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Volume 54, 1923
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Genus Hudsonia n. gen.

Allied to Macromastix, and agreeing with that genus in its venation and its non-verticillate antennae, but (1) the antennae are reduced in both sexes, both in length and in the number of joints, the reduction being greater in the female; (2) the male hypopygium is large and normal in position, not small and twisted to one side as in Macromastix; (3) the ovipositor has long horny anal valves as in Tipula, Holorusia, Longurio, &c.; (4) in both sexes the sensory pits on the praescutum (pseudosutural foveae of Alexander) are larger than in Macromastix; (5) the palpi are short, the fourth joint scarcely longer than the third.

This genus is erected for the single species Tipula heterogama Hudson, which cannot be satisfactorily placed in Macromastix, on account of the horny ovipositor; in Holorusia, on account of the venation of the male; or in Longurio or Tipula, on account of the absence of verticils on the antennae. I have much pleasure in dedicating the genus to Mr. G. V. Hudson, the discoverer and describer of the type species. I do not consider the name Hudsonia to be preoccupied by Hudsonius Gir., 1856 (Pisces).

The reduction of the female antennae is very interesting, and seems to foreshadow the somewhat similar condition in the Brachycera. I do not know of another case among the Nematocera in which reduction of the wings is accompanied by reduction of the antennae.

Hudsonia heterogama (Hudson). (Fig. 122.)

Tipula heterogama Hudson, Trans. N.Z. Inst., vol. 45, p. 68, 1913.

♂. Head ochreous, greyish round eyes, with some dark hair, especially noticeable on the moderately large and somewhat bifid frontal tubercle. Front a little over a quarter as broad as head. Rostrum shorter than head, with broad rather long nasus. Antennae brownish-ochreous, tips of second and third joints pale. First scapal joint rather short, not much more than twice as long as the second. Flagellum with eight joints; first about three times as long as broad, second equally broad but rather shorter, third narrower and only about half as long as second; the remaining five all minute and together shorter than second and third together. Palpi not much longer than rostrum, brown; last joint not much longer than third. Thorax brownish, dusted with grey, except on the four somewhat shining dark reddish-brown praescutal stripes. Pronotum and interspaces of praescutum with some dark pubescence. Abdomen rather brightly shining, reddish-brown, apical portion of segment 2 and segments 3–5 darker. Segment 2 more than twice as long as segment 3. Pubescence rather spare, except on tergites 8 and 9, where it is shorter and much denser, and points forwards instead of backwards. Hypopygium as long as segments 6–8 together and slightly broader; ninth tergite considerably narrower than side-pieces, and only half as long, with a small median V-shaped emargination, corners somewhat produced; claspers large; reaching back over margin of ninth tergite, short-haired, without strong spines, an upper

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flattened lobe covering a lower narrower ones. Legs rather stout, ochreous-brown, tips of femora and tibiae and last two or three tarsal joints blackish. Last tarsal joint cylindrical, unmodified. Wings with slight brown tinge, more obvious in costal cell; stigma shortly oval, dark brown. Sc1 present, oblique (as in several species of Macromastix). Rs very short, curved, slightly longer than R2+3, which is only half as long again as R2; r meeting R2 near the middle; stem of cell M1 about one-third as long as cell; cell Ax rather narrow, of even width throughout. Halteres rather light brown. Length of body, 13 mm.; wing, 18 mm.; hind leg—femur 10 mm., tibia 10·5 mm., tarsus 20 mm.

♀. Differs from the male as follows: Front slightly broader, quite one-third as broad as head. Antennae further reduced, flagellum club-shaped, very narrow at base, hardly any longer than scape, composed of five indistinctly separated joints, of which first is as long as the other four together. Thorax narrow, no broader than base of abdomen, dark brown, unstriped, with hardly any grey dusting. Abdomen blackish, except towards tip, second segment not much longer than third. Ovipositor long and horny; anal valves straight, stout, with rounded tips, longer than rather long anal segment; genital valves also long, straight, reaching beyond middle of anal valves. Legs much shorter, stouter, with slightly longer pubescence; front femora somewhat stouter than others. Wings reduced to narrow rudiments. Halteres blackish, twisted, apparently in the first stage of degeneration. Length of body, 16 mm.; wing, 2 mm.; hind leg—femur 5 mm., tibia 5·2 mm., tarsus 5·5 mm.

North Island: Wellington district, April—May (G. V. Hudson, No. 170).

The male bears a rather striking resemblance to Macromastix zeylandiae Alexander, and it is not at all surprising that Hutton should have considered the two as varieties of one species. H. heterogama may be a development from this group of Macromastix.

Plate 27.

  • Fig.

  • 1. Anisopus notatus (Hutton), ♂. × 6.

  • 2. Anisopus notatus (Hutton), ♀. × 6.

  • (From Lamb's type of Rhyphus undulatus.)

  • 3. Anisopus neozelandicus (Schiner), ♀. × 6.

  • 4. Tanyderus annuliferus Hutton, ♂. × 3.

  • 5. Tanyderus annuliferus Hutton, ♀. × 3.

  • 6. Tanyderus neptunus n. sp., ♂. × 3.

  • 7. Tanyderus forcipatus O.-S., ♀. × 3.

  • 8. Tanyderus varipes n. sp., ♂. × 3.

  • 9. Tanyderus varipes n. sp., ♀. × 3.

  • 10. Dicranomyia flavidipennis n. sp., ♂. × 5.

  • 11. Dicranomyia sublacteata n. sp., ♂. × 5.

  • Fig.

  • 12. Dicranomyia decincta n. sp., ♂. × 5.

  • 13. Dicranomyia melanogramma n. sp., ♂. × 5.

  • 14. Dicranomyia cubitalis n. sp., ♂. × 5.

  • 15. Dicranomyia huttoni n. name, ♀. × 5.

  • 16. Dicranomyia crassipes n. sp., ♂. × 5.

  • 17. Dicranomyia repanda n. sp., ♀. × 3.

  • 18. Dicranomyia fumipennis (Butler), ♂. × 3.

  • 19. Dicranomyia hudsoni n. sp., ♀. × 3.

  • 20. Dicranomyia tenebrosa, n. sp., ♂. × 3.

  • 21. Dicranomyia conulifera n. sp., ♂. × 5.

  • 22. Dicranomyia aegrotans n. sp., ♀. × 6.

  • 23. Dicranomyia conveniens (Walker), ♂. × 6.

  • 24. Dicranomyia luteonitens n. sp., ♀. × 6.

  • 25. Dicranomyia gracilis n. sp., ♂. × 6.

Plate 28.

  • 26. Discobola ampla (Hutton), ♂. × 3.

  • 27. Discobola dohrni O.-S., ♀. × 3.

  • 28. Discobola dicycla n. sp., ♀. × 3.

  • 29. Discobola gibbera n. sp., ♀. × 3.

  • 30. Discobola venusta (O.-S.), ♂. × 3.

  • 31. Discobola picta (Hutton), ♀. × 3.

  • 32. Discobola striata n. sp., ♂. × 3.

  • 33. Discobola tesselata (O.-S.), ♀. × 3.

  • (From the type.)

  • 34. Discobola tesselata O.-S., ♀. × 5.

  • 35. Elephantomyia zealandica n. sp., ♂. × 6.

  • 36. Ceratocheilus levis (Hutton), ♀. × 6.

  • 37. Ceratocheilus ochraceum n. sp., ♀. × 6.

  • 38. Gonomyia (Lipophleps) nigrohalterata n. sp., ♀. × 6.

  • Fig.

  • 39. Amphineurus insulsus (Hutton), ♀. × 6.

  • 40. Amphineurus fatuus (Hutton), ♂. × 6.

  • 41. Amphineurus perdecorus n. sp., ♀. × 6.

  • (Denuded wing, to show venation)

  • 42. Amphineurus perdecorus n. sp., ♂. × 2·5. (Showing markings.)

  • 43. Amphineurus hudsoni n. sp., ♀. × 5.

  • 44. Amphineurus horni n. sp., ♂. × 9.

  • 45. Molophilus pulcherrimus n. sp., ♀. × 9.

  • 46. Molophilus variegatus n. sp., ♂. × 9.

  • 47. Gnophomyia (Aphrophila) neozelandica n. sp., ♀. × 3.

  • 48. Gnophomyia (Astelobia) rufa (Hudson). × 3.

Plate 29

  • 49. Orolimnophila eluta n. sp., ♂. × 3.

  • 50. Tinemyia margaritifera Hutton, ♂. × 3.

  • 51. Rhamphophila (n. gen) obscuripennis (Hudson), ♂. × 3.

  • 52. Rhamphophila lyrifera n. sp., ♂. × 2.

  • 53. Ulomorpha nebulosa n. sp., ♂. × 3.

  • 54. Ulomorpha fuscana n. sp., ♂. × 7.

  • 55. Limnophila leucomelas n. sp., ♂. × 7.

  • 56. Limnophila hudsoni n. sp., ♂. × 3.

  • 57. Limnophila marshalli Hutton, ♂. × 3.

  • 58. Limnophila chryser hoea n. sp., ♂. × 3.

  • 59. Limnophila nigrocincta n. sp., ♀. × 3.

  • Fig

  • 60. Limnophila cyatheti n. sp., ♂. × 2.

  • 61. Limnophila geographica Hutton, ♂. × 3.

  • 62. Limnophila crassipes Hutton. × 2.

  • 63. Limnophila oculata n. sp., ♂. × 2.

  • 64. Limnophila lambi n. sp., ♂. × 3.

  • 65. Limnophila argus Hutton, ♀. × 3.

  • 66. Limnophila delicatula Hutton, ♀. × 3.

  • 67. Limnophila subinterventa n. sp., ♂. × 3.

  • 68. Limnophila skusei Hutton, ♂. × 2.

  • 69. Limnophila skusei Hutton var, ♂. × 3.

  • 70. Limnophila skusei Hutton var.,

Plate 30.

  • Fig.

  • 71. Gynoplistia magnifica n. sp., ♂. × 3.

  • 72. Gynoplistia notata n. sp., ♂. × 3.

  • 73. Gynoplistia nebulosa n. sp., ♂. × 3.

  • 74. Gynoplistia subfasciata Walk., ♂. × 3.

  • (From Walker's type.)

  • 75. Gynoplistia arthuriana n. sp., ♀. × 3.

  • 76. Gynoplistia canterburina n. sp., ♀. × 2.

  • 77. Gynoplistia fimbriata Alex., ♂. × 2.

  • 78. Gynoplistia tridactyla n. sp., ♀. × 3.

  • 79. Gynoplistia speciosa n. sp., ♀. × 3.

  • 80. Gynoplistia sackeni Alex., ♀. × 3.

  • 81. Gynoplistia wakefieldi Westw., ♂. × 3.

  • 82. Gynoplistia conjuncta n. sp., ♂. × 3.

  • 83. Gynophistia bona Alex., ♀. × 3.

  • Fig.

  • 84. Gynophistia cuprea Hutton var., ♂. × 3.

  • 85. Gynophistia cuprea Hutton, ♀. × 3.

  • 86. Gynophistia violacea n. sp., ♀. × 5.

  • 87. Gynophistia nigronitida n. sp., ♂. × 7.

  • 88. Gynophistia polita n. sp., ♂. × 5.

  • 89. Gynophistia trifasciata n. sp., ♂. × 7.

  • 90. Gynophistia clavipes n. sp., ♂. × 7.

  • 91. Gynophistia albicincta n. sp., ♀. × 7.

  • 92. Cerozodia plumosa O.-S., ♂. × 2.

  • (From Osten-Sacken's type.)

  • 93. Cerozodia paradisea n. sp., ♂. × 2.

  • 94. Cerozodia pulverulenta n. sp., ♂. × 2.

  • 95. Cerozodia hudsoni n. sp., ♂. × 2.

Plate 31.

  • Fig.

  • 96. Dolichopeza par vicauda n. sp., ♂. × 3.

  • 97. Holorusia novarae (Schin.), ♀. × 2.

  • (Slight development of stump in cell M.)

  • 98. Holorusia novarae (Schin.), ♂. × 2.

  • (Complete cross-vein in cell M.)

  • 99. Holorusia fulva (Hutton), ♂. × 2.

  • 100. Longurio dux (Hudson), ♂. × 2.

  • 101. Longurio legatus n. sp., ♂. × 2.

  • 102. Macromastix aperta n. sp., ♂. × 2.

  • 103. Macromastix clara (Kirby), ♂. × 2.

  • 104. Macromastix maori Alex., ♂. × 2·5

  • (From Alexander's type.)

  • 105. Macromastix binotata Hutton, ♂. × 3.

  • 106. Macromastix lunata Hutton, ♂. × 3.

  • 107. Macromastix variegata n. sp., ♂. × 3.

  • 108. Macromastix decorata n. sp., ♂. × 3.

  • 109. Macromastix huttoni n. sp., ♂. × 3.

  • 110. Macromastix vittata n. sp., ♂. × 3.

  • Fig.

  • 111. Macromastix alexanderi n. sp., ♂. × 2.

  • 112. Macromastix fumibasis n. sp., ♂. × 2.

  • 113. Macromastix cubitalis n. sp., ♀. × 3.

  • 114. Macromastix albistigma n. sp., ♀. × 2. (A pale specimen.)

  • 115. Macromastix obligua n. sp., ♂. × 2.

  • 116. Macromastix vulpina (Hutton), ♂. × 2.

  • 117. Macromastix rufiventris n. sp., ♂. × 2.

  • 118. Macromastix minoi n. sp., ♂. × 2.

  • 119. Macromastix dichroithorax Alex., ♂. × 2.

  • 120. Macromastix sinclairi n. sp., ♂. × 2.

  • 121. Macromastix zeylandiae Alex., ♂. × 3.

  • 122. Hudsonia heterogama (Hudson), ♀. × 3.

Picture icon

Fig.
123. Tanyderus neptunus n. sp.: penis. × 42.
124. Dicranomyia flavidipennis n. sp.: hypopygium from above. × 42.
125. Dicranomyia decincta n. sp.: hypopygium from above. × 42.
126. Dicranomyia sublacteata n. sp.: hypopygium from above. × 42.
127. Dicranomyia cubitalis n. sp.: hypopygium from above. × 42.
128. Dicranomyia melanogramma n. sp.: hypopygium from above. × 42.
129. Dicranomyia octava n. sp.: hypopygium from above. × 42.
130. Dicranomyia vicarians Schin.: hypopygium from above. × 42.
131. Dicranomyia conveniens (Walk.): hypopygium from above. × 42.
132. Dicranomyia sulphuralis n. sp.: hypopygium from above. × 42.
133. Dicranomyia gracilis n. sp.: hypopygium from above. × 42.
134. Discobola striata n. sp.: hypopygium from above. × 42.

Picture icon

Fig.
135. Amphineurus perdecorus n. sp.: hypopygium from below (tergal view). × 42.
136. Amphineurus bicinctus n. sp.: hypopygium from below (tergal view). × 42.
137. Amphineurus hudsoni n. sp.: hypopygium from below (tergal view). × 42.
138. Amphineurus insulsus (Hutton): hypopygium from below (sternal view). × 42.
139. Amphineurus insulsus (Hutton): hypopygium from below (tergal view). × 42.
140. Amphineurus fatuus (Hutton): hypopygium from below (tergal view). × 42.
141. Molophilus inornatus n. sp.: side-piece and claspers from inner side. × 42.
142. Orolimnophila eluta n. sp.: hypopygium from above. × 42.
143. Ulomorpha nebulosa n. sp.: hypopygium from above. × 42.
144. Tinemyia margaritifera Hutton: hypopygium from above. × 42.

Picture icon

Fig.
145. Rhamphophila obscuripennis (Hunds.): hypopygium from below. × 27.
146. Rhamphophila obscuripennis (Huds.): side-piece from inner side. × 27.
147. Rhamphophila obscuripennis (Huds.): apex of ninth tergite. × 27.
148. Rhamphophila lyrifera n. sp.: hypopygium from below. × 27.
149. Rhamphophila lyrifera n. sp.: process of ninth tergite. × 27.
150. Limnophila skusei Hutton: hypopygium from above. × 42.
151. Limnophila delicatala Hutton: yopopygium from above. × 42.
152. Limnophila leucomelas n. sp.: hypopygium from above. × 42.
153. Limnophila subinterventa n. sp.: hypopygium from above. × 42.
154. Limnophila subinterventa n. sp.: aedoeagus from above. × 42.
155. Limnophila geographica Hutton: hypopygium from above. × 42.
156. Limnophila oculata n. sp.: hypopygium from above. × 27.

Picture icon

Fig.
157. Limnophila lambi n. sp: hypopygium from above. × 42.
158. Limnophila chrysorrhoea n. sp.: hypopygium from above. × 27.
159. Limnophila marshalli Hutton: hypopygium from above. × 42.
160. Limnophila cyatheti n. sp.: hypopygium from above. × 27.
161. Limnophila cyatheti n. sp.: basal lobes of side-pieces from below. × 27.
162. Limnophila hudsoni n. sp.: hypopygium from above. × 27.
163. Gynoplistia subfasciata Walk.: hypopygium from above. × 42.
164. Gynoplistia glauca n. sp.: hypopygium from above. × 42.
165. Gynoplistia fuscoplumbea n. sp.: hypopygium from above. × 42.
166. Gynoplistia incisa n. sp.: hypopygium from above. × 42.

Picture icon

Fig.
167. Gynoplistia tridactyla n. sp.
168. Gynoplistia violacea n. sp.
169. Gynoplistia clavipes n. sp.
170. Gynoplistia trifasciata n. sp.
171. Gynoplistia polita n. sp.
172. Macromastix virescens n. sp.
173. Macromastix viridis (Walk.).
Fig.
174. Macromastix albistigma n. sp.
175. Macromastix holochlora (Now.).
176. Macromastix obliqua n. sp.
177. Macromastix zeylandiae Alex.
178. Macromastix dichroithorax Alex.
179. Macromastix sinclairi n. sp.
180. Macromastix obscuripennis (Kirby).
Hyoppygium from above. × 42.