Art. XXII.—Descriptions of New Native Flowering-plants.
[Read before the Auckland Institute, 11th December, 1917; received by Editors, 24th December, 1917; issued separately, 10th June, 1918.]
1. Myosotis cinerascens sp. nov.
M. perennis, follis culmisque pilis albis subrigidis appressis cinerascentibus. Culmi a radice complures graciles simplices vel parce divisi, 10–20 cm. alti, paene ad racemorum basim foliosi. Folia radicalia anguste obovato-spathulata ± 4 cm. longa circa 8 mm. lata obtusa, petiolis laminas aequantibus; caulina approximata consimilia ± 1.5 cm. longa lineari – obovata sessilia acuta. Racemi plenunque ± divisi breves subcapitati, raro simplices ac elongati. Flores albi breviter pedicellati; calyx ± 4.5 mm. longus, pilis rigidis patentibus subuncinatis dense hispidus, lobis brevibus acutis; corolla calyce subduplo longior anguste infundibuliformis, tubo lobis rotundatis brevibus ter quaterve longiore; stamina lineari-oblonga filamentis gracillimis duplo longiora, ad squamarum apices pertinentia; stylus maturus calycem ter quaterve superans. Nuculi oblongi, ter longiores quam lati, tenues brunnei.
Perennial, ashy-grey in all its parts. Culms from the root few or several, erect or ascending at the base, simple or sparingly branched, 10–20 cm. (4–8 in.) high, slender, leafy to near the base of the inflorescence. Radical leaves narrow obovate-spathulate, ± 4 cm. (1 ⅔ in.) long, 8 mm. (⅓ in.) broad, gradually narrowed into petioles about as long as the blades, obtuse rather membranous, densely hispid on both surfaces with rather long stiff appressed whitish hairs; midrib little conspicuous; cauline all much alike, closely placed and usually overlapping, linear-obovate acute sessile, about 1.5 cm. (⅔ in.) long. Racemes usually closely branched, short subcapitate, more rarely simple and more or less elongated. Flowers white on strongly hispid pedicels; calyx ± 4.5 mm. (⅙ in.) long, equalling or exceeding the pedicels, densely hispid with stiff spreading more or less hooked hairs, cut for about one-third its length into narrow acute lobes; corolla narrow funnel-shaped, nearly twice as long as the calyx, cut into broadly rounded lobes one-fourth as long as the tube; stamens linear-oblong, twice as long as the very slender filaments, reaching quite to the top of the scales; style slender, longer than the corolla, and ultimately about thrice as long as the calyx. Nutlets oblong, thrice as long as broad, thin, dark brown, shining.
Hab. — Limestone shingle-slip, Trelissick Basin, North Canterbury; 730 m. alt.: L. Cockayne! Broken River, on limestone debris, Canterbury Alps; 2,400 ft.: D. P.
This species is allied to Myosotis Traversii Hook. f. as Cheeseman understands that species, but by no means closely. It was almost certainly included in M. Traversii by Hooker f., and is no doubt the plant from the “Waimakeriri Valley” mentioned on page 195 of the Handbook of the New Zealand Flora. The specimen examined do not show the radical leaves in good condition, while the corollas are more or less withered and shrivelled.
2. Myosotis saxatilis sp. nov.
Perennis, pilis subtilibus brevibus albidis appressis viridi-incana. Caules pauci graciles, 8–12 cm. alti, supra pro parte tertia nudi. Folia radicalia obovato-lanceolata, 3 cm. longa 8 mm. lata, obtusa v. subacuta in petiolum sublatum angustata; caulina pauca subdistantia sessilia lineari-lanceolata acuta, radicalibus ½ breviora. Racemi conferte ramosi, ramia brevibus; flores 9 mm. longi, albi. Calyx alte 5-partitus, lobis linearibus; corolla infondibuliformis, tubo sublato calycem duplo superante; stamina corollae tubo vix breviora ad faucis squamas pertinentia; filamenta antheris breviora.
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Perennial, greenish-grey, everywhere hoary with delicate short appressed whitish hairs. Stems few from the root, slender, more or less ascending at the base then erect, the upper third naked, 8–12 cm. (3 ½–5 in.) high. Radical leaves rather numerous forming a compact rosette, obovate-lanceolate, 3 cm. (1 1/5 in.) long, 8 mm. (5/16 in.) broad near the tips, rather membranous, obtuse or subacute slightly apiculate, gradually narrowed into rather broad petioles about as long as the blades; midrib evident, otherwise nerveless: cauline leaves few, rather distant, sessile or the lowermost shortly petiolate, linear-lanceolate, acute or subacute, about half as long as the radical. Racemes compactly branched, branches short; flowers crowded, fairly numerous, almost sessile, 9 mm. (⅜ in.) long, white. Calyx deeply 5-partite, lobes linear acute, sparsely clothed with more or less spreading hairs; corolla funnel-shaped, the tube rather wide and twice as long as the calyx, lobes spreading shortly oblong obtuse; stamens slightly shorter than the corolla-tube, reaching just to the level of the prominent throat-scales; anthers short, broadly linear, filaments about half as long as the anthers; style as long as the corolla and elongating after flowering. Mature nutlets not seen.
On dry rocks, Shingly Range, Awatere Basin, Marlborough; about 4,000 ft.
For the opportunity of examining this plant I am indebted to Dr. L. Cockayne, F.R.S., F.L.S., of Wellington. Perhaps its closest ally is my M. oreophila.
3. Myosotis diversifolia sp. nov.
Species M. petiolatae Hook. f. affinis; differt caulibus erectis, foliis valde membranaceis, caulinis acutis, pilis foliorum culmorumque longioribus tenuloribus vix rigidis minus arcte appressis, pedicellis brevioribus, corollae tubo duplo longiore, antheris angustioribus subapiculatis.
Perennial; stems several (usually three or four) from the top of the moderately stout root, slender leafy to near the base of the racemes, simple or forked at the point of origin of the inflorescence, rather sparingly clothed with soft spreading white hairs, 8 in. (20 cm.) high or less. Leaves very membranous, sparsely clothed on both surfaces and along the edges with soft loosely appressed rather long delicate hairs, with a conspicuous rather fine submarginal nerve running all round; radical ± 2 ½ in. (6.3 cm.) long, blade elliptic 1 ⅛ in. (2.8 cm.) long, ¾ in. (2 cm.) broad, apiculate, sharply contracted into rather narrow softly pilose petioles somewhat longer than the blades; cauline progressively smaller and narrower, distant (internodes ½–¾ in. long), the lower shortly petiolate, the upper sessile by a broad base, all acute and apiculate. Racemes ± 3 in. (7.5 cm.) long, simple or forked many-flowered Flowers ¼ in. (6–7 mm.) long, white, shortly pedicelled
(pedicels shorter than the calyx), ebracteate, crowded in the earlier flowering stage; calyx ± ⅙ in. (4 mm.) long, cut half-way down into narrow ovate acute strongly ciliated lobes with short spreading hooked hairs at the basal part; corolla funnel-shaped, tube about twice as long as the calyx, narrow below, cut above into rather short rounded lobes; scales of the throat short and broad; stamens inserted a little below the scales, reaching as high as the clefts of the corolla-lobes; anthers narrow, scarcely elongated, almost apiculate, about three times as long as the very slender filaments to which they are attached a little below the middle; style short, scarcely exceeding the calyx even in fruit; nutlets (scarcely mature) pale brown, suborbicular.
Hob.—Ruahine Mountain-range, above the forest-belt.
Collected by Mr. H. Hill, B.A., of Napier, to whom I am indebted for specimens.
Mr. Cheeseman (Manual, p. 468) referred this plant to M. petiolata Hook. f. The latter is a coastal form, and it is doubtful if it ever grows inland or at considerable elevations. In my view M. petiolata is a purely coastal form, the montane plants referred to it probably belonging elsewhere. I am not certain of the colour of the corolla when fresh.
4. Myosotis tenericaulis sp. nov.
M. annua(?) M. spathulatae Forst. f. affinis; differt caulibus primo suberectis demum ± late diffusis, 20–30 cm. longis, valde tenuibus flaccidisque; caulibus foliisque cineraceis; floribus minoribus; corollae tubo limbi divisuras bis terve superante; antheris anguste oblongis subapiculatis filamenta excedentibus, vix ad faucis squamas breves latas manifestas pertinentibus; nuculorum integumentis pallide flavidis, nucleo atriore per integumenta ± pellucida manifesto.
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Annual (?); stems numerous from the root, 20–30 cm. (1 ft. or less) long, suberect below, spreading and diffuse above, sparingly and distantly branched, branches elongated (internodes long), very thin and flaccid, like the leaves ashy-grey, slightly hispidulous with short appressed whitish hairs, rarely nearly glabrous. Leaves very thin, often subapiculate, with evident midrib, sparsely sprinkled with short appressed whitish hairs, the lower surface sometimes nearly glabrous; radical 3.5 cm. (± 1 ¼ in.) long, the blades elliptic-obovate contracted into slender petioles nearly twice as long as the blades; cauline distant, smaller, narrow-obovate, sessile by a broad base, obtuse or the uppermost subacute. Flowers solitary distant, usually opposite the axils of the upper cauline leaves, on very slender pedicels about as long as the calyx; calyx narrow ± 2 mm. (about 1/14 in.) long, cut nearly to the base into linear subulate segments, sparingly hispidulous with the usual appressed hairs; corolla narrow funnel-shaped, the tube about one-half longer than the calyx, lobes broadly rounded about one-third as long as the tube; throat-scales short and broad; stamens narrow-oblong subapiculate, scarcely reaching to the level of the scales; filaments very short, affixed to the anthers near the base; style slender, equalling the corolla, and scarcely elongating in fruit. Nutlets shining, pale yellow especially at the thin margins, elsewhere brownish owing to the deeper colour of the nucleus of the seed showing through the rather pellucid integuments.
Hab.—Inch-Clutha, Clutha County. The exact locality where this plant was collected can be easily found. It is about a mile from the Romahapa
station on the Catlins River railway line, where the line crosses the Puerua Stream and enters the alluvial plain of Inch-Clutha. It grew in moist spots alongside the creek.
Mr. Cheeseman (Manual, p. 467) doubtfully referred this plant to M. spathulata Forst. f., which is its nearest ally, though not a very close one. He further mentions that it has been collected by the late Mr. T. Kirk near Winton, Southland County. It appears to be confined to the southern lowlands of Otago.
5. Myosotis macrantha Hook. f. var. westlandica var. nov.
A forma typica differt foliis radicalibus longioribus multo tenuioribus anguste obovatis molliter pilosis, venis a costa ad venam submarginalem conspicuam oblique progredientibus, culmis longioribus gracilioribusque, floribus flavis.
In a moist shady ravine on Rangi Taipo, Jackson's, Taramakau River; about 4,000 ft.: L. Cockayne, D. P.
When better known this form may prove to be a distinct species; for the present it seems better to rank it as a variety.
6. Pterostylis areolata sp. nov.
Gracilis glabra ± 15 cm. alta. Folia pauca, caulina, sessilia, culmum amplectentia; inferiora scariosa squamiformia; superiora (plerumque 3) valde tenuia, in siccitate pellucida, lanceolata v. oblongo-lanceolata, acuta v. subacuta, plurinervia manifeste areolata, 3.5–4 cm. longa 1 cm. lata; culmo folium summum longe excedente. Flores solitarii 3.5 cm. longi ± 1.5 cm. lati. Galea pro parte majore erecta, pro parte tertia recurva; sepalum superius in apicem brevem acutum haud filiformem desinens, petalis acutis paulo longius; labii inferioris divisurae anguste obcuneatae, in apices subulato-filiformes summam galeam haud excedentes desinentes; labium subcrassum lanceolato – oblongum subacutum; columna gracilis elongata galeae partem erectam aequans.
Slender, glabrous, ± 15 cm. (6 in.) high. Leaves 4 or 5 (in the specimens seen), rather distant, sessile and sheathing the stem; the lower reduced to scarious sheathing scales; the upper very thin, pellucid when dried, 3.5–4 cm. (± in.) long, 1 cm. (± ⅜ in.) broad, lanceolate or oblong-lanceolate, acute or subacute, entire, narrowed towards the base, with conspicuous veins running nearly straight along their whole length and connected by delicate more or less oblique veinlets into an open network; the uppermost leaf placed about half-way up the stem and reaching about half-way up to the flower. Flowers solitary, 3.5 cm. (± 1 ½ in.) long, ± 1–5 cm. (⅝ in.) broad, green more or less streaked with reddish-brown; galea erect for two-thirds its length, then sharply bent forwards; upper sepal ending in a short more or less acute non-filiform tip, a little longer than the acute petals; lower lip narrow-cuneate for nearly half its length, forking widely into narrow obcuneate subulate-filiform-tipped lobes that do not exceed the top of the galea; lip brownish when dried, rather thick and firm, lanceolate-oblong, subacute with exserted tip; column slender, as long as the erect part of the galea, the lower lobe of its wings large long obtuse.
Hob.—Base of Shingle Peak, Awatere Valley, Marlborough; 3,000 ft.; in shade: L. Cockayne! Bealey, Waimakariri Valley, Canterbury: T. Kirk!
This appears to be a well-marked species. The late Mr. Kirk referred his specimens, which are in fruit and are rather stouter than Dr. Cockayne's, to P. micromega Hook, f., but they are destitute of radical leaves, while the cauline leaves are much larger and broader than those of P. micromega, and do not extend above the middle of the stem. As I have seen, only dried specimens, the details of the structure of the column may be imperfectly sketched here.
7. Poa campbellensis sp. nov.
Species P. pusillae Berggren affinis: differt foliis numerosis erectis v. suberectis conduplicatis apice obtusis; ligulis longioribus ± oblongis laceris v. erosis; spiculis paulo majoribus viridibus colore purpureo-spadiceo ± tinctis; glumis florigeris subacutis basi pilis crispulis brevibus exigue instructis; paleis apice subalte bifidis ac a carinis delicate ciliatis.
Culms very slender densely tufted, leafy below and usually clothed by the sheaths of the cauline leaves to above the base of the panicle, 5–10 cm. (2–4 in.) rarely 15 cm. (6 in.) high. Basal leaves much shorter than the culms, erect or slightly spreading, narrow blunt-pointed, smooth, folded, rather stiff; sheaths about as long as the blades, broad, thin, loose, membranous, striate; ligules variable in length, shortly oblong (rarely longer and narrowed upwards), thin and scarious, erose or lacerate at the tops.' Panicle small, 2.5–4 cm. (1–1 ½ in.) long, narrow-ovate, of 6–9 spikelets placed on rather long glabrous or slightly scabrid pedicels; branches capillary, the lower much longer. Spikelets ± 7 mm. (¼ in.) long, ovatelanceolate, greenish, faintly stained with purplish-brown, 2–3-flowered; outer glumes slightly unequal, about two-thirds as long as the flowering-glume immediately above, smooth or slightly scabrid along the keel; the lower narrow-ovate acute faintly 3-nerved, the upper broader subacute strongly 3-nerved; flowering-glumes ovate-oblong, subacute, thin, more or less scarious around the tops, smooth except on the finely scabrid keel, with a scanty tuft of delicate crisped hairs at the callus, 5-nerved; the two lateral nerves faint, the median nerve alone reaching the top; palea a little shorter than the flowering-glume, rather deeply bifid at the top, finely ciliate along the nerves.
Hob.—Campbell Island, and Port Ross in the Auckland Islands: B. C. Aston! (January, 1909).
The Port Ross specimens are considerably taller than those from Campbell Island. In my report on the Gramina in vol. ii of The Subantarclic Islands of New Zealand this grass was united with my Poa incrassata. I am now satisfied that this treatment of the plant is incorrect. The original description (Trans. N.Z. Inst., vol. 34, p. 394, 1902) and that given in Mr. Cheeseman's Manual (p. 911) are therefore the correct ones. Mr. Cheeseman has noted that Poa incrassata is most nearly allied to Poa exigua Hook. f. The present species has its nearest ally in Poa pusilla Berggren.