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Volume 54, 1923
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III. The Axinea Group.

Glycymeris trelissickensis n. sp. (Plate 5, fig. 4; Plate 6, fig. 4.)

Shell of moderate size, very inflated, almost globular; beaks low, with a very wide angle; outline equilateral, roundly trigonal in youth, longitudinally oval in the adult; anal depression obsolete; sculpture, surface appearing almost smooth, but there are fine close radial riblets over the greater part of the disc, crossed by growth-lines; underneath these, and shown on weathering, is a system of narrow radial ribs; inner margin finely crenate, ligamental area small, closely striated by 4 or 5 grooves (a paratype of advanced age shows a wide area with 12 grooves); teeth small, numerous, about 6 on each side free of the area; muscular impressions subequal.

Height of holotype (right valve), 42 mm.; length, 45-5 mm.; thickness, 16 mm.

Holotype in collection of N.Z. Geological Survey.

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Localities:—

  • 239—“ Fan coral bed, Porter and Thomas Rivers, Trelissick Basin ” (McKay, 1879). (Holotype.)

  • 237—“ Upper part Mount Brown limestone,” Trelissick.

  • 238—“ Mount Brown limestone, Coleridge Creek,” Trelissick.

  • 449—“ Lower beds, Trelissick Basin ” (Enys, 1880).

The matrix and preservation of this shell are similar to that of the holotype, so it is probably from the same horizon, and not from lower beds as, stated in McKay's manuscript.

The species is distinguishable from the others of the group by its wider beaks and greater inflation towards the margins.

Glycymeris waipipiensis n. sp. (Plate 5, fig. 5; Plate 6, fig. 5.)

Shell very large, regularly inflated, beaks wide and prominent, outline almost circular, anterior regularly rounded, posterior slightly straightened above; sculpture consisting of very fine radial riblets upon a system of obsolete narrow ribs, about 60 in number; waved growth-lines are prominent towards the margin; inner margin very finely crenate; ligamental area moderate, closely grooved: teeth” small,. 7 on each side free of the area; anterior muscular impression only slightly larger than the posterior.

Height of holotype (complete individual), 77 mm.; length, 79 mm.; thickness of one valve, 27 mm.

Holotype in the collection of Dr. Marshall.

Localities:—

Waipipi (Marshall). (Also collected by Murdoch.)

996—Kaawa Creek (small specimens). (Also collected by Bartrum.)

This species is much larger than any of the others in the Axinea group, described in this paper. It is intermediate in character between G. trelissickensis and G. shrimptoni. Small specimens would be distinguishable from the former by their more prominent and narrower beaks, and from the latter by their slightly stronger beaks and higher shoulders.

Glycymeris shrimptoni n. sp. (Plate 5, fig. 3; Plate 6, fig. 3.)

Shell of moderate size, inflated, beaks low; outline almost circular; anal depression obsolete; sculpture—the whole disc is covered with fine closely—placed radial riblets, about 6 per millimetre, superimposed upon narrow obsolete ribs each bearing about 10 of the lines at the margin, near which especially there are concentric growth-lines; inner margin finely crenate; ligamental area moderate, closely striated by 8 grooves; teeth small, 7 on each side free of the area; muscular impressions subequal.

Height of holotype (left valve), 44 mm.; length, 43 mm.; thickness, 15 mm.

Holotype in collection of N.Z. Geological Survey.

Localities:—

191—Shrimpton's, Ngaruroro. River, Hawke's Bay. (Holotype.)

? 231—McLean's, Ngaruroro River. (Very much worn.)

This species is closely related to G. trelissickensis, but the beak is a shade higher and has a narrower angle; there is also less inflation towards the margins.

Some specimens develop a certain amount of obliquity, but they can always be separated from such species as G. callaghani and O. waitakiensis by their wider shoulders.

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In N.Z. Geol. Surv. Pal. Bull. No. 8, p. 19, this species is listed as G. striatularis (see below, p. 77).

Glycymeris waitakiensis n. sp. (Plate 5, fig. 2; Plate 6, fig. 2.)

Shell of moderate size, fairly solid, roundly trigonal almost equilateral, inflated; beaks broad, prominent, but only moderately projecting above the hinge-margin; anterior end steeply sloping above, broadly rounded below; posterior end similar, slightly attenuated ventrally; sculpture- the surface is almost smooth but weathering shows very narrow radials corresponding to the fine crenulations of the inner margin; ligamental area short and broad, closely striated by 7–8 grooves; teeth small and fine, about 12 on each side, 6 of which are free of the area.

Height of holotype (left valve), 39 mm.: length, 40–5 mm.; thickness, 15 mm.

Holotype in collection of N.Z. Geological Survey.

Locality.—483—Hutchinson's Quarry greensand, Wharekuri (McKay).

Material, one specimen, in very good preservation.

Remarks.—This shell may be distinguished from G. trelissickensis by its trigonal shape, and from G. callaghani by its approach to an equilateral disposition.

Listed by Suter in N.Z. Geol. Surv. Pal. Bull. No. 8, p. 69, as G. cordata (?), in Mr. Uttley's collection.

Glycymeris. callaghani n. sp. (Plate 5, fig. 1; Plate 6, fig. 1.)

Shell large, solid, inequilateral, obliquely ovate, inflated; beaks moderate, narrow: anterior end semicircular; posterior end attenuated ventrally: sculpture-surface covered with very fine radial riblets, underlain by obsolete primary ribbing, showing plainly on weathering, these ribs are about 50 in number; inner margin with crenulations of moderate size; ligamental area short but wide, closely striated by 8–10 well-incised grooves; teeth fairly strong; muscular impressions subequal, the posterior with a very strong ridge in front of it.

Holotype in collection of N.Z. Geological Survey.

Height of holotype (right valve), 50 mm.; length, 51 mm.; thickness, 18 mm.

Localities:—

154 (in part)—Greensand, Callaghan's Creek, Kanieri district (N.Z. Geol. Surv. Rep. No. 9, 1877, p. 84). (Holotype.) (Collected also by Mr. Bartrum.)

649—Paparoa Rapids, Wanganui River, (The beaks of these specimens are broader than those of the type.)

? 899—Taumatamaire, Awakino. (A small specimen filled with matrix.)

This species may be distinguished from others of the group by its obliquity. The outline is like that of G. manaiaensis of the globosa group, but the fine ornamentation and closely grooved area easily distinguish it.

Dr. Henderson kindly procured some Callaghan's Creek specimens from the Reefton School of Mines. These show the interior clearly. The ridge in front of the posterior muscular impression is stronger than in any other species of the genus.

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Glycymeris modesta (Angas). (Plate 5, fig. 6; Plate 6, fig. 6.)

  • 1879. Axinea modesta Angas, Proc. Zool. Soc, p. 418, pl. 35, fig. 4.

  • 1873. Pectunculus striatularis Lam.: Hutton, Cat. Ter. Moll., p. 28 (not of Lamarck).

  • 1908. Glycymeris velutina Suter, Trans. N.Z. Inst., vol. 40, p. 354, pl. 30, figs. 1, 2.

  • 1913. Glycymeris modesta. (Angas): Suter, Man. N.Z. Moll., p. 852.

For further synonymy and description see Suter's Manual. It must be remembered that Suter mistook the anterior for the posterior of the shell, so his description of the beaks should read “ slightly curved backwards,” and so on.

  • Localities:—

  • Holotype, Recent.

  • 207, 208, 768—Shakespeare Cliff, &c, Wanganui.

  • 720—Upper band, Petane limestone.

  • 191—Shrimpton's, Ngaruroro River, Hawke's Bay.

  • 996—Kaawa Creek, Auckland. (Small specimens.) (Also collected by Bartrum.)

The lowest horizon in the Wanganui district recorded for this shell by Marshall and Murdoch is Nukumaru.

Beyond the characters of the “ group,” this shell shows no close affinities with any other New Zealand shell, and, as the original stock apparently became established in this area about the .commencement of the Wanganuian, it would be interesting to know from whence it came. Suter mentions its occurrence at the Kermadec Islands; but it is omitted by Oliver (1915, p. 567) in his list of mollusca from that locality.