is really red-tinged, but appears purplish, and is allied to the two latter, but larger than either, darker, and easily known by the grey hindwings (in the other two whitish) and differently formed median band of forewings. Probably it varies considerably, like several of the others.

Asaphodes stephanitis, n. sp.

♂. 26–27 mm. Head and thorax yellow-ochreous, with a few blackish scales. Forewings triangular, costa gently arched, termen bowed, oblique, slightly waved; clear light yellow-

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ochreous; fasciÆ formed by dentate striÆ of blackish irroration, first and second separated by a suffused whitish line, second obsolete except on extremities; third reduced to a single curved stria, preceded by a white line, and followed by white suffusion above and below middle; fourth of three striÆ, posterior edge with acute short triangular projection in middle, followed by a strong white line; fifth of two striÆ, posterior sometimes very thick and suffused, followed by white subterminal line, sometimes partially obscured; sixth of one stria following subterminal line, terminated above by an oblique subapical suffusion; a blackish terminal line: cilia white, barred with blackish irroration. Hindwings light ochreous-yellow; terminal line and cilia as in forewings.

Invercargill, on sandhills (Philpott); two specimens received from Mr. Hudson. Recalls a small Xanthorhoe clarata, but the resemblance is only superficial.

Xanthorhoe dionysias, n. sp.

♂. 28 mm. Head, palpi, and thorax pale-ochreous tinged with brown-reddish. Forewings somewhat elongate-triangular, costa gently arched, subsinuate in middle, termen rather bowed, oblique, not waved; pale greyish-ochreous, towards costa suffusedly tinged with reddish-ochreous; basal area indistinctly striated with dark fuscous irroration; median band defined anteriorly by two curved similar striÆ, posteriorly by three curved dark striÆ enclosing two lines, first pale, second slightly tinged with reddish-ochreous; within median band are two suffused striÆ connected by a transverse dark-fuscous discal dot, first obsolete in middle; terminal area irrorated with dark fuscous; an interrupted dark-fuscous terminal line: cilia pale-greyish-ochreous, barred with dark-fuscous irroration. Hind-wings elongate, termen rounded, faintly waved; pale grayish [ unclear: ] ochreous, thinly irrorated with grey; a blackish discal dot; a cloudy grey postmedian line; cilia pale greyish-ochreous mixed with grey.

Old Man Range, Dunedin (Lewis); one specimen received from Mr. Hudson. A distinct species, probably allied to aegrota, but not very nearly.

CrambidÆ.

Crambus apselias, n. sp.

♂ 28–30 mm., ♀ 23 mm. Head ochreous - white, face rounded. Palpi 4 ½-5, brownish-ochreous, internally and at base beneath, white, in ♀ very slender. AntennÆ in ♂ filiform, shortly ciliated. Thorax brownish-ochreous, with ochreous-white central stripe. Abdomen whitish-ochreous. Forewings

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very elongate-triangular, costa posteriorly moderately arched, apex round-pointed, termen sinuate, oblique; shining brassy-ochreous; a moderate white longitudinal median streak from base, split into three branches posteriorly, and two similar detached branches above these; very indistinct narrow streaks of whitish suffusion towards costa between veins, in ♀ more suffused; two or three broader and sometimes confluent streaks of white suffusion towards dorsum between veins: cilia in ♂ whitish, on lower half of termen more or less suffused with ochreous, in ♀ white. Hindwings fuscous-whitish, in ♂ more fuscous-tinged towards apex; cilia in ♂ ochreous-whitish, in ♀ white.

Invercargill, in December; two specimens taken by myself, and three others received from Messrs. Philpott and Hudson. Closely allied to ramosellus, from which it is very easily separated by the entire absence of the black terminal dots and of the blackish inferior edging of the white median streak.

Crambus conopias, n. sp.

♂. 26 mm. Head white, with a central ochreous line, face forming a moderate conical projection. Palpi 5, brownish-ochreous, white at base and internally. AntennÆ filiform, simple. Thorax light ochreous, with suffused whitish dorsal stripe. Abdomen whitish-ochreous. Forewings very elongate, narrow, somewhat dilated posteriorly, costa slightly arched, apex tolerably acute, termen hardly sinuate, oblique; pale brownish-ochreous; a moderate white longitudinal median streak from base to termen, posterior extremity angularly produced upwards on termen, margined with blackish suffusion above from ⅓ to near termen, and beneath almost from base to middle; veins posteriorly marked with some dark-fuscous scales; median and second lines indicated by angulated series of ill-defined dark-fuscous longitudinal marks on veins, obsolete near costa; terminal blackish dots on veins: cilia white, somewhat mixed with brownish-ochreous. Hindwings fuscous-whitish, more infuscated posteriorly; cilia white.

Dunedin (Lewis); one specimen received from Mr. Hudson. Apparently more allied to ramosellus than to any other New Zealand species, but very distinct by the frontal cone; there would seem to be undoubted affinity to the European inquinatellus.

Tauroscopa glaucophanes, n. sp.

♀. 29–31 mm. Head and palpi ochreous-whitish mixed with blackish, palpi 3 ½; a naked dark-grey space between eye and palpi. Thorax dark grey suffused with light bronzy-

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blue-greenish, and mixed with ochreous-whitish and pale ochreous. Forewings elongate, moderate, slightly dilated, costa gently arched anteriorly, almost straight posteriorly, apex obtuse, termen rounded, somewhat oblique; dark grey, more or leas mixed suffusedly with ochreous-whitish or whitish-ochreous, with a brassy-blue-greenish tinge; first and second lines pale, angulated, margined with darker shades, but sometimes almost wholly obsolete: cilia grey mixed with ochreous-whitish. Hindwings grey, lighter towards base, suffused with darker towards termen; cilia grey, tips whitish.

Lake Wakatipu, 4,000–5,000 ft. (Hudson); two specimens. Larger than the other two species, with the forewings much less triangular, rather oblong, and otherwise quite distinct.

PyraustidÆ.

Scoparia xysmatias, n. sp.

♂. 19 mm. Head dark fuscous. Palpi 2 ½, dark fuscous, lower longitudinal half ochreous-whitish. AntennÆ very minutely pubescent. Thorax dark fuscous, sprinkled with ferruginous scales. Forewings elongate, gradually dilated, costa almost straight, apex obtuse, termen faintly sinuate, rather oblique; dark fuscous, irregularly strewn except on margins with yellowish-ferruginous scales, and partially suffused with black, especially about margins of first and second lines and towards dorsum anteriorly; first and second lines represented by straight undefined series of whitish scales surrounded with yellowish-ferruginous suffusion, strongly converging towards dorsum; a small spot of white scales in middle of disc, followed by a black spot; a well-marked black fascia beyond second line; a few white scales indicating subterminal line: cilia ochreous-whitish, with dark-fuscous basal and grey median lines. Hindwings without hairs in cell; dark grey sprinkled with blackish; cilia ochreous-whitish, with dark-grey basal line.

Old Man Range, Dunedin (Lewis); one specimen received from Mr. Hudson. Probably intermediate between hemicycla and ergatis, but very distinct.

Scoparia autochroa, n. sp.

♂. 21–23 mm. Head and thorax brown, face dark fuscous. Palpi 2⅔, pale brownish mixed with dark fuscous, base whitish. AntennÆ dark fuscous, ciliations ⅔. Abdomen fuscous. Forewings elongate-triangular, costa hardly arched, apex obtuse, termen little rounded, somewhat oblique; brown sprinkled with rather dark fuscous: cilia pale brownish, with rather

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dark fuscous subbasal line. Hindwings without hairs in cell; fuscous; cilia light greyish-ochreous, with fuscous subbasal line.

Invercargill, in open swampy situations, in November (Philpott, Hudson); three specimens. Very distinct, of somewhat doubtful affinity, possibly nearest encapna.

Scoparia choristis, n. sp.

♂. 17 mm. Head and thorax fuscous, somewhat whitish-sprinkled. Palpi 2¼, fuscous, basal area white beneath, edged with dark fuscous. Antennal ciliations ½. Forewings elongate-triangular, costa posteriorly moderately arched, termen slightly sinuate, rather oblique; white, thinly sprinkled with fuscous; base suffused with fuscous, with a few black scales, and a short very oblique black streak from base of costa; first line white, undefined anteriorly, posteriorly edged by a black streak from ⅓ of costa to ⅖ of dorsum, rather curved near costa, followed by an undefined band of fuscous suffusion; discal spot 8-shaped, faintly outlined with fuscous; second line slender, white, from beyond ⅔ of costa to dorsum near tornus, edged anteriorly with fuscous irrorated with black, subsinuate inwards near costa and outwards near dorsum and very obtusely angulated above middle; terminal area fuscous mixed with blackish, subtermmal line represented by some whitish suffusion towards apex, and an oblique suffused white bar below middle; a terminal series of cloudy whitish dots. Hindwings without hairs in cell; light grey with brassy reflections, paler towards base; cilia whitish, with two pale-grey shades.

Kaitoke, Wellington, in November (Hudson); one specimen. Very similar to colpota, but immediately distinguished by quite different form of second line, especially by the peculiar outward sinuation near dorsum; also allied to periphanes, in which, however, the lower half of second line is straight and oblique.

Scoparia asaleuta, n. sp.

♀. 21–23 mm. Head, palpi, and thorax prismatic bronzy-grey, suffusedly irrorated with white; palpi 2¼. AntennÆ dark grey, suffusedly ringed with white. Abdomen pale ochreous, sprinkled with grey. Forewings very elongate, narrow, posteriorly dilated, costa subsinuate in middle, slightly arched posteriorly, apex obtuse, termen nearly straight, hardly oblique, rounded beneath; iridescent pale ochreous mixed with dark bluish-grey, wholly suffused or densely irrorated with white except dark markings as under—viz., an undefined subbasal fascia; a nearly direct fascia representing first line, dilated posteriorly above middle; an 8-shaped discal mark; a broad

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terminal fascia, on which the second and subterminal lines appear as whitish shades confluent in middle and sometimes partially obsolete: cilia whitish-ochreous, tips whitish, with narrow basal and broader postmedian grey shades. Hindwings without hairs in cell; pale greyish-ochreous, with suffused dark-grey terminal fascia; cilia ochreous-grey-whitish, with grey basal line.

Lake Wakatipu (Hudson); two specimens. An elegant species, intermediate between cataxesta and tetracyla; to the naked eye obviously bluish-tinged.

Scoparia augastis, n. sp.

♂. 28–29 mm. Head fuscous, crown mixed with whitish. Palpi 3, fuscous, base white. AntennÆ fuscous, ciliations ½. Thorax light fuscous. Abdomen pale greyish-ochreous. Forewings very elongate, gradually dilated, costa posteriorly gently arched, apex obtuse, termen faintly sinuate, rather oblique; rather light fuscous, more or less irrorated finely with whitish; costa and all veins marked by more or less distinct somewhat darker fuscous lines: cilia whitish, with two fuscous lines, anterior interrupted. Hindwings with long hairs in cell; very pale brassy-ochreous; cilia whitish, with very faint greyish subbasal line.

Invercargill, in March, on flowers of Senecio after dark (Philpott); three specimens. Very distinct, perhaps nearest to the Australian nephelitis.

PterophoridÆ.

Platyptilia campsiptera, n. sp.

♂. 17 mm. Head and thorax yellowish-white, metapleura with an oblique black streak, frontal cone of scales moderately long. Palpi 3 ½, white, apical ⅔ externally irrorated with dark fuscous. Abdomen pale whitish-yellow, with a black lateral dot near base, and a few black lateral scales posteriorly. Legs white, banded with dark fuscous. Forewings cleft from ¾, upper segment rather narrow, apex produced, pointed, lower segment much broader, posteriorly dilated; whitish, tinged with pale yellow; costa towards base shortly strigulated with fuscous irroration; a small triangular fuscous spot irrorated with dark fuscous on costa before fissure, not reaching across first segment, and a smaller similar mark on costa between this and apex: cilia ochreous-whitish, spotted with fuscous round lower angle of first segment, and upper angle and termen of second, with a small black scale-tooth on dorsum at ⅔. Hindwings reddish-fuscous; cilia whitish, slightly reddish-tinged, with a blackish basal mark on lower half of termen of first segment, and mere traces of black scales in middle of dorsum of third segment.

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Humboldt Range, Lake Wakatipu, at 3,600 ft. (Hudson). Mr. Hudson writes: “In this species the second digit of the forewing is held almost at right angles pointing downwards from the first digit during life; after death the digit assumes the normal position.” This would appear to be a very singular characteristic, which would repay further investigation. The species is allied to deprivatalis, but very distinct.

Platyptilia deprivatalis, Walk. 946.

This name supersedes Haasti, Feld.; I have examined the types.

TortricidÆ.

Epagoge, Hb.

This genus, which differs from Capua only by the absence of the costal fold in ♂, has not been hitherto known from New Zealand, but is well represented in Australia.

Epagoge cyclobathra, n. sp.

♂ ♀. 16–18 mm. Head, palpi, and thorax rather dark fuscous. Abdomen pale grey, Forewings elongate, suboblong, costa moderately arched, apex obtuse, termen hardly rounded, rather oblique; fuscous, slightly purplish-binged, sprinkled with dark fuscous, towards middle of costa suffused with dark ashy-fuscous, towards termen mixed with reddish-ochreous and strigulated with dark fuscous; basal ⅖ whitish-ochreous, marked with several deeper ochreous striÆ, outer edge curved; within this patch an irregular dark-fuscous streak from base of dorsum along costa to ¼, thence proceeding as a curved transverse streak to sub-median fold: cilia grey, with dark fuscous median line. Hind wings light grey; cilia whitish-grey, with darker subbasal line.

Invercargill, in December and January (Philpott); two specimens.

Trachybathra, n. g.

AntennÆ in ♂ moderately biciliated. Palpi moderate, porrected, rough-scaled above and beneath. Thorax smooth. Forewings with rough scales at base, in ♂ with costal fold; 2 from ⅔, 7 and 8 stalked, 7 to termen, 11 from middle. Hindwings with vein 4 absent, 6 and 7 stalked.

Allied to Capua, from which it differs by the rough basal scales of forewings, and the absence of vein 4 of hindwings.

Trachybathra scoliastis, n. sp.

♂. 18 mm. Head, palpi, and thorax brownish irrorated with grey-whitish and dark fuscous. Abdomen fuscous. Forewings

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elongate, hardly dilated, costa slightly bent before middle, apex obtuse, termen faintly sinuate beneath apex, bowed, oblique; pale brownish, partially suffused irregularly with whitish, costa and dorsum strigulated with dark fuscous; outer edge of basal patch indicated by a blackish line in disc, obsolete towards extremities; an irregular incurved fuscous streak marked with black from ⅖ of costa to below middle of disc, followed by whitish suffusion; an irregular dark-fuscous spot above tornus, and some dark-fuscous strigulÆ towards lower part of termen: cilia grey-whitish mixed with dark grey. Hindwings fuscous, strigulated with darker; some undefined ochreous-yellowish suffusion in centre of disc and towards costa in middle; cilia pale grey, with dark-grey subbasal line.

Lake Wakatipu (Hudson); one specimen.

Pyrgotis tornota, n. sp.

♂. 18 mm. Head and thorax pale ochreous sprinkled with dark ferruginous. Palpi 2½, whitish-ochreous, irrorated wit dark ferruginous. Abdomen pale whitish-ochreous. Forewings elongate, posteriorly somewhat dilated, costa strongly arched towards base, faintly sinuate in middle, posteriorly almost straight, apex obtuse, termen concave, little oblique; reddish-brown, suffusedly mixed with pale leaden-grey, and strigulated with whitish-yellowish; a few minute dots of blackish scales, especially on dorsum; a moderately broad curved darker streak, suffusedly irrorated with yellow-ochreous, from ⅓ of costa to below middle of disc, thence curved evenly upwards, confluent towards costa with a round similar blotch on middle of termen, so that the ground-colour appears to form a narrow whitish-edged projection upwards between them: cilia reddish-brown, darker towards apex, tips whitish-ochreous. Hindwings whitish, with a few grey strigulÆ on dorsal half; cilia whitish.

Invercargill, in August (Philpott); one specimen. The whitish hindwings are a special characteristic.

Adoxophyes conditana, Walk.

I have received from Mr. Philpott three examples of a curious melanic form of the male of this-species, stated to be common near Invercargill. The forewings are mainly suffused with dark purplish-grey irrorated or strigulated with blackish, except some small variable whitish spots towards middle of dorsum and sometimes towards apex and termen; whilst the hindwings are densely irrorated with blackish. I at first thought it a distinct species, but after careful comparison with my long series of varieties of this extraordinarily variable species am satisfied that it is only a melanic southern form; I have not

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however, yet seen the corresponding females. Several other species from Invercargill show the same tendency to melanism, which should be borne in mind when considering insects from that region.

Cacoecia acrocausta, n. sp.

♂. 19–21 mm. Head and thorax brownish-ochreous or yellow-ochreous. Palpi. 3, fuscous, externally suffused with ferruginous. Antennal ciliations 1 ½. Abdomen whitish-ochreous, beneath ferruginous, and tuft mixed with dark grey. Forewings elongate-triangular, costa slightly arched, apex obtuse, termen rounded, little oblique; whitish-ochreous or yellow-ochreous, with scattered blackish-grey strigulÆ, basal ¾ more or less tinged or suffused with brown; costal edge ferruginous: cilia whitish-ochreous, on upper half of termen dark grey, on costa yellowish-ferruginous. Hindwings ochreous-whitish, strigulated with pale grey, more distinctly towards base; cilia ochreous-whitish.

♂. 22 mm. Head and thorax whitish. Palpi 3 ½, ochreous-whitish, externally fuscous-sprinkled. Forewings more elongate than in ♂, ochreous-whitish, sprinkled with very pale fuscous; central fascia indicated by an undefined grey very zigzag shade; a small grey spot towards termen in middle: cilia whitish-ochreous, becoming fuscous on upper part of termen. Hindwings as in ♂.

I took a male and female together at Christchurch in September, but had not ventured hitherto to describe them; I have, now received four additional males from Mr. Philpott, taken at Invercargill, where the species is common in October and November. It is allied to excessana, but quite distinct.

Cacoecia orthropis, Meyr.

Examples from Invercargill sent by Mr. Philpott have the forewings much greyer than Christchurch and Wellington specimens, and the hindwings are also grey; they appear to constitute a geographical form only, and to afford an instance of the tendency to a darker colouring mentioned above.

Tortrix molybditis, n. sp.

♂. 12 mm. Head and palpi rather dark fuscous, palpi 2. Antennal ciliation 1. Thorax dark glossy leaden-fuscous. Abdomen dark fuscous. Forewings elongate, posteriorly dilated, costa gently arched, apex obtuse, termen straight, rather oblique; rather dark glossy leaden-grey; markings blackish-fuscous; four small spots on costa alternating with principal markings; a stria marking outer edge of basal patch, strongly angulated in

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middle; a small spot of pale - yellowish projecting scales on dorsum neat base; upper half of central fascia well marked, lower half obsolete; a small triangular costal patch, from near which proceed two irregular striae to tornus and lower part of termen, edged with a few pale-yellowish scales: cilia dark fuscous, tips paler. Hindwings dark fuscous; cilia pale - greyish, with dark-grey basal line.

Wellington (Hudson); one specimen. Very distinct; has some superficial resemblance to Dipterina hemiclista, but easily distinguished by the short antennal ciliations.

Eurythecta paraloxa, n. sp.

♂. 12–13 mm. Head and thorax whitish-ochreous, more or less suffused with ferruginous, with a few black scales. Abdomen dark fuscous, apex pale ochreous. Forewings elongate, rather narrow, costa somewhat arched anteriorly, apex obtuse, termen somewhat rounded, oblique; vein 7 present; ochreous-whitish, more or less suffused with yellow-ochreous, palest on costa; markings yellow-ochreous or ferruginous, variably mixed with dark fuscous; basal patch darkest towards outer edge, which is oblique, rounded - prominent in middle, more or less-excavated beneath this; central fascia moderate, anterior edge nearly straight, posterior edge irregularly prominent below middle, so as to appear concave on upper half; four small spots on costa posteriorly, first three sometimes confluent into a small triangular blotch; an irregular blotch along termen: cilia ochreous-whitish, more or less distinctly barred with dark fuscous. Hindwings dark grey; cilia grey.

Invercargill, common on sandhills in January (Philpott); three specimens. This species differs from the other two in having vein 7 of forewings present; but as it possesses the other characteristic structural points of the genus, and is obviously nearly allied in all respects, it seems unnecessary to form a new genus for its reception. The genus is a development of Proselena, and the present species is an early form of it.

Prothelymna niphostrota, n. sp.

♂. 15 mm. Head whitish. Palpi, thorax, and abdomen pale fuscous. Forewings elongate, costa moderately arched, apex obtuse, termen slightly rounded, oblique; white; basal patch pale fuscous, marked with spots of blackish irroration; dorsal half from this to tornus marked with coarse grey strigulÆ irrorated with black; an oblique grey patch on middle of costa; a smaller dark-grey spot on costa at ¾; a grey apical patch, marked with coarse blackish-grey strigulÆ, and extended as an irregular subterminal stria to tornus: cilia whitish, round

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apex greyish-tinged and spotted with blackish irroration. Hindwings pale grey, veins partially dark grey; cilia grey-whitish.

Invercargill, in January (Philpott); one specimen. This, the second known species of the genus, is easily known from the other by the white ground.

PhaloniadÆ.

Heterocrossa iophaea, n. sp.

♀. 18–19 mm. Head, palpi, and thorax dark fuscous irrorated with whitish, face and palpi internally pale ochreous, palpi 4. Abdomen grey, two basal segments whitish-ochreous. Forewings elongate, narrow, costa gently arched, apex round-pointed, termen almost straight, oblique; dark fuscous irrorated with whitish, sometimes more or less mixed with pale ochreous; a series of small dark spots along costa; tufts brownish-ochreous suffusedly edged with black and posteriorly margined with whitish, viz., two near base sometimes surrounded with ochreous suffusion, a transverse angulated series beyond ¼, and five arranged round middle of disc, enclosed space sometimes blackish; a more or less defined angulated dark subterminal line: cilia rather dark fuscous irrorated with whitish. Hind-wings grey: cilia whitish-grey.

Invercargill, amongst bush, from October to February (Philpott); four specimens. Much the darkest species of the genus.

Heterocrossa gonosemana, Meyr.

Further material has convinced me that epomiana, Meyr., is only a variety of this species, and must sink accordingly.

GelechiadÆ.

Gelechia acrodactyla, n. sp.

♂. 15–17 mm. Head whitish-ochreous. Palpi ochreous-whitish; basal joint, lower third of second joint, and subapical ring of terminal joint dark fuscous. AntennÆ serrulate, pubescent, pale ochreous dotted with dark fuscous. Thorax whitish-ochreous, tinged or irrorated with brownish. Abdomen grey. Forewings elongate, narrow, costa gently arched, apex obtuse, termen rounded, rather strongly oblique; whitish-ochreous, irregularly irrorated with brown; plical and first discal stigmata rather large, blackish, plical rather before first discal; brown irroration forms a suffused costal patch beyond middle, and a narrow terminal fascia; cilia whitish-ochreous, with dark-grey subbasal line. Hindwings over 1, grey; cilia pale whitish-ochreous, with grey subbasal line.

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Invercargill, amongst bush, local, in November (Philpott); three specimens. Allied to achyrota, but, apart from other characters, achyrota has two blackish rings on terminal joint of palpi, acrodactyla only one.

œcophoridÆ.

Borkhausenia armigerella, Walk.

Specimens sent by Mr. Philpott from Invercargill, in conjunction with the material already possessed, appear to show conclusively that actinias, Meyr., is only a strongly marked form of this species, and the name should therefore be reduced to a synonym.

Borkhausenia perichlora, n. sp.

♂. 21–22 mm. Head whitish-ochreous mixed with fuscous. Palpi whitish-ochreous, irrorated throughout with rather dark fuscous. AntennÆ dark fuscous. Thorax dark fuscous, apical half of patagia yellow-ochreous. Abdomen grey. Forewings elongate, costa gently arched, apex round-pointed, termen very obliquely rounded; ferruginous-brown, towards costa somewhat paler and more ochreous; a yellow-ochreous streak along dorsum from base to near tornus posteriorly whitish, upper edge triangularly indented before middle, with some blackish scales in indentation: cilia ferruginous-ochreous. Hindwings light grey, margins narrowly whitish; cilia whitish.

Invercargill, abundant among Leptospermum, in November and December (Philpott); three specimens. At first sight very similar to basella, but with termen of forewings more oblique, and easily distinguished by the fuscous-mixed head, uniformly infuscated palpi, and whitish cilia of hindwings.

Borkhausenia basella, Walk. (Incurvaria basella, Walk. 492; œcophora ademptella, ib. 698.)

♂. 18–21 mm. Head ochreous-yellow. Palpi ochreous-yellow, second joint except apex, and subbasal and subapical rings of terminal joint dark fuscous. AntennÆ dark fuscous. Thorax dark brown, apex of patagia and a posterior spot ochreous-yellow. Abdomen fuscous. Forewings elongate, costa moderately arched, apex obtuse, termen almost straight, oblique; ochreous-brown irrorated with darker, with a slight purplish gloss; costal edge finely ochreous-yellow except towards base; a whitish-yellowish streak, partly suffused with deep ochreous-yellow, along dorsum from base to ¾, upper edge broadly triangularly indented before middle, with a blackish dot in in-

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dentation: cilia brown, irrorated with darker. Hindwings and cilia fuscous-grey.

Wellington (Hudson); six specimens. Distinguished by the uniform dark colouring and pale dorsal stripe; phegophylla, Meyr., is very similar, but much brighter-coloured, and has terminal joint of palpi wholly yellow, without dark fuscous rings; politis, Meyr. (of which I have both sexes) is much more mottled in appearance, with dark stigmata and subterminal line always apparent, and (though occurring in the same neighbourhood) truly distinct, since the antennal ciliations of ♂ are appreciably longer, obviously over 1, whilst in basella they are only 1.

Borkhausenia pronephela, n. sp.

♂. 16–17 mm. Head and thorax ochreous-yellowish. Palpi light yellowish, lower half of second joint dark fuscous. AntennÆ dark fuscous. Abdomen grey. Forewings elongate, rather narrow, costa moderately arched, apex obtuse, termen rounded, rather strongly oblique; yellow-ochreous; a pale yellow dorsal streak from base to near tornus; basal third of wing suffused with brown or dark fuscous except on dorsal streak which is indented by the dark colouring before middle, indentation partially whitish-edged and containing a blackish mark; two ill-defined fasciÆ of fuscous suffusion from middle of dorsal streak and tornus respectively, meeting on middle of costa; stigmata cloudy, dark fuscous, plical beneath first discal; some dark fuscous scales indicating an angulated subterminal line: cilia yellow-ochreous mixed with fuscous, with a fuscous postmedian shade. Hindwings grey; cilia whitish-grey.

Invercargill, on outskirts of bush, in December (Philpott); three specimens. A distinct species, intermediate between the basella and griseata groups.

PlutellidÆ.

Glyphipteryx metasticta, n. sp.

♂ ♀. 11–12 mm. Head and thorax bronzy - fuscous sprinkled with dark fuscous. Palpi ochreous-whitish, with two rings on second joint, and two rings and an anterior streak on apical portion of terminal joint blackish. AntennÆ dark fuscous. Abdomen fuscous. Forewings elongate, rather narrow, costa moderately arched, apex round-pointed, termen very obliquely rounded; 7 and 8 stalked; in ♂ ochreous-fuscous, sometimes partly suffused with bronzy-ochreous, in ♀ dark fuscous with a cloudy whitish streak along dorsum

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and posterior half obscurely marked with undefined whitish strigulÆ, partly edged with blackish; second discal stigma round, blackish, sometimes with a smaller similar dot before and above it; two or three short whitish strigulÆ from costa posteriorly, in ♂ very undefined, in ♀ longer and blackish-edged: cilia fuscous, on termen white with a blackish basal line. Hindwings and cilia rather dark grey, darker in ♀.

Invercargill, in damp situations on sandhills, in October (Philpott); three specimens. A distinct species, allied to the following.

Glyphipteryx aulogramma, n. sp.

♀. 15–16 mm. Head and thorax light shining bronzy-ochreous. Palpi ochreous-whitish, second joint with two black subapical rings, terminal joint with black anterior line. AntennÆ and abdomen dark fuscous. Forewings elongate, narrow, costa gently arched, apex acute, termen faintly sinuate, very oblique; 7 and 8 stalked; light shining bronzy-ochreous; costal edge whitish towards middle; a whitish-ochreous streak along submedian fold from ¼, variably extended posteriorly, and sometimes undefined whitish streaks on veins posteriorly, the interspaces sometimes fuscous-tinged; five silvery-grey-whitish fuscous-edged diversely oblique streaks from posterior half of costa, and two from about tornus, first costal connected with first tornal by some black scales, second tornal partly black-edged and subconfluent with fourth costal in a straight line: cilia ochreous-whitish. Hindwings dark grey, posteriorly blackish-irrorated; cilia grey, round costa and apex grey-whitish.

Invercargill, in damp situations on sandhills, in October (Philpott); two specimens. Allied to transversella, Walk., but a much less brilliant insect.

Simaethis exocha, n. sp.

♂. 20 mm. Head, palpi, and thorax grey-whitish mixed with blackish. AntennÆ pale grey, ringed with blackish. Abdomen fuscous, segmental margins mixed with whitish. Fore-wings elongate, moderate, posteriorly rather dilated, costa gently arched, apex obtuse, termen faintly sinuate, oblique; 7 and 8 connate; olive-fuscous, costa and dorsum broadly suffused with dark fuscous; basal area irrorated with whitish except a narrow fascia preceding first line; first line whitish, acutely angulated near costa, followed by a very irregular fascia of whitish irroration, which sends a triangular projection above middle to centre of disc; second line white, sharply defined, running from middle of costa to ¾ of disc, thence acutely angulated to beyond middle of dorsum, somewhat sinuate inwards towards

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costa and dorsum; an evenly broad fascia of white irroration from ⅘ of costa to ⅘ of dorsum, resting on second line in discal portion, terminating in a white spot on costa, and edged with a white line from this to angle of second line: cilia grey mixed with whitish, and indistinctly barred with dark fuscous irroration. Hindwings grey, darker posteriorly; indications of a cloudy whitish dot towards termen below middle; cilia grey mixed with whitish, with dark grey basal line.

Humboldt Range, Lake Wakatipu, at 3,600 ft. (Hudson); one specimen. This large and interesting form is allied to combinatana, Walk., but very distinct.

Art. VIII.—The Increase of Nitrogen in certain Soils due to Nitroculture.

[Read before the Philosophical Institute of Canterbury, 14th November, 1906.]

This paper is the record of an attempt to determine what increase of nitrogen takes place in soils which have grown a leguminous crop, with and without the aid of nitro-culture, and with and without the addition of sulphate of potash and superphosphate.

  • The nitro-culture was obtained from the United States Department of Agriculture, the experiments being carried out with the garden-pea grown on various soils.

  • Three soils were experimented with about 1,500 grams of soil being used in each pot.

The following are the conditions under which each soil was treated and cultivated:—

  • Experiment 1. The crop grown without fertiliser or nitro-culture.

  • Experiment 2. The crop grown with 1 gram sulphate of potash added to the soil.

  • Experiment 3. The crop grown with 1 gram sulphate of potash and 1 gram superphosphate added to the soil.

  • Experiment 4. The crop grown with the seed inoculated with nitro-culture.

  • Experiment 5. The crop grown with the soil inoculated with nitro-culture.

  • Experiment 6. The crop grown with the seed inoculated with nitro-culture and 1 gram sulphate of potash added to the soil.

  • Experiment 7. The crop grown with the seed inoculated with nitro-culture and 1 gram of sulphate of potash and 1 gram superphosphate added to the soil.

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The following tables show the increase or decrease of nitrogen in each soil when the various experiments were completed. The original percentage of nitrogen in each soil is shown at the top of each table. All determinations are calculated to the water-free sample.

Table I.
Soil A.
Original N. = .188 per cent.
Experiment No. Percentage of Nitrogen. Increase or Decrease.
1 .175 —.013
2 .180 —.008
3 .190 + .002
4 .212 + .024
5 .211 + .023
6 .218 + .030
7 .222 + .034
Table II.
Soil B (peaty).
Original N. = .332 per cent.
Experiment No. Percentage of Nitrogen. Increase or Decrease.
1 .334 + .002
2 .336 + .004
3 .340 + .008
4 .360 + .028
5 .392 + .060
6 .367 + .035
7 .381 + .049
Table III.
Soil C.
Original N. = .088 per cent.
Experiment No. Percentage of Nitrogen. Increase or Decrease.
1 .081 —.007
2 .082 —.006
3 .092 + .004
4 .113 + .025
5 .114 + .026
6 .116 + .028
7 .120 + .032
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Table IV.
Soil C + 10 grams Starch.
Original N. = .088 per cent.
Experiment No. Percentage of Nitrogen. Increase or Decrease.
4 .121 + .033
5 .146 + .058
6 .135 + .047
7 .142 + .054

From these experiments it would appear that there is a decided increase of nitrogen when the seed is inoculated with the nitro-culture; further, the increase is greater when sulphate of potash and superphosphate are added to the soil.

Except in the case of peaty soil (Table II), and when combined carbon was added (Table IV), there does not appear to be any special increase of nitrogen by inoculating the soil; and from the results presented in Table II, experiment 5, it is probable that the cellulose in the soil-humus is utilised to promote the growth of the organism and the increase of nitrogen.

The experiments presented here indicate that to a certain extent nitro-culture can be utilised to assist the replacement of the nitrogen removed by crops from the soil.

For permission to publish this paper I have to express my thanks to the Management of the Christchurch Meat Company (Limited), in whose laboratory most of the work was carried out.

Art. IX.—Notes on New Zealand Echinoderms; with Description of a New Species.

[Read before the Wellington Philosophical Society, 1st August, 1906.]

These notes will serve to clear up some points which were doubtful at the time when my “List of New Zealand Echinoderms” was published (Proc. Linn. Soc. N.S.W., 1898, p. 305); to describe a new ophiuran discovered by my colleague Mr. Arthur Haylock, who has made a fine collection of New Zealand echinoderms; and to correct several omissions in the “Index FaunÆ NovÆ-ZealandiÆ.”

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Although it is stated in the preface to the “Index” that I revised the Echinoidea, star-fishes, and Hydrozoa, I am not entirely responsible for the lists of these groups. When the late Captain Hutton was compiling the work, in reply to his request I sent copies of my papers to him, and some notes on the nomenclature of several species, and I offered to revise the lists when compiled, but I did not see them until the book was published.

Ophiuroidea.

Ophionereis schayeri.

This species, which is abundant near Wellington, Nelson, and Auckland, is very widely spread. Professor Kirk, of Victoria College, found it at the Chatham Islands; Mr. Haylock obtained specimens at the East Cape and at the Kermadec Islands; Dr. Ludwig has determined specimens from Juan Fernandez as our species (Zool. Record, 1898, Ech., p. 67); this or a very nearly allied form was described by Mr. E. A. Smith in the Proceedings of the Zoological Society, 1877, p. 92, from the Galapagos Islands, under the name Ophionereis albomaculata; and it occurs freely on the eastern coast of Australia. This is one of the two New Zealand littoral ophiurans which extend northwards into the Indo-Pacific region, the other being Ophiomyxa australis. I have found young specimens on the roots and stems of seaweed (Lessonia and Macrocystis), and it probably came to New Zealand across the Tasman Sea on floating seaweed. The southern branch of the Pacific Equatorial Current, which strikes the north-eastern coast of Australia and is diverted thence southwards and eastwards across the Tasman Sea, assisted by the prevailing westerly winds, has apparently been the means of adding many species to our littoral marine fauna. Our hydroid fauna especially shows the effect of this commingling of Australian forms with those which are peculiarly New-Zealandian.

The distribution of this sea-star appears to indicate a very great amount of vitality. From New Zealand (probably via the Chatham Islands) it has spread right across the South Pacific Ocean and up the western coast of South America in the line of distribution of the great sea-tangle of the Southern Ocean, Macrocystis pyrifera, a line of distribution followed by a considerable number of other southern forms, as pointed out by Professor D'Arcy Thompson in the Proc. Roy. Soc. Edin., vol. xxii, p. 313. The holothurian Colochirus brevidentis is in the same category if Ludwig's identification of specimens from Juan Fernandez is correct. Professor Kirk also found Ophiomyxa australis at the Chatham Islands.

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Amphiura elegans.

I have followed Professor Bell in retaining Leach's specific name elegans (Catl. Brit. Ech., p. 119), although Lyman preferred that of Delle Chiaje—squamata (Chg. Rpts., vol. v, p. 136). This is an exceedingly interesting little species, on account of its remarkable geographical distribution, being widely diffused in the temperate seas of both hemispheres. It occurs freely on seaweed in rock-pools near Wellington, and I found great numbers on seaweed in rock-pools at Gisborne, as noted by me in the Linnean Society's Journal (Zool.), vol. xxvi, p. 191. It also occurs in deeper water among sand and gravel, Mr. H. Suter having dredged up a quantity of specimens in Lyttelton and Akaroa Harbours.

Ophiactis nomentis, n. s.

The disc is roundly subpentagonal in form, somewhat swollen above, constricted above the arms, and the edge thick and rounded between the arms; about 9 mm. in diameter. The arms increase slightly in width to a short distance from the disc and then taper evenly to a fine extremity; they are about 46 mm. in length, and the greatest width is 2 mm. The disc is closely covered above with small, rounded, irregular, imbricating scales, somewhat elongated towards the edge of the disc, armed with small, bluntly pointed, smooth spinelets, thickly placed at the edge of the disc between the arms, and a few scattered irregularly around the radial shields and sometimes towards the middle. The radial shields are seed-of-pear-shaped, 2 mm. long and a little less than 1 mm. wide, separated their whole length by a narrow wedge of small elongated scales. One rounded leaflike mouth-papilla on each side of the base of the mouth-angle. Mouth-shields round, with a very slight peak without. Side mouth-shields triangular with re-enteringly curved sides, meeting or almost meeting within. The teeth are short, stout, and wedgelike, the uppermost one being often rounded and leaflike. The upper arm-plates are transversely oval, much broader than long, covering the whole of the upper part of the arm; some of them are usually split into two or three pieces in adult specimens. The under arm-plates are circular, slightly truncated within. The side arm-plates, which do not meet, bear four short, stout, blunt, divergent arm-spines, and one round, leaflike tentacle-scale. The colour of dried specimens is brownish-grey above, the arms being banded or variegated with darker grey, and yellowish-white beneath. In life the disc is reddish, becoming bright-red towards the edge of the disc between the arms.

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A number of specimens were dredged up by Mr. Haylock at Cape Maria van Diemen, between the little island on which the lighthouse stands and the mainland, in about 4 fathoms of water, among seaweed and stones.

Ophiopeza danbyi.

This species is given in the “Index” without a reference. It was described by me in the Linnean Society's Journal (Zool.), vol. xxvi, p. 189 (1897), from a specimen collected by Mr. Danby at the Kermadecs, which is now in the Canterbury Museum.

A number of echinoderms are given in the “Index” which are not in my “List.” The reason of this is that Captain Hutton included the species found at the Kermadec Islands and “Challenger” Stations Nos. 170 and 170a. These may be included in our fauna, as the Kermadecs are part of this colony, and their land fauna and flora are essentially New-Zealandian.

Asteroidea.

Gnathaster miliaris.

1846.

Astrogonium miliare, Gray. Proc. Zool. Soc., p. 80.

1866.

" " Synopsis of Starfish, p. 10.

1872.

Pentaceros rugosus, Hutton. Proc. Zool. Soc., p. 812.

1872.

Astrogonium rugosum, Hutton. Cat. Ech. N.Z., p. 7.

1876.

Pentagonaster miliaris, Perrier. Révis. Stell. Mus., p. 220.

1878.

" " Archiv. Mus. Hist. Nat., 2e série, vol. 1, p. 84.

1889.

Gnathaster miliaris, Sladen. Chger. Rpts., vol. xxx, pp. 286, 750.

1897.

Gnathaster rugosus, Farquhar. Jrn. Linn. Soc. (Zool.), vol. xxvi, p. 194.

Having compared Gray's description and figure and Hutton's description of this species, and of his supposed new species (N. rugosus) with Perrier's original description of Goniodon dilatatus, and with Hutton's type specimens in the Colonial Museum, I have come to the conclusion that Hutton's N. rugosus is N. miliaris, Gray; and the specimens identified by Hutton as Gray's species are specimens of Perrier's Goniodon dilatatus. The synonymy of Gray's species G. miliaris will therefore stand as above.

Goniodon dilatatus.

1872.

Astrogonium miliare, Hutton (non Gray). Cat. Ech. N.Z., p. 7.

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1876.

Pentagonaster dilatatus, Perrier. Archiv. Zool. Expér., vol. v, p. 33.

1889.

Gnathaster dilatatus, Sladen. Chger. Rpts., vol. xxx, pp. 286, 750.

1894.

Goniodon dilatatus, Perrier. Stellérides du Travailleur, p. 244.

1901.

Goniodon dilatatus, Loriol. Notes pour servir à l'Etude des Echinodermes, ix, p. 43.

My late friend Mr. A. Reischek kindly sent a copy of Perrier's original description of this species to me, which may not be accessible to New Zealand naturalists. It runs thus: “Espèce reamarquable par sa forme rappelant un peu celle du Pentagonaster pulchellus, bien que les plaques marginales soient disposée tout autrement. Le corps est plat, les bras courts, larges, dilatés au sommet en forme de spatule, et terminées par un bord arrondi. R=2r, d=90 millimètres. Les deux faces dorsale et ventrale sont uniformément couvertes de granules assez gros et bien distincts les uns des autres. L'ensemble des granules correspondant à chaque ossicule dorsal est nettement séparé des granules voisis, de sorte que les ossicules sont parfaitement distincts les uns des autres; sur la face ventrale, au contraire, il est difficile de distinguer les limites des ossicules squelettiques. Les plaques marginales augmentent graduellement de largeur depuis le sommet de l'arc interbrachial jusque vers le sommet des bras, les dernières seules diminuant de nouveau. La longueur de toutes ces plaques demeure au contraire sensiblement constante. C'est à l'élargissement graduel des plaques marginales qu'est due la form spatulée des bras; l'aire limitée par les plaques, qui sont au nombre de dix-huit, présente comme d'habitude la forme d'un pentagone à côtés concaves. La plaque madréporique est grande et subcentrale. Sur la face ventrale les piquants ambulacraires sont disposés sur trois rangs dont le niveau supérieur dépasse à peine celui de la granulation générale. Les piquants du premier et du seconde rang sont un peut plus gros que ceux du troisième, dont les dimensions transversales sont identiques à celles des granules ventraux.”

Asteropsis imperialis.

This species is given in the “Index” without a reference, and wrongly spelt “Asterospis.” It was described by me in the Linnean Society's Journal (Zool.), vol. xxvi, p. 193. The unique type specimen, which is from the Kermadecs, is in the Canterbury Museum.

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Asterias rodolphi.

This Kermadec species is also given in the “Index” without a reference. A few notes were added by me in the Linnean Society's Journal (Zool.), vol. xxvi, p. 192, to M. Perrier's brief original description, which runs thus: “Very like A. gracialis, L., from which it differs chiefly in the number of rays, which is seven, and the position of the ventral spines near the ambulacral spines, which form a triple and not a double series as in the European species”: Ann. Mag. Nat. Hist., ser 4, vol. xvii, p. 34 (1876). The type, which is in the British Museum, was collected by Macgillivray during the voyage of H.M.S. “Herald” in 1854.

Cribrella ornata.

Professor H. B. Kirk found this species in abundance at the Chatham Islands. He also observed Asterias calamaria and Asterias scabra at the same place.

Echinoidea.

Echinobrissus recens.

This extremely interesting little echinoid occurs in Tasman Bay; specimens may be found on the beach at “The Sands,” near Richmond, after northerly gales. It also occurs pretty freely in the channel at the entrance of Port Nicholson. Mr. Haylock has collected a good many specimens on the beach between Day's Bay and Pencarrow Light, some of them being quite fresh with the spines on. The spines on the abactinal surface are short, slightly tapering, with rounded tips, and longitudinally striated, 1.6 mm. long; those on the actinal surface are longer and finer, the longest, 3.5 mm., being near the mouth. There is a specimen in the Canterbury Museum from the Chatham Islands, and one from Stephen Island. It also occurs in Foveaux Strait.

The genus Echinobrissus attained its maximum development in the later Jurassic and early Cretaceous periods, when its forms were numerous and widely diffused. Two species occur in the Australian Tertiaries, and one (E. papillosus) in the New Zealand Eocene. If this species occurs in Madagascar, as stated by Agassiz in the “Revision,” its distribution is not so remarkable as would appear at first sight, for it is evidently a very old form, and in far distant times—probably before the Cretaceous—Madagascar was much nearer to New Zealand zoologically than it is now. This is shown by the relationship

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between the extinct Dinornithidœ of New Zealand and the extinct Æpiornithidœ of Madagascar. Our fern-bird belongs to a genus (Sphenœacus) found only in New Zealand and South Africa, with a near relation in Madagascar; and the beetle Sternaulax and the marine shell Littorina mauritiana occur in New Zealand and Madagascar, the latter being also found in Australia. (See Introduction to the Index FaunÆ N.Z., p. 6.)

Centrostephanus rodgersii.

Strongylocentrotus erythrogrammus.

New Zealand specimens of both these species are in the Colonial Museum at Wellington, as noted by me in the Linnean Society's Journal (Zool.), vol. xxvi, p. 189. These are omitted from the “Index,” although Strongylocentrotus tuberculatus, which is noted as a New Zealand form on the same page, is retained. C. rodgersii ranges from New Caledonia to Tasmania and New Zealand, and S. erythrogrammus occupies the same area, with extensions to Japan and Chili. S. tuberculatus was found at the Kermadecs by Mr. Haylock.

Sphærechinus australe.

This species, which is given in my “List” and omitted from the “Index,” occurs in Australia, Tasmania, Mauritius, and the Society Islands. It is recorded by Agassiz as from New Zealand, and is one which we might expect to find here; as, however, it is not known to New Zealand naturalists, it may be omitted from our list, at any rate for the present.

Holopneustes inflatus.

I gave this species in my “List” on the authority of Mr. Agassiz (Rev. Ech., pp. 136, 483; and Chger. Repts., iii, p. 274), but as it is not known here it may be correctly omitted from the “Index.” Dr. Ramsay says, “Three species—H. inflatus, H. porosissimus, and H. purpurescens—are somewhat doubtfully recognised by Agassiz, and without a large series of specimens it is almost impossible to distinguish the varieties: it is quite likely that they all belong to one species. As Holopneustes occurs on the east and south coasts of Australia, it is not unlikely to occur in our seas. Some dredging operations have recently, I believe, been carried on in the neighbourhood of Dunedin and some of the species now omitted from our list may have been rediscovered, but I do not know what echinoderms were obtained.

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Echinus margaritaceus.

Peronella rostrata.

Metalia sternalis.

These three species are omitted from the “Index.” I included them in my “List” on Mr. Agassiz's authority (Chger Repts., iii, pp. 274, 275). Echinus margaritaceus is widely spread in southern seas, having been found at the Straits of Magellan, Juan Fernandez, Kerguelen Land, and Heard Island, and we may expect to find it in our southern waters; but as it is not known to New Zealand naturalists it may be omitted for the present. Peronella rostrata is a Zanzibar form which does not occur here. We have a species of Peronella, however, specimens of which are in the Otago Museum, from the West Coast Sounds. Two species of Peronella occur in Australian waters, P. decagonale and P. peronii. The former appears to be confined to the Indo-Pacific region, and the latter ranges from the Philippine Islands to Tasmania. Being unacquainted with these, I cannot identify our species. Metalia sternalis is a tropical form, widely diffused in the Indo-Pacific region, and does not, I think, occur in our seas.

Brissopsis luzonica.

I do not know of the occurrence of this species here. It is a tropical form, widely spread in the Indo-Pacific region, and I do not think it belongs to our fauna. It is given in the “Index,” although I marked it “doubtful” in my “List” for both New Zealand and Australia.

Salmacis.

We have a species of Salmacis in New Zealand, but I do not think it is S. globator, which is given in my “List” and in the “Index.” Specimens are in the Otago Museum from Stewart Island, and one denuded test is in the Colonial Museum. I believe it belongs to Bell's S. alexandri (“Alert” Report, p. 118), which is a very variable form, and occurs freely in Port Jackson (Catl. Ech. Aust. Mus. (1891), p. 50).

Phyllacanthus dubia.

A specimen of Phyllacanthus, which I take to be P. dubia, was found by Mr. Haylock at the Kermadec Islands. This is an Indo-Pacific species, having been found also in New Caledonia, North Australia, the Red Sea, Zanzibar, and the Bonin Islands.

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Art. X.—Note on the Bipolarity of Littoral Marine Faunas.

[Read before the Wellington Philosophical Society, 1st August, 1906.]

The littoral marine fauna of New Zealand, in common with the land fauna, consists of several distinct elements. In all or nearly all the groups which have been worked up we find two comparatively small elements, which are nevertheless more interesting than the other parts of our fauna—namely (1) an autochthonic element, consisting of species which are peculiarly Neozelandian in type, and, having no relations in any other parts of the world, are entirely distinct from all other forms; these have arisen in the New Zealand area in extremely remote geological times; and (2) a representative element, consisting of species which are identical or closely allied to species inhabiting the northern temperate or Arctic regions. The latter are known as bipolar forms. The autochthonic element is stronger in the land fauna, but it is also well marked among our marine animals; and, although only a faint trace of the northern element is found among our terrestrial animals, it is much more strongly marked in the marine fauna.

The bipolar forms are thought by some naturalists to be the remnant of a fauna which was cosmopolitan in very early times, when a more equable climate probably obtained all over the world.

Of Hydroida we have six littoral species in New Zealand identical with European forms: Obelia geniculata, Sertularia operculata, Sertularella polyzonias, Plumularia setacea, Antennularia antennina, and Tubiclava fruticosa. Campanularia caliculata var. makrogona, which occurs freely in Wellington harbour, is a distinct species, for it always has the large type of gonangia figured by Bale in the Proc. Linn. Soc. of New South Wales, 1888. It is closely related to the European species C. caliculata, which occurs in Australia. Aglaophenia filicula occurs here and at the Azores; and almost all the genera are European.

The sea-anemone Actinia tenebrosa, which is only found between tide-marks, is the southern representative of, and closely allied to, A. equina. The genera are almost all common to both regions. Two New Zealand species known to me are entirely distinct from all others—namely, Halcampactis mirabilis,

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and a very beautiful, large, undescribed species with large kidney-shaped lumps on the body-wall, giving it somewhat the appearance of a bunch of grapes, for which a new genus will have to be established.

The two lime-sponges, Leucosolenia clathrus and Leucosolenia cerebrum, have been found here by Professor Kirk; the former occurs in the English Channel and the latter in the Adriatic.

Of echinoderms, Amphiura elegans (A. squamata) is widely spread in the North Atlantic; Stichaster insignis is the southern representative of the far northern form S. albulus—the two species are remarkably near, and both increase by subdivision; and Cribrella compacta is a near ally of C. occulata. The little heart-shaped sea-urchin Echinocardium australe extends into the North Pacific, but this has a great bathymetric range, having been obtained from a depth of 2,675 fathoms off Japan. It is very closely related to the Atlantic species E. cordatum. The only genus of echinoderms peculiar to New Zealand is Ophiopteris, with but one species, O. antipodum.

The large gephyrean worm Echiurus neozelanicus, which occurs freely in Wellington harbour, is nearly allied to its congener E. unicinctus of Japan. Priapulus caudatus, which occurs along the coasts of Greenland, Norway, and Great Britain, and in both the North and Baltic Seas, was found by the “Southern Cross” Expedition in the Antarctic off Cape Adair. Mr. Shipley says, “The genus, too, seems also bipolar in its distribution. P. bicaudatus lives in the North Sea and Arctic Ocean, and is represented in habits and its two tails by M. de Guerne's Piapuloides australis from the neighbourhood of the Magellan Straits” (Rpt. “Southern Cross” Collections, p. 285). Mr. A. Willey, in his report on the Polychæta of the “Southern Cross” Expedition, says, “Perhaps the most interesting feature of the collection is the addition of the characteristic northern maldanid, Rhodine loveni, Magn., to the Antarctic fauna. Besides this, two other genera not hitherto recorded in the south are represented by species slightly different from their northern congeners—namely Gatlyana (= Ngchia) cristata, n. sp., and Malmgrenia crassicirris, n. sp. (p. 262).

The stalked ascidian Boltenia pachydermatina, which is abundant at low water on our southern coasts, in the colder water of the Antarctic Drift, occurs also on the coasts of Greenland.

Eighteen species of New Zealand Bryozoa are identical or closely related to European forms—namely, Scrupocellaria scrupea, Bugula neritina, B. avicularia, Membranipora membranacea, M. pilosa, M. lineata, M. solidula, Microporella ciliata,

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M. malusii, Mucronella variolosa, Retepora cellulosa, Membraniporella nitida, Hippothoa flagellum, Schizoporella hyalina, Crisia denticulata var., Idmonea serpens, Entalophora raripora, and Diastopora patina. I have obtained these names by comparing Hutton's revised list of New Zealand species (Trans. N.Z. Inst., xxiii, 102) with Miss Jelly's “Synonymic Catalogue,” assuming that those with a reference to Hinck's “British Marine Polyzoa,” Fleming's “British Animals,” or Johnston's “British Zoophytes” are European forms. Some of them are probably cosmopolitan. Tenison-Woods described a fossil bryozoan, Fasciculipora ramosa, from New Zealand Tertiary beds so nearly allied to a species from the Lower Pliocene of Europe that if it had been found in the same locality it would have been regarded as a mere variety (“Palæontology of New Zealand,” pt. iv, p. 31). Smittia landsborovi occurs in the Arctic Ocean, European Seas, and the Antarctic off Cape Adair.

Our marine crustacean fauna has evidently a considerable amount of affinity with that of northern Europe. Myers, in the introduction to his “Catalogue of New Zealand Crustacea” (1876), says, “The remarkable resemblance between the carcinological fauna of New Zealand and that of Great Britain has been adverted to by Dana and other authors, and is sufficiently striking.” This generalisation was quoted by Dr. Chilton in his presidential address to the Philosophical Institute of Canterbury, 1904, on “Arctic and Antarctic Faunas,” which has not been published. Nearly all the species of the genus Gnathia are European, and one species, G. polaris, occurs in the Antarctic off Cape Adair. I hope Dr. Chilton will presently give us an account of the New Zealand species which are identical with and nearly related to northern forms, as he and Mr. G. M. Thomson have worked up this group. The character and affinities of our terrestrial Crustacea are extremely interesting. Mr. Thomson kindly gave me an account of these some time back, which I embodied in a continuation of my paper on “The New Zealand Zoological Region” (Nature, vol. lxi, p. 246). This continuation was never published, and unfortunately both Mr. Thomson's and my own notes were lost in the confusion of changing residence from Wellington to Auckland and then back to Wellington again.

The following list of marine Mollusca which occur in New Zealand and the north temperate region has been kindly furnished to me by Mr. H. Suter: Crepidula crepidula, Mediterranean, Atlantic; Tritonium costatum, Mediterranean, West Indies, Africa, Brazil; Tritonium rubicundum, Great Britain, Mediterranean, India to Japan; Venericardia corbis, Mediterranean, Pliocene fossil in Italy; Trivia europœa, Kellia suborbicularis,

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and Thyasira flexuosa, Atlantic; Cassidea pyrum, Lima bullata, and a variety of Cassidea labiata, Japan; Ancilla rubiginosa, Japan, China, Malacca, Madagascar; Mytilus edulis and Saxicava arctica, cosmopolitan. Mr. Suter says the above list is probably far from complete.

Of shore fishes we have six species which are identical with those of the European seas—namely, Trachurus trachurus, Zeus faber, Conger vulgaris, Scymnus lichia, Echinorhinus spinosus, and Acanthias vulgaris. Our species of Cyttus and Polyprion are representatives of the North Atlantic forms; and the southern mackerel Scomber australasicus is either identical with or very closely allied to Scomber colias. The genus Argentina, which is characteristic of the seas of northern Europe and the Mediterranean, is represented in New Zealand waters by A. decagon.

These facts show that there is a good deal of evidence in the character of the littoral marine fauna of New Zealand which makes for the support of the bipolar theory.

Professor D'Arcy Thompson, in his critical review of Sir John Murray's paper, maintains “that an actual community of forms is not proven, save for a very few forms, some peculiar to the extreme depths of the sea, and others that inhabit the surface of the ocean in colder latitudes while represented in the deeper and colder waters of tropical seas” (Proc. Roy. Soc. Edin., xxii, p. 312). The forms enumerated by me, with the exception of Echinocardium australe and perhaps one or two molluscs, are, however, essentially littoral species, which do not extend into deep water, and do not appear to be represented in the intervening seas.

I have gathered together these scraps, which are, no doubt, far from complete, and if they contain any mistakes I hope they will be corrected by others.

If there is anything of value in the bipolar theory, and in view of the fact that there are immense differences in the variation of species—some forms remaining unchanged or changing very little through vast geological periods, while others are unstable and change very rapidly—we should expect to find a few species in the Antarctic and sub-Antarctic regions identical with those of the corresponding northern regions, a considerable number of representative species and the genera for the most part common to both areas, with a few genera in all the groups peculiar to each area of distribution in both regions; and this is, I think, what we find in a comparison of the New Zealand littoral marine fauna with that of the North Atlantic. Near alliance of species is probably more favourable to bipolarity than actual identity.

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I know little or nothing of the North Pacific fauna, but a comparison on the same lines would be extremely interesting. Then we need an Australian naturalist to give us an account of the character and affinities of the Australian marine fauna. And when our Tertiary fossils have been worked up, a comparison with those of the northern temperate and sub-Arctic regions will perhaps shed a little more light on this exceedingly interesting problem.

Art. XI.—The New Zealand Plateau.

[Read before the Wellington Philosophical Society, 3rd October, 1906.]

Plate V.

At the time when the “Challenger” Reports were published only a very few deep-sea soundings had been made with the improved sounding apparatus in the neighbourhood of New Zealand, and the New Zealand Plateau was only approximately sketched on the map published with the reports. But since the “Challenger” Expedition a great deal of sounding-work has been done in the South Pacific by H.M.S. “Penguin” and other vessels, in connection with the laying of the Pacific telegraph-cable from British Columbia to New Zealand and Australia; and a line of soundings was run by Sir James Hector from the Bluff southward to the Macquarie Islands, north-eastward to the Chathams, and westward to Lyttelton. The contour of the sea-bed surrounding this country is therefore fairly well known, and we can now map out the Plateau correctly, except at one or two points.

The map now published is compiled chiefly from the Admiralty chart of the Pacific Ocean with the latest additions, and a map of the Pacific, showing soundings, published by the India-rubber, Guttapercha, and Telegraph Works Company.

This great Plateau forms a very irregular area. It extends far to the south and south-east beyond the outlying islands (Auckland, Macquarie, Campbell, Antipodes, Bounty, and Chatham), but the boundary-line has not been certainly fixed here and may extend a little beyond that marked on the plan.

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North-westward the Plateau extends in an immense submerged ridge, which may be called the “Lord Howe Island Ridge,” and joins or almost joins the New Caledonian Plateau. The deep channel which runs along the south-western side of New Caledonia may extend northwards between the D'Encrecasteaux and Chesterfield Reefs, thus cutting off the New Caledonian Plateau from that of New Zealand by a narrow channel; but this is not very probable, and a few soundings are needed here to decide the point.

This great ridge is the most striking feature of our Plateau, and it has profoundly affected the character and distribution of the fauna and flora of New Zealand—a great stream of life having come in this way.

At the southern extremity of the Lord Howe Island Ridge, and lying to the north-westward, there are two smaller ridges, that lying farthest north being the extension of what is now known as the Auckland Peninsula. Between these ridges lie two valleys or depressions, in which, no doubt, large rivers flowed at the times of elevation.

The great backbone ridge of New Zealand, with its divergent ranges, must have presented a truly grand appearance during the periods of elevation—the beginning of the Tertiary era and early Pliocene—when the land stood five or six thousand feet higher than now, with its vast snowfields and glaciers grinding out the great valleys and fiords, and forming the moraines which lie scattered about the country.

North of the main body of the Plateau and east of the Lord Howe Island Ridge a large depression known as the “Gazelle Basin” lies between New Zealand and the Fiji Islands, and further eastward again, beyond the Kermadec Ridge, the “Aldrich Deep” culminates near the Kermadecs at a depth of over 5,000 fathoms, plumbed by Captain Aldrich. Its western boundary is an immense mural escarpment, running southward from the Tonga Islands past the Kermadecs, and down the eastern side of the North Island towards Cook Strait, where it turns abruptly eastward to the north of the Chatham Islands. On the west our Plateau is separated from Australia by the “Thomson Deep,” which is evidently one of the great primordial depressions of the earth's crust, and accounts for the fundamental difference between the animals and plants of Australia and New Zealand. Eastward from the Australian coast-line the sea-bed rapidly plunges down to a great depth (2,600 fathoms), and then rises gradually in a long undulating slope to New Zealand.

The evidence of the former great extension of New Zealand, and of the vast changes that have occurred in this region, as

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seen in the geology and in the character and relationships of the fauna and flora, has been admirably discussed by the late Captain Hutton in the “Transactions of the New Zealand Institute,” and in the introductions to the “Index FaunÆ” and “Animals of New Zealand,” and by Messrs. Laing and Blackwell in the introduction to their beautiful “Plants of New Zealand.” And if these works are studied with the map of the Plateau as now developed the relations and reasons of most of the observed facts may be understood.

Art. XII.—On the Occurrence in New Zealand of Platalea regia, Gould.

[Read before the Philosophical Institute of Canterbury, 11th July, 1906.]

In vol. ix of our Transactions, Dr. (now Sir) Walter Buller recorded the first occurrence of Platalea regia, Gould, in the colony. That specimen was shot in April, 1875, near the mouth of the Manawatu River, and is now in the collection of birds in the Colonial Museum. In vol. xxvii Sir Walter Buller stated that he had been informed by Mr. Townson, of Westport, that another specimen had been shot, on the Buller River, about January, 1892, and was preserved in Dr. Gaze's collection.

I have now to put on record another occurrence of this fine bird, making the third in a space of thirty years. The specimen, which I now exhibit, was shot on a lagoon near Greytown North, in the Province of Wellington, in the month of May, 1905. This specimen is an immature bird.

As the bird was set up as you now see it when I heard of it, I can only give you the words of the man who shot it. He said, “I was coming home by the side of the lagoon from rabbit-shooting when the bird rose from a hollow just in front of me. I shot it, and was surprised at the bird when I picked it up. I took it along to a friend of mine who dabbles in bird-stuffing, and he set it up for me. I found that it had been seen for some weeks previously flying about the lagoon, and several attempts had been made to stalk it, but all had been unsuccessful owing to its wariness. It had even been christened ‘the white hawk.’”

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Art. XIII.—New Zealand Ctenophores,

[Read before the Otago Institute, 14th August, 1906.]

Plate VII.

Hitherto the only member of that group of “meoln-jellyfish” that has been attributed to our seas is a pleurobrachid, named Cydippe dimidiata, Lesson. * This was obtained in the Tasman Sea—or, as it was termed, “the channel between New Zealand and New South Wales”—in 1770, by Banks and Solander, during Captain Cook's first voyage. It was originally named by them “Beroe biloba,” but by Lesson was placed in his genus Eschscholthia and described by him as E. dimidiata. It had been figured previously by Eschscholtz and Chun, presumably judging from this figure, identifies it as a member of the genus Cydippe. § It has not been studied further, and nothing more is known of it than the short account given by Lesson. I think that it may well be removed from our list.

During the last few years I have collected or received a few specimens of ctenophores which belong to two genera—Beroe, Browne, 1756; and Euplokamis, Chun, 1880—for each of which I propose, though with considerable diffidence, owing to lack of recent literature, a new specific name, though I recognise that these may be found superfluous.

Beroe shakespeari, n. sp.

I received a number of individuals, well preserved in formalin, from Mr. R. Shakespear, the Curator of Little Barrier Island, to whom my thanks are given for his kindness and thoughtfulness in presenting them to me.

The specimens, some of which have a faint roseate hue, are subglobular, but slightly compressed in the transverse or infundibular plane, so that the “sagittal” diameter is rather greater than the “transverse.”

The fifteen individuals vary from 27 mm. to 62 mm. in height, and from 20 mm. to 50 mm. in diameter across the base, being

[Footnote] * Lesson, Ann. Sci. Nat. (Zool.). 1836, ser. 2, vol. v, p. 254.

[Footnote] †;Lesson, Hist. Nat. Zoophytes, Acalephes, 1843, p. 102.

[Footnote] ‡ Eschscholtz, Syst. der Acalephen, 1829, pl. ii, fig. 2.;

[Footnote] § Chun, Fauna u. Flora des Golfes v. Neapel. Ctenophoren, p. 287 (1880).

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rather greater than this near the equator. One or two have a more oval form, the diameter being proportionally smaller, but in the majority the vertical and horizontal diameters are approximately equal.

The mouth does not occupy the whole of the base—i.e., it is not relatively so large as in the two well-known Mediterranean species, B. ovata and B. forskalii. In the present species the oral moiety of the body is somewhat contracted, sloping downwards and inwards, so as to be nearly horizontal around the mouth.

The costÆ, even in the largest individuals, do not extend down to the margin of the mouth. This I supposed at first to be related to the immaturity of the specimens, but I find that the largest ones contain fully developed spermatozoa in the meridional canals and young ova, so presumably they are practically full-grown, or at any rate adult. These gonads are limited to the meridional canals, and extend down to their lowest limits. The costÆ taper off to a fine point orally, about two-thirds of the total height of the body. The eight costÆ are not absolutely equidistant; the pair at each end of the sagittal plane run at first parallel with each other and are closer together than are the other costÆ; they then diverge rather suddenly, so that at the equator the distance separating the pair of subsagittal costÆ is only slightly less than that between other pairs. Thus, in the largest specimen, the distance between the pair of subtransverse costÆ at the equator is 12 mm., and between the subsagittals 10 mm. But at their origin these distances are 4 mm. and 1.5 mm. respectively.

The subcostal or meridional canals give off, right and left, branched cœca, which do not anastomose but terminate in slight dilatations; and to some extent those of one canal interdigitate with those of a neighbouring canal—as in B. ovata. The two stomodæal canals also give origin to a few fine outgrowths which take a horizontal direction on the wall of the stomodæum and branch slightly; these open into a series of inwardly directed branches from the meridional canals. These canals and branches are readily seen in the specimens, and I injected them in one large individual in order to confirm the observation. The four meridional and the stomodæal canal of each side open into a semicircular marginal canal at the edge of the mouth; but the two marginal canals do not meet at the ends of the sagittal plane. This fact is also in agreement with Chun's observations.

I have obtained specimens of this species of Beroe from the following localities: (a.) Little Barrier Island (Shakespear); 1906. (b.) Off D'Urville Island, off the north coast of the

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South Island (G. M. Thomson); 1900. (c.) Tauranga, on the east coast of the North Island (which is without date, and was collected during my predecessor's time).

The lot (a) include forms varying in size from 27 mm. to 62 mm. in length; the largest I take to be adults. The lots (b) and (c) are quite young, attaining a length of 9 mm., 6 mm., and even less. They exhibit a series of stages of growth corresponding to some of those figured and described by Agassiz * for B. roseola, on pp. 36, 37.

The youngest of my specimens has costÆ extending only a short way down the body (cf. fig. 52 of Agassiz); the meridional canals are relatively wide, have no branches, and open below into a semicircular canal (cf. figs. 57 and 58 of Agassiz); but in the former figure no marginal canal has appeared, while in the latter there are already indications of cœca which are absent in my specimens.

In the lot (a), even in the smallest, the meridional canals are provided with branched cœca, though the branches in the smaller individuals (20–25 mm. in length) are not so much ramified as in the older. (This measurement was taken after transference from formol to alcohol, in which the specimens shrink somewhat—losing about 20 per cent. of their former size). In all of them the costÆ extend more than half-way down.

Remarks.—The question arises, Is a new specific name required for this form? Chun (p. 306) has pointed out in his memoir the immense amount of synonymy—both generic and specific—that occurs in the family BeroidÆ, and comes to the conclusion that, at any rate in the Northern Hemisphere, not more than three species exist—namely, Beroe ovata, B. forskalii, and B. cucumis. But he acknowledges that the forms that have been gathered in the Pacific Ocean, and figured by older authors, are insufficiently described to enable him to decide as to their validity or otherwise.

More recently, a species B. australis has been described by Agassiz and Mayer from Fiji, which, however, is quite distinct from the present form, and appears nearly related to B. forskalii.

But from each of the three species from the Northern Hemisphere the present one differs in the following points: From B. forskalii in form and sundry details which serve to distin-

[Footnote] * Agassiz, N. American AcalephÆ, Ill. Catal. Mus. Harvard, 1865.

[Footnote] † Agassiz and Mayer, “The Acalephs of Fiji,” Bull Mus. Harvard, 32, 1899, p. 177.

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guish this from B. ovata. From B. cucumis (= B. roseola, Agass.) in the presence of network of canals on the wall of the stomodæum—just that feature which most distinctly separates the North Atlantic form from B. ovata.

We are reduced, then, to a comparison of our New Zealand species with B. ovata, which, though it descends into the tropical and even the southern regions of the Atlantic, does not appear to have been met with eastward of Africa. From B. ovata our species differs in its more globular form, for even in the individuals in which the horizontal diameter is least in relation to the vertical (e.g., 60 mm. × 40 mm.) the proportions are very different from those of B. ovata, in which the length is more than twice the breadth; secondly, the approximation of the upper ends of the subsagittal costÆ; thirdly, in the cessation of the costÆ before the lower margin is reached (which may possibly be due to the specimens being not yet fully grown); fourthly, in the fact that the mouth is not at the widest part—that the lower margin of the bell is directed inwards, so that the mouth is smaller, relatively, than in B. ovata. It may be that in life the mouth of our species is capable of a certain amount of mobility and distension.

Reference should perhaps be made to Agassiz's *Idyiopsis clarkii” from South Carolina, which in its more rounded form (see fig. 63, p. 39, loc. cit.) certainly resembles our species. Agassiz emphasizes the short vertical axis, the compressed body, the prominent costÆ, and depressed intercostal regions as distinctive of the genus, which later authors include in Beroe. But though there is a certain degree of resemblance, it is not probable that the same species would occur in the Atlantic and the Pacific. In “Idyiopsis clarkii” the polar area is, from Agassiz's figure, much more compact than in the present species.

The only Pacific species, besides B. australis, is Beroe macrostoma, Peron, which was obtained south of New Guinea during the voyage of the “Coquille.” Its shape is, however, like that of B. ovata, but Chun states that it is insufficiently described for identification. At any rate it is quite unlike our New Zealand species.

Euplokamis australis, n. sp.

The body is cylindrical, though slightly narrower at the aboral pole; the mouth is situated on a short and probably mobile tube, considerably narrower than the body.

[Footnote] * Agassiz, Illust. Cat. Harvard Mus., 1865, p. 39.

[Footnote] † Person, Voy. de la Coquille, Zool., p. 105, pl. xy, 2.

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The characteristic features upon which this genus was founded are (1) the great extent of the costÆ and the large size of the costal plates; (2) the tentacles issue from the tentacular sheath very high up the sides.

The present species differs from E. stationis, described by Chun, p. 283, in a few very evident characters. The “tentacle-base” is about one-third the length of the body, and extends considerably below the level of the “infundibulum”; the tentacillÆ or accessory filaments do not roll up into spirals, and I am unable to detect the transversely striated muscle within them which Chun describes for E. stationis.

Localities.—(a.) A single individual: off the north coast of the South Island, near Farewell Spit (G. M. Thomson). (b.) Several young ones: Otago Harbour (Cottrell). *

The largest individual is 20 mm. in length, with a greatest diameter of 12 mm. rather below the equator, while near the aboral pole its diameter is 8 mm.

The specimen preserved in formalin exhibited the internal anatomy well (see Plate VII, fig. 1). The gonads are present, but the gametes not fully developed. The long “infundibular canal,” large “infundibulum,” the radiating canals from it, are all shown. Those from Otago Harbour are young ones, and in them the enormous breadth of the costÆ and the great size of the individual costal plates can be seen.

These young ones measure, on an average, 5 mm. × 3 mm. The costÆ almost touch one another, and each is made up of 12 plates. In these young ones the aperture of the tentacle-sac is much nearer the apex, and the length of the tentacle-base is about half the length of the animal, and its lower end reaches nearly to the lower end of the costa: this is perhaps due to unequal contraction of the parts: these had been fixed in corrosive sublimate and preserved in alcohol, which causes the jelly to shrink to a much greater extent than does formalin.

Remarks.—A comparison of the figures and description of E. stationis with those of our species makes it evident that there are considerable differences between them. But since the publication of Chun's monograph on the Mediterranean ctenophores two other species belonging to this genus have been described. E. cucumis, Mertens, has been redescribed by Vanhoffen, to whose account, however, I have not access. It was obtained near Unalaschka, in the Behring Sea. The short account given by Mertens is referred to by Chun (footnote, p. 284), where it is

[Footnote] * I have also a specimen, 9 mm. in length, from Port Jackson.

[Footnote] † Vanhoffen, “Ctenophoren,” in Nordische Plankton, 1903.

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stated that the tentacle-base is situated in the neighbourhood of the stomodæum. E. californiensis has been described by Torrey, * whose account of it I have been unable to obtain. It was captured off San Diego, on the coast of California. It is possible that our New Zealand species is identical with it.

In addition, there is an earlier species, E. elongata, of Quoy and Gaimard, which was placed by them in the genus Beroe, then by Lesson (loc. cit., p. 103) in the genus Janira (as J. quoyi), but has been by Chun recognised as a Euplokamis, who resuscitated the original specific name.

This species was taken in the Atlantic, off the coast of Africa; in it the tentacle-base is more than half the length of the body.

From this species, also, mine is evidently quite distinct.

Explanation of Plate VII.

  • (Figs. 14 refer to Beroe shakespeari; figs. 5–7 refer to Euplokamis australis.)

  • Fig. 1. Beroe shakespeari, natural size of preserved specimen: a, canals containing gonads; b, costÆ; c, mouth. The apex is retracted.

  • Fig. 2. The same, view of apical pole (natural size), showing the compression of the animal, and the apical approximation of the sub-sagittal costÆ (d).

  • Fig. 3. A younger individual (natural size).

  • Fig. 4. A very young specimen (enlarged), showing short costÆ, wide canals, without the cæcal outgrowths.

  • Fig. 5. Euplokamis australis, adult individual (x 2½), showing, by transparency, the internal anatomy. The tentacles are partially retracted.

  • Fig. 6. A young specimen (X 4). The costÆ are wide; the tentacles, almost fully protruded, issue much nearer the apex than in the adult.

  • Fig. 7. A portion of a young one bisected to show the great length of the tentacle-sac at this stage as compared with adult (perhaps due to differences of preservation). b, the tentacle-base.

[Footnote] * Torrey, Univ. California Publications, 1904, vol. ii. p. 46.

[Footnote] † Quoy and Gaimard, “Voy. Astrolabe,” iv, p. 37, pl. vi, Mollusques; and Atlas, pl. 90, figs. 9–14.

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Art. XIV.—On the Presence of another Australian Frog in New Zealand.

[Read before the Philosophical Institute of Canterbury, 14th November, 1906.]

Our New Zealand fauna can only boast of one amphibian—namely, the indigenous frog, Liopelma hochstetteri—which was rare at the time of its discovery, but now is rarer still, if not almost extinct, only being found occasionally on the Coromandel Peninsula.

About the year 1868 several batches of the common green Australian frog, Hyla aurea, were liberated in different parts of New Zealand, as Christchurch, Wellington, &c.: these have increased so much that they are now to be seen in hundreds in many parts of the country.

Since the introduction of Hyla aurea there is only one other recorded instance of the introduction of frogs into New Zealand—namely, in 1898, when the Agricultural Department liberated another kind of Australian frog in this country.

Mr. T. W. Kirk, Government Biologist, writing in his report, says, under the heading of “Climbing-frogs,” “A consignment of six dozen of these insect-destroyers was also obtained and liberated at suitable spots in the following districts: West Coast (North Island), Wellington Province, Wairarapa, Hawke's Bay, and Auckland. This frog is similar to the ordinary common frog, so common in many parts of New Zealand, except that it has a very considerable advantage over that species in that its toes are provided with suckers, which enables the animal to climb trees and houses in search of insects. In Sydney I have seen these frogs at the top of a wall four stories high.” Unfortunately, Mr. Kirk does not mention the name of the frogs, and so far I have been unable to obtain it.

As late as 1904 Captain F. W. Hutton included only Hyla aurea in his list of naturalised amphibians, inserted at the end of the “Index FaunÆ NovÆ-ZealandiÆ” (page 348). However, for the last thirty years there has been living and increasing in West-land, especially around Greymouth, another species of Australian frog, which, though well known to the residents, was not thought to be very different from the common green frog of Canterbury. My brother, Mr. F. G. Marriner, was the first person to draw my attention to the presence of this amphibian around Greymouth. He told me that the frogs in the district had a

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peculiar whistle, which, when a large number joined in chorus, could be heard for some distance.

This at once aroused my interest, and through the kindness of Mr. H. West, Greymouth, I received five live specimens on the 3rd February, 1906.

On inspection I found that they were small frogs from 1¼ in. to 1 ½ in. in length, and of a brown colour; and in order to get them properly named I forwarded two live ones to Mr. J. J. Fletcher, of Sydney, who has done much work on Australian frogs.

He described it as Hyla ewingii, D. and B., var. calliscelis, and stated that it was included in the British Museum Catalogue of Batrachia (1882), page 406; he also informed me that it is one of the commonest frogs of eastern Australia and Tasmania.

On the 1st March I received about forty more from Mr. H. West, and so had a good opportunity of noticing any variation that might occur between different individuals. When exposed to light they are of a light-brown or even a very pale brown colour; there is usually a broad dark band running down the middle of the back, with two lighter broad bands on each side. The under-surface is lighter, and on the ventral surface of the thighs there is a yellow streak. When buried in the earth, or not exposed to a bright light, they go to dark reddish-brown colour—almost to a dark chocolate; and if one is buried with only a part of its body exposed to bright light, the buried portion turns a dark reddish-brown colour, and the exposed portion, no matter how small it may be, keeps its very light colour, the line of demarcation being very definite.

In the specimens that I received the male seems to be about ¼ in. smaller than the female, but otherwise they seem to be the same externally. The average size of my specimens was about 1 ½ in., but one specimen was about 2 in. in length. The head is large, eyes prominent, and snout short; hind feet are webbed, but fore feet are not so; all the digits have suckers at their tips.

The frogs in my case seem to be more strictly nocturnal than Hyla aurea, for they seldom come out in the daytime, except in wet weather. I kept a large number among some grass in a bell jar for some days; at night-time they could be seen climing all over the sides of the jar, but in the daytime they were almost all invisible. Since then I have kept them in a glass aquarium, with a dish of water and turf, but I seldom see them out in the daytime. Mr. H. West tells me that they are best caught at dusk or later, when they crawl over the grass, &c., and can be located by their peculiar cry, which somewhat resembles a whistle when compared with the hoarse croak of Hyla aurea.

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Arboreal Habits.

Hyla ewingii is a true climbing-frog, but according to Mr. J. J. Fletcher it has, at least in Australia, altogether or nearly lost the arboreal habits of a tree-frog. In Westland, however, it still seems to do a fair amount of climbing.

Mr. A. P. Harper, of Greymouth, in a letter to me, gives the following account of their climbing propensities: “I have personally seen these frogs (Hyla ewingii) crawling over black-berry bushes at a height of from six to eight feet above the ground, and also in the middle of a patch of berry, five yards by three at least. They even crawl along the thorny stems. I have also seen them on the macrocarpa and in the branches of the natural creepers on a dead tree-stem. The highest these frogs climb above the ground is, I should say, about eight feet. On one old tree-stem covered with creepers, ferns, &c. (as one so often sees here) there are nearly always some singing-frogs. It is just above a pool which exists in wet weather only. I rather think they climb when the pools are dry, but I am not sure.”

Several that escaped from me on the 9th March I afterwards found clinging to the top of grass-stems about two feet from the ground. Whether the abundance of bush in Westland has stimulated the frog to make use of a power which it has almost lost in Australia is difficult to decide. At any rate, the frog's environment in Westland would certainly be conducive to tree-climbing, and the abundance of undergrowth and creepers would make it very easy work.

Breeding Habits.

Hyla ewingii evidently extends its breeding season into the autumn, for on the 1st March two lots of spawn were laid in the aquarium.

On the 11th and 12th most of the eggs hatched out, and the tadpoles, dark-blue in colour, after swimming about for a little time, finally fastened themselves to the water-weeds or to the sides of the aquarium.

After about three weeks of normal development the tadpoles had grown to about 17 mm. in length.

On the 22nd March I put five into an aquarium, which I shall call A; it was out in the open, about 2 ft. by 2 ft. by 1 ft. in size, and held 4 cubic feet of running water. Five others I put into an aquarium (B) situated in a hothouse. The tank was 2 ft. 8 in. by 15 in. by 7 ½ in., and held about 2 ⅓ cubic feet of standing water. During the autumn they were always visible swimming about in the aquarium, but when the cold weather

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came they all disappeared. However, in aquarium B, whenever warm weather prevailed a solitary tadpole would appear from time to time, but in the coldest part of the winter no tadpoles were visible. It seems to me that when tadpoles remain as tadpoles all the winter, they must either bury themselves in the mud or else hide away among the water-weeds at the bottom of the aquarium, for I could never see any sign of them during the cold weather.

In the spring two tadpoles appeared in tank B, and only one in tank A, though no doubt several crayfish, which were living in the same aquarium, were partly responsible for the smallness of the number of survivors in the latter. Though the aquariums had plenty of water-weeds and submerged rocks, there was no place where the tadpoles could find a shallow landing-place where they would need their limbs for crawling, and the result has been that neither the lungs nor the limbs have been developed.

In autumn they would often come to the surface and take in large mouthfuls of air, as most tadpoles do, but since the winter they seem to have no inclination to come to the surface, but act to all intents and purposes similar to fish.

It has been stated that not only will tadpoles keep as tadpoles when reared under such conditions, but that they will increase in size in proportion to the size of the tank in which they are kept. Though three specimens are not sufficient to prove anything, yet the tadpoles in my aquariums do seem to uphold this fact. All of them were about 17 mm. in length when put into the tanks, and now the two in the tank holding 2 ⅓ cubic feet of water are only 25 mm. and 30 mm. respectively, while the one in the tank holding 4 cubic feet of water is now 45 mm. in length.

Distribution.

It seems almost certain that these frogs first made their appearance at Greymouth, whence they have extended inland and south.

From Greymouth they have spread either naturally or artificially up the Grey River for at least twenty-four miles to Ahaura, and very likely further, but so far all the places where I have found them to be present are on the south bank of the river, and none on the north, though it is not at all unlikely that they are on the opposite bank also.

At Brunnerton, eight miles from Greymouth along the river, they are plentiful, and seem to be increasing. No other kinds of frogs are said to be found there.

At Stillwater, nine miles from Greymouth and a mile past

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Brunnerton, according to Mr. Mallock there are both the green and brown kinds present, though the latter are more numerous and are increasing.

At Ahaura, twenty-four miles up the river, only the brown frogs are present, and these appear to be increasing.

At the centre of their dispersion, Greymouth, they are said to exist in very large numbers, and seem to be increasing.

The only other place where I have them recorded is Hokitika, twenty-two miles south of Greymouth, and there is no reference to them being found in between these towns, so that it is very unlikely that they travelled overland—indeed, rumour says that they were brought by an unknown person from Greymouth.

At Hokitika Hyla ewingii is now very scarce, if not quite extinct. Mr. James King, of Hokitika, says that ten years ago the big green frog (I suppose Hyla aurea) was introduced, and perhaps this has something to do with the decrease of the brown ones. From what I have seen of the Hyla aurea it would find the small brown frog very eatable, and if it does not stop at eating its own kind there is very little chance of it sparing the small strangers. It is evident that they have not extended in any direction from Hokitika, for I have no place outside of that town where they are known to be present.

Its Introduction.

The next question is to find out how this frog came to New Zealand.

I thought that at first it might be an indigenous frog, but there seems to be no evidence whatever to support this theory. There is no record of it in the early days of settlement, and at all the places where it has been found its origin can be traced directly or indirectly to Greymouth.

As the frog was so common on the east coast of Australia I thought that probably it came over among the ballast of some ship trading between that continent and Greymouth, but this theory was upset on making further inquiries in Westland.

Several of my correspondents say that these brown frogs, Hyla ewingii, were brought from Tasmania by a Mr. W. Perkins in 1875, and through the kindness of Mr. T. Eldon Coates, of Greymouth, I received the report which is generally accepted in that town. He states that a Mr. W. Perkins, who was a barrister in Greymouth for some time, brought some of these frogs from Tasmania in a glass bottle in 1875. They were liberated in a drain in Alexandra Street, Greymouth, whence they have spread to the surrounding country. They have remained in