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Volume 53, 1921
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Ischyrocerus anguipes Kröyer.

Podocerus cylindricus Kirk, 1879, p. 402. Wyvillea longimana Haswell, 1879, p. 337, pl. 22, fig. 7; Stebbing, 1906, p. 648. Podocerus longimanus Chilton, 1884, p. 255, pl. 17, fig. 2 a—e. Ischyrocerus anguipes Sars, 1894, p. 588, pl. 209; Stebbing, 1906, p. 658.

This species was first recorded from New Zealand by T. W. Kirk in 1879 from specimens collected at Worser Bay, Wellington, which were by him identified as Podocerus cylindricus Say, the identification, however, being subsequently questioned by Miers (1880, p. 125). In the same year Haswell had described Wyvillea longimana from Port Jackson, establishing for it the new genus Wyvillea. In 1884 I identified specimens taken at Lyttelton as being the same as Haswell's Wyvillea longimana, and pointed out that his generic description had apparently been based on a misinterpretation of the terminal uropods, and that the animal in question was the same as the specimens referred by Kirk to Podocerus cylindricus, which I had been able to examine. Owing, however, to Miers's doubt as to the possibility of an Arctic species being found also in New Zealand, I adopted Haswell's specific name, and therefore named the species Podocerus longimanus. In 1888 Stebbing in his notice of Haswell's paper says, “The

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Picture icon

Fig. 4.—Jassa frequens (Chilton).
A. Second gnathopod of male.
B. Fourth peraeopod of the same
C. Second gnathopod of female.
D. Fourth peraeopod of same.

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figure which Mr. Haswell gives much resembles Ischyrocerus (Podocerus) anguipes Kröyer. Mr. Chilton supposes that the description given of the pleopoda [uropoda] is the result of an oversight, and that the genus must be cancelled in favour of Podocerus. It must, however, be observed that Mr. Haswell's description of the maxillipeds is quite inconsistent with this conclusion.” In Das Tierreich Stebbing (1906, p. 648) retains the genus Wyvillea Haswell, describing the maxillipeds as “exunguiculate, inner and outer plates rudimentary, palp three-jointed,” and to this genus he ascribes two species—viz., W. longimana Haswell and W. haswelli (G. M. Thomson). This description of the maxillipeds must, I think, be based on Haswell's original description, which was apparently incorrect. In the specimens from Lyttelton, which I feel sure are rightly referred to Haswell's species,* the maxillipeds are normal and closely resemble the figure given by Sars for Ischyrocerus anguipes. I have also been able to compare my specimens with an Arctic one from Davis Strait sent to me by Dr. W. T. Calman, and have no hesitation in identifying them both as belonging to the one species. I have already pointed out (1920, p. 6) that the other species, Wyvillea haswelli (G. M. Thomson), is a species of Eurystheus. In this the maxillipeds are also normal. Consequently the genus Wyvillea must be finally dropped.

The Lyttelton specimens are all rather small, the largest about 6 mm. long; but those examined by Kirk were very much larger, the second gnathopod (now in my collection) of one specimen being itself 5 mm. long. Stebbing gives the length as varying from 4 mm. to 15 mm. The Davis Strait specimen that I have examined is about 12 mm. in length.

The differences between the male and female, as pointed out by me in 1884 from New Zealand specimens, closely agree with those described and figured by Sars in 1894. The special characters of the second gnathopod of the male are only acquired when the animal is fully adult, the immature stages being at first similar to those of the female. I have one immature male specimen in which the gnathopod closely resembles the figure given by Sars of Ischyrocerus minutus Lilljeborg; a species which Stebbing considers a synonym of I. anguipes Kröyer.

Ischyrocerus anguipes has been recorded from South Africa by Barnard, and is another example of an amphipod first described from northern seas which proves to be also widely distributed in the Southern Hemisphere.

[Footnote] * Since this was printed specimens quite similar to those from Lyttelton have been sent to me from Coogee, close to Port Jackson, New South Wales, the type locality for Podocerus longimanus Haswell.