
Art. 30.—A Contribution to the Study of New Zealand Leaf-hoppers and Plant-hoppers (Cicadellidae and Fulgoroidea).
[Read before the Wellington Philosophical Society, 25th October, 1921; received by Editor 31st December, 1921; issued separately, 30th April, 1923.
Probably few New Zealand insects have been so generally neglected by entomologists as those usually small but abundant forms which, comprise the division Anchenorrhyncha of the Hemiptera-Homoptera. Although the comparatively large and at least audibly conspicuous Cicadidae have received some considerable attention, the other families have been so little collected that a comprehensive monograph of the division is at present out of the question. The present paper is an attempt to bring up to date our knowledge of a few of the subfamilies as a basis fox future taxonomic and biological work.
It will be advisable firstly to indicate briefly the exact connotation of the popular terms “leaf-hopper” and “plant-hopper.” The following classificatory sketch of the Auchenorrhyncha will make this clear:—
| Superfamily 1. Cicadoidea. | |
| Family Cicadidae | Cicadas, or singers. |
| Family Cercopidae | Frog-hoppers. |
| Family Cicadellidae | Leaf-hoppers. |
| Family Membracidae* | Tree-hoppers. |
| Superfamily 2. Fulgoroidea | Plant-hoppers. |
In this paper a revisional account is given of three subfamilies of the Cicadellidae and one family of the Fulgoroidea. The relative value of the families and subfamilies is a debatable matter, in which I have followed Mr. F. Muir, to whom I am deeply indebted for specimens, determinations, literature, and unpublished information, which he has generously and unreservedly placed at my disposal. A list of those who have helped me in New Zealand, particularly with collections, would include most of the working entomologists of the country. Mr. E. H. Atkinson has also collected much valuable material. Since this was written Professor C. F. Baker has been most generous with help and advice.
I. Cicadellidae (Leaf-hoppers).
Syn. Jassidae, Tetigoniidae, auctt.
Of this large family only two species have hitherto been described from New Zealand, of which one is extremely rare, while the description of the other, founded on damaged material, is difficult to identify with any known species. The following key includes all the subfamilies of Cicadellidae known to me to be represented in New Zealand, but new species are continually coming to light. The Cephalelinae, Typhlocybinae, and Tettigoniellinae are dealt with in the following pages; the other three subfamilies will be treated, it is hoped, in a later communication. With regard to the venation I have followed Tillyard in regarding vein 1st A of Metcaif (1913) as really Cu2.
[Footnote] * Not, as far as is known, represented in New Zealand.

| 1 | Head very strongly produced; legs feebly spinose or only bristly | Cephalelinae. |
| Head not strongly produced; hind tibiae strongly spinose | 2 | |
| 2. | Veins of tegmen running without branching to apical crossveins | Typhlocybinae. |
| Veins of tegmen branching on the disc | 3 | |
| 3. | Ocelli situated on dorsal part of head, away from anterior margin | Tettigoniellinae. |
| Ocelli on border of vertex or on margin between vertex and frons | Jassinae. | |
| Ocelli on front, distinctly on ventral side of head | Bythoscopinae. | |
| Ocelli in groove below anterior edge of vertex | Paropiinae. |
Subfamily 1. Tettigoniellinae.
Genus 1. Diedrocephala Spinola. (Fig. 2.)
Diedrocephala cassiniae n. sp. (Fig. 4.)
Short and squat, especially the male. Pale olivaceous brown with whitish streaks and spots. Ocelli pink.
♂. Vertex more than twice as wide as medianly long. A fairly deep depression lateral to each ocellus; a depressed, median longitudinal line; a slightly raised circular area on each side ornamented with more or less concentric striae; several rather deep longitudinal striae on each side of medial longitudinal striation; both anterior and posterior border slightly raised, the former narrowly edged with fuscous. Pale testaceous with three fuscous spots about middle of fore-border and a few yellowish-brown blotches on the disc. Edge between vertex and frons acute. Fore-border of vertex roundly and very obtusely angulate.
Pronotum transversely striate, pale testaceous irregularly marked with yellowish-brown and fuscous, the latter towards posterior margin, which is long and sinuately but slightly emarginate.
Scutellum divided transversely into an anterior striate portion with depression on each side of median longitudinal line, and blunt posterior apical portion very shiny and less striate. Scutellum somewhat shorter than pronotum.
Tip of abdomen barely visible beyond folded tegmina.
Tegmen extremely variegate with white, fuscous (almost black), and brownish. Veins white, edged either interruptedly or continuously with fuscous. Discal portion of each cell yellowish or olivaceous brown. White spots alternating with fuscous spots along the costal margin, each spot extending back to first vein (radius). First apical cell wholly white.
Hind tibiae with two rows of long amber spines with fuscous bases, about ten spines in each row.
Frons elliptical, gently convex. Antennae-pits fuscous. Joints of antennae amber.
Grenitalia fuscous with bands of dark yellow.
Length, 4 mm.
♀. As male, except in following characters: Vertex as long as pronotum. Whole of last ordinary abdominal tergum visible beyond tips of folded tegmina. Often two oblique white bands on each corium.
[Footnote] * These subfamilies are not of equal rank. The Cephalelinae and Typhlocybinae are closely related to the Jassinae

Genitalia — Lateral plates yellowish spotted with fuscous. Ovipositor fuscous, scarcely projecting beyond last unmodified tergite. Dorsal surface of abdomen fuscous. Length, 5 mm.
Nymph. — Varying in colour from brownish - olive to green, thickly sprinkled with short stubbly glistening hairs. A darker depressed spot on each side of each abdominal tergite. Vertex produced much more than in imago; anterior border thin, foliaceous, with fuscous-tipped acute angle projecting cephalad of each eye, and with a row of five or six more or less distinct brownish blotches along anterior edge.
Described from eleven males and nine females.
Common in vast numbers on tauhinu (Cassinia leptophylla) and Olearia Solandri on the sea-coast, Wellington, October, November, June.
Holotype and allotype, with eighteen paratypes, in Myers collection, Biology Laboratory (No. 25).
D. maorica n. sp. (Fig. 6.)
General macroscopic appearance uniform yellowish-testaceous. A very distinct species.
♂. Vertex pale stramineous slightly clouded with yellowish. Median suture strongly marked. Coarsely but sparsely punctate, with a few striations near each eye. Ocelli pink. Eyes uniform fuscous. Vertex about twice as broad as medianly long—if anything, a little less. Fore-border acute but rounded. Edge between frons and vertex very acute.
Pronotum a little longer medianly than vertex. Colour similar to that of vertex. Anterior half with a median striation and a few coarse punctures; posterior half transversely and distinctly striate. Hind-border sinuately but very slightly emarginate.
Scutellum small and short, pale-yellowish, with a brown-tipped, suddenly narrowing but rather long apex.
Abdominal terga uniform pale-olivaceous.
Tegmen narrowing slightly towards tip, which is rounded. Almost uniform pale-yellowish, with indistinct whitish spots along the veins (eight, for example, along claval suture), and one or two cross-veins similarly pale; but all these markings very indistinct.
Wings far less hyaline than in the other species; cells filled with pale-brownish, veins themselves quite clear.
Hind tibiae armed externally with two rows of yellowish spines with brownish bases on an otherwise colourless tibia—eleven in one row and six or seven in the other.
Frons elliptical, pale uniform stramineous, as are also the antennal pits.
Genitalia concolorous with the rest of abdomen and likewise shining.
Length, 5·4 mm.
Described from one male.
Wainuiomata, 12th February, 1921; Mr. G. V. Hudson.
Holotype in Myers collection, Biology Laboratory, Wellington (No. 192).
D. zealandica n. sp. (Fig. 3.)
A large and easily recognized though very variable species. General colour brownish, speckled with fuscous and white.
♂. Vertex almost as long laterally as medianly—i.e., only very slightly, produced; about four times as wide as long. Ocelli red. Fore-border rounded, edge fuscous and slightly upturned. Colour warm brownish, with a few irregular smoother yellowish areas. Very slightly granulate on disc.

Pronotum three times as long as vertex. Anterior portion with a wide shallow depression on each side; transversely sinuately striate; pallid-testaceous, heavily marked with blotches of coffee-brown and irregular spots and streaks of fuscous; fore-border rounded; hind-border with a slight very obtusely angulate emargihation in middle and abrupt angular bend cephalad on each side.
Scutellum brownish with a fuscous spot on each side of base. Apex acute and narrow, nearly as long as pronotum.
Tegmen long, narrower at tip, which is rounded. Testaceous, all veins fuscous, dotted with hyaline white roundish spots, very distinct-about eleven along costal margin and four on termen. A large white spot at end of subcosta connecting by white irregular markings larger than vein-spots to a large white spot in distal angle of clavus. These two spots are very distinct.
Hind tibiae with two rows of fuscous-based amber spines, about nine m each row.
Frons elliptic-oblong, practically plane. Juncture with vertex acute edged venbrally with yellow, followed by an irregular fuscous band notched upward in middle and followed in its turn by a second yellow band curving above antennae-pits.
Length, 6 mm.
♀. Resembles ♂; but vertex is slightly longer and the white areas on tegmen frequently more extensive and less sharply defined, sometimes rendering half the area of tegmen hyaline. The closed tegmina completely cover the abdomen even in this sex. Length, 7 mm.
Described from one male and four females.
Dun Mountain, Nelson, 2,000 2,500 ft., January, March; Mr. A. Philpott! Wellington, January; Mr. H. Hamilton! J. G. Myers.
Holotype and paratypes in Myers collection, Biology Laboratory (No. 190). Allotype in Cawthron Institute collection, Nelson.
D. tararua n. sp.
General colour testaceous or dark greyish-olivaceous. Two oblique white bands on corium, one tipping the tegmen distally.
♀. Vertex considerably less than twice as wide as medianly long; strongly punctate with large shallow punctures. One specimen slightly granulated on disc of vertex. Ocelli pinkish-red. Vertex considerably produced but rounded.
Pronotum — Anterior portion somewhat depressed, with two deeper depressions on each side and a median longitudinal striation extending rather less than third of distance from fore-border. Posterior portion of pronotum somewhat transversely striate, with distinct circular deep dark punctations along each furrow. A median longitudinal area somewhat free of these striae and punctations. Hind-margin sinuate, slightly and roundly emarginate, with indications of four obscure fuscous patches evenly spaced.
Scutellum about three-quarters length of pronotum, which latter is itself co-equal with vertex. Anterior portion of scutellum with a median longitudinal groove meeting a transverse groove at about ⅓ to form an indented inverted T. Posterior portion more elevated, curving suddenly at ⅔ from base of scutellum to form a long, narrow, acute apex which forms a third of total length of scutellum. Whole scutellum with indications of transverse striation.

Tegmen short and broad, apically rounded. Pale-brownish, with the following areas fuscous: two or three irregular patches near base (some- times absent), three more or less confluent spots making an irregular oblique band at half-way, and veins of ante-apical cells also fuscous. At about ⅓ is a more or less hyaline patch, followed by two more very distinct hyaline areas forming a band at ⅔ and at tip respectively. Outer margin of clavus or dorsum of tegmen with two fuscous areas separated by two whitish spots.
Hind tibiae with two rows of dark-amber spines sometimes with dark bases, eight or nine spines in each row.
Frons oblong, slightly convex. Antennae-pits nearly black. Basal joints of antennae yellow.
Genitalia uniformly testaceous or brownish.
Closed tegmina leave exposed the whole of the last unmodified tergite and much of the penultimate.
Length, 5·5–6 mm.
Male unknown.
Described from two females.
Tararua Range (Mount Alpha and Bull Mound), Wellington Province, 3,300–3,600 ft., February.
Holotype and paratype in Myers collection, Biology Laboratory (No. 189).
D. dunensis n. sp.
A small species easily distinguished by the very pale greenish colour almost obsoletely flecked with fuscous, and by the dark eyes.
♀. Vertex twice as wide as medianly long. Median suture distinct. Disc irregularly corrugate and punctate as in D. hudsonica n. sp. A very pale sea-green with a few indistinct smudges of pale brown. Ocelli red and prominent. Eyes almost wholly black, with a slight edging of greenish. Fore-border of vertex with fuscous edging narrower than usual, acutely rounded, but not so much produced as in D. hudsonica. Two small deep pits, one on each side of caudal end of median suture. Actual edge of vertex and frons quite abrupt, but angle not acute.
Pronotum about as long as vertex. Anterior third with a median line interrupted by a deep and wide median depression flanked antero-laterally by two calluses. Posterior portion transversely striate, with fuscous spots in grooves. Disc pale olivaceous-green, with a few brownish smudges. Hind-border very slightly not angulately emarginate, sinuate.
Scutellum large, with a smooth glassy anterior portion separated by a convex line from the short sharp apex, as in D. hudsonica. Disc pale olive-green. Median length about that of pronotum.
Abdomen pale greenish-olivaceous with segmental margins paler.
Tegmen very pale greenish, almost colourless, with very infrequent small fuscous spots scattered irregularly and surrounded narrowly by pale-brownish smudges. Outer edge of clavus with beginning of a broken dark-brownish line. Tip of tegmen somewhat truncate.
Wing as in D. hudsonica.
Hind tibiae armed externally with two rows of spines containing respectively ten and eight.
Frons shield-shaped as in D. hudsonica, with pale stripe on cephalad portion extending along cephalad edge of antennae-pits, which are pale,

edged with brownish. Disc of frons pale olivaceous with brownish lateral transverse striations not meeting across middle.
Genitalia concolorous with rest of abdomen, except ovipositor, which is darker and projects slightly, and ultimate ordinary tergite, which is laterally marked with brownish.
Length, 5 mm.
Described from one female.
Dun Mountain, Nelson, 3,000 ft.; Mr. A. Philpott.
Holotype in Myers collection, Biology Laboratory (No. 197).
D. hinemoa n. sp. (Fig. 5.)
Short and squat, the tegmina and wings small. Tegmen with two white areas separated by a transverse dark patch.
♂. Vertex considerably more than twice as broad as medianly long. Anterior border smooth, slightly upraised, and narrowly bordered with fuscous; disc of vertex irregularly corrugated, with a depression on side near each eye and three longitudinal striae between ocelli, which are dull-pinkish. Vertex uniform olivaceous. Fore-border roundly and obtusely angulate. Edge between vertex and frons acute but passing immediately ventrad into a rounded curve. Eyes variegate, olivaceous and fuscous.
Pronotum irregularly transversely striate; a median transverse depression, shallow, but wide near fore-border; on each side of this a smaller thumb-print-like smooth depression. Pale olivaceous, the above depressions brownish with flecks and spots of castaneous and fuscous irregularly disposed on disc, especially along striae. Posterior border slightly emarginate in an angulately sinuate manner. Pronotum longer than vertex.
Scutellum somewhat shorter than pronotum; finely and evenly corrugate, with transverse line at ⅓ from apex and large cloudy fuscous spot on each side of base.
Abdominal terga—First two pale semitransparent olivaceous; remainder fuscous, almost black, except genitalia, which are paler.
Tegmen—Base with a few cloudy brown marks and spots, especially round bristles; next a wholly white area, in which even veins are not picked out; then follows a yellowish-brown portion in which veins are marked and widely bordered with fuscous and punctations even darker, followed by another colourless area of varying width and irregular shape, after which veins at tips and for a short distance before are widely fuscous, the space between them being brownish-yellow except on termen itself, which is wholly but narrowly edged with fuscous. Anal angle filled with brownish-yellow, the punctations fuscous.

Fig. 1.—Diedrocephala hudsonica n. sp.: female.
Fig. 2.—Diedrocephala sp.: last nymphal instar.
Fig. 3.—Diedrocephala zealandica n. sp.: ventro-caudal view of male pygophor. a, anal segment; v, valve.
Fig. 4.—D. cassiniae n. sp.: ventro-caudal view of male pygophor.
Fig. 5.—D. hinemoa n. sp.: ventro-caudal view of male pygophor.
Fig. 6.—D. maorica n. sp.: ventro-caudal view of male pygophor.
Fig. 7.—D. hudsonica n. sp.: tegmen. } Sc = subcosta; R = radius; M = media;
Fig. 8.—D. hudsonica n. sp.: wing.} Cu = cubitus; A = anal.

Hind tibiae armed externally with two rows of long amber spines with dark bases, eight in one row and nine in the other. Internally a row of bristles.
Frons oblong, very gently convex; olivaceous with numerous semitransverse irregular black marks. Antennae-pits black.
Genitalia olivaceous marked with fuscous. Eighth abdominal sternite yellowish.
Length, 5 mm.
♀. Resembles male except in the following characters: Eyes wholly fuscous except for olivaceous edging. Colour altogether more greenish on head and thorax, with numerous fuscous flecks on vertex. Darker areas of tegmen have occasional or even frequent clear white roundish spots. Ultimate and penultimate ordinary terga mottled with muddy brown. Length, 5·5 mm.
One male, Nelson, January; Mr. A. Philpott. Three females, Mount Arthur Tableland, Nelson, February; Mr. A. Philpott.
Holotype (male) in Myers collection, Biology Laboratory (No. 191). Allotype in Cawthron Institute collection.
D. hudsonica n. sp. (Figs. 1, 7, 8.)
A rather large species with pointed vertex and pale tegmina with a broad chocolate band along the outer border of clavus.
♀. Vertex less than twice as broad as medianly long. Irregularly and coarsely corrugated and punctured. Median suture distinctly marked by several ridges and depressions. Fore-border, as usual, edged narrowly with shining fuscous; acutely rounded. Eyes variegate with fuscous and grey, anterior portion wholly fuscous. Ocelli pinkish with pale centres, each ocellus somewhat on the side of a wide and fairly deep depression. Disc of vertex shining yellowish-brown inclining to olivaceous. Edge between vertex and frons abrupt and fairly acute, but passing immediately ventrad into a rounded curve.
Pronotum about as long as vertex or very slightly shorter; anterior half with almost smooth surface, but with median longitudinal line flanked on each side by a transverse rounded callus, which forms on each side the anterior border of a slight depression. There is a deeper depression lateral to each callus. Posterior portion transversely striate with wide grooves containing occasional fuscous punctations. Disc of pronotum concolorous with vertex. Hind-border slightly emarginate and sinuate with no trace of angularity.
Scutellum large, with a roundly curved anterior surface of pale olivaceous clouded faintly with fuscous, and separated from sharp brownish apex by a curved transverse depressed line with its convexity cephalad. Disc of scutellum with faint indications of irregularly concentric striation. As long as or a little shorter than pronotum.
Abdomen olivaceous, with segmental margins paler; very shining.
Tegmen coarsely punctate, especially along veins. Corium almost immaculate, whitish except for a pale-brownish suffusion on costal portion of basal third, and a few chocolate flecks about middle of claval suture and near bases of ante-apical cells. Clavus with anal angle and a wide irregular band reaching in width more than half-way to suture, dark brown or chocolate; remainder colourless.
Wings with the cells speckled with fuscous, but not heavily enough to mar the transparency. Veins clear.

Hind tibiae armed externally with two rows of yellowish rather short spines with dark bases, about eight in one row and thirteen in the other. Internally a row of bristles.
Frons shield-shaped, with cephalad portion somewhat pointed, passing caudad into a rounded swelling at each side, followed by slight constriction at ⅓, where the typical elliptical curvature is resumed. Pale-olivaceous slightly clouded with pale-brownish. Antennae-pits pale-brownish.
Genitalia concolorous with rest of abdomen, except ovipositor, which is darker and projects slightly beyond ultimate ordinary tergite.
Length, 6·4 mm.
One female.
Karori, Wellington, February; Mr. G. V. Hudson.
Holotype in Myers collection, Biology Laboratory (No. 193).
Subfamily 2. Cephalelinae.
Tribe Dorydini Cogan, Ohio Journ. Sci., vol. 16, p. 184, 1916.
Tribe Cephalelini Kirkaldy, Bull. 3, H.S.P.A. Exp. Sta., p. 72, 1907.
This interesting subfamily is closely related to the Jassinae, of which it is frequently ranked as a tribe. I follow Kirkaldy in elevating it to equal rank with the other primary divisions of the Cicadellidae, although I consider all of them as subfamilies instead of tribes. It is distinct not only on account of the elongate or foliaceous head—a characteristic of widely separated genera in other subfamilies—but also in the “degradation of the flight-organs,” the reduced venation, and the only feebly bristly tibiae. The hind tibiae of Paradorydium westwoodi F. B. White, however, bear, in addition to the bristles, a row of quite considerable spines. In connection with the reduced venation and alar degradation it is noteworthy that in Cephalelus hudsoni n. sp., although the tegmina are well developed and extend considerably beyond the abdomen, the wings are completely atrophied.
It is no exaggeration to say that the members of the two Cephaleline genera represented in New Zealand present as striking examples of Poulton's “constant, special protective resemblance” as are to be found. With regard to the African Cephalelus infumatus Perch., the description of which would apply in essentials to our own species, Osborn noticed that “the protective feature comes in from the fact that the aborted leaf-sheaths on the stem of the plant form sharp spines occurring at intervals along the length of the stem, and these are perfectly reproduced in the form and colour of the insect. So close is the resemblance that when a number of the spines are mounted separately alongside of the insects it is very difficult to distinguish them without the most careful scrutiny.” The plant mimicked by this African species is the rush Dovea tectorum Masters. The stems are green, while the aborted leaf-sheaths are dark brown (Cogan, 1916). The disguise of the New Zealand species is no less perfect.
Of the four species of Cephalelinae so far found in New Zealand, one was described in 1879 by F. B. White, while the other three are new.
| Frons distinctly convex, often even strongly carinate | Paradorydium. |
| Frons concave; deeply excavated | Cephalelus. |

Genus 1. Paradorydium Kirkaldy.
Kirkaldy, Entom., vol. 34, p. 339, 1901.
Paradorydium westwoodi (F. B. White).
Dorydium westwoodi F. B. White, Ent. Mo. Mag., vol. 15, p. 215, 1879; Signoret, Ann. Soc. Ent. France (5), vol. 10, p. 43, 1880; Kirby, Trans. Ent. Soc. Lond., p. 411, 1894; Hutton, Trans. N.Z. Inst., vol. 30, p. 185, 1898. Paradorydium westwoodi Kirkaldy, Entom., vol. 34, p. 339, 1901; ibid., Trans. N.Z. Inst., vol. 41, p. 28, 1909.
F. Buchanan White's description is as follows: “Pale yellowish-ochraceous; head with a very slight tinge of brown; tarsi claws pale brown. Head and thorax finely punctate; tegmina coarsely reticulate-punctate or cribose between the veins. One example has a brown intramarginal streak from the base to the apex of the tegmina. Length, 14; breadth at base of tegmina, 2; length of head, 4 mm. Three specimens from Mr. Wakefield, with the ticket, ‘Pound by Mr. Fereday near Christchurch.’ This curious species strongly resembles the seed of one of the larger grasses.”
By the courtesy of Mr. G. Archey, of the Canterbury Museum, I have been enabled to examine specimens from the Fereday collection and from the Hutton collection.
Fereday's specimen measures 14 mm. in total length, with the head 4·5 mm. long, and agrees exactly with White's description, which may be amplified as follows: Head inclined upwards, but practically straight in itself except for a very slight upturning at the tip, with a faint rounded median longitudinal ridge. Tegmina tapering gradually to a very sharp point. Legs bristly, but in addition the hind tibiae have a row of seven or eight fairly strong spines on the upper outer edge. There is, unfortunately, no date, locality, or number fixed to this specimen.
The Hutton collection contains three specimens almost exactly similar except in size, their lengths being 11 mm., 11 mm., and 12 mm. respectively. The head is also rather more upturned at the tip and inclined to be very slightly spatulate apically. This is a sexual difference in P. menalus Kirk from Australia (Kirkaldy, 1906, p. 339), so that I suspect these are males and the other (Fereday collection) a female; but all four are gummed so tightly by the whole of the ventral surface that T refrained from attempting to remove them for examination of the genitalia. The Hutton specimens have the date 1865, but no locality legible.
This is evidently a rare species. Besides the specimens referred to above, Signoret received some from John Scott, which according to Kirby (1894, 412) appeared to be “darker than the types” and were only 11 mm. long. Finally, a nymph and a male imago were sent to Kirby at the British Museum in 1894, with the following notes by Mr. H. Clarke, of Christchurch: “I found it on the rushes which grow in damp situations. In colour and shape it so much resembles a piece of dried rush that I have never been able to find a specimen except by switching the net amongst them. I think they are scarce and very local, as the specimens I have captured were taken in a space a few yards square, and I have never been able to collect them elsewhere. The place where they were found was in a plantation of Pinus insignis, about a quarter of a mile from the sea and a few chains from the river. The time of appearance is from the beginning of November to the end of the year.”

I quote these local particulars from Kirby (1894) in the hope that collectors in Christchurch will endeavour to rediscover this interesting leaf-hopper.
Since this was written Mr. J. F. Tapley has taken two typical specimens at Governor's Bay, Banks Peninsula, and presented them them to me.
P. philpotti n. sp. (Fig. 9.)
♂. Pale olivaceous, without markings. Scutellum with a conspicuous black spot on each side. Elongate-fusiform, narrowing suddenly cephalad. Head more than half as long as nota and tegmina together, converging suddenly and then tapering to narrowly rounded apex; sides acutely carinate for at least anterior two-thirds. Punctation of head, pronotum, and scutellum almost obsolete. Tegmina coarsely but shallowly punctate, in a linear but inconspicuous manner; much longer than abdomen; convex; not sharply pointed at apex. Eyes pale. Frons and clypeus very convex. Legs relatively longer than in the other species. Ventral surface pale, with deep blackish lateral areas, especially on abdominal sternites. Genital valves uniformly pallid.
Length of body, 8 mm.; head, 3 mm.; greatest width, 1·4 mm.
A very distinct little species, for the only specimen of which, a male, I am indebted to the discoverer, Mr. A. Philpott, after whom I have much pleasure in naming it.
Hump Range, 3,000 ft., December.
Holotype in Myers collection, Biology Laboratory.
Genus 2. Cephalelus Percheron.
Percheron, Mag. Zool., vol. 2, pl. 48, 1832.
Cephalelus hudsoni n. sp. (Figs. 10, 11, 12, 14.)
♂♀. Pale olivaceous varying through greyish-stramineous to yellowish-testaceous and greyish-brown, irregularly mottled with fuscous on sides of vertex, disc of pronotum, and tegmina. Mesonotum often greenish, with a dark spot laterally. Sterna buff, with abdominal sternites, rostrum, and genitalia darker. Genital valves, however, are pallid. The whole surface shows occasional black specks. Shape elongate-fusiform. Head exactly half as long as nota and tegmina together; gradually converging towards apex, which is extremely slightly spatulate. Head concave beneath and convex above. Head, pronotum, and scutellum sparsely and almost obsoletely punctate, except a median longitudinal line on vertex which is smooth. Head exactly porrect. Tegmina with larger, but shallow, dark-coloured punctations. Eyes black, not protuberant. Tegmina much longer than abdomen, tapering rather suddenly to a very sharp point. Wings practically absent, represented by two white membranous flaps less than ⅕ mm. long. Legs short and similar, concolorous with the body. The whole upper surface, when tegmina are folded, is remarkably smooth, convex, rounded, and free from keels. Upper surface of abdomen dark.
Length: Male, total length, 10mm.; head, 3 mm.: female, 11–11·5 mm.; head, 3·7 mm.
Wellington, October to March.
Discovered by Mr. G. V. Hudson, to whom I have much pleasure in dedicating it, and subsequently collected by myself, on a small patch of the jointed rush (Leptocarpus simplex) at Breaker Bay, Wellington.

A second locality is Gollan's Valley, Wellington, also near the sea. I have not found it on any other food-plant, nor even on the same food-plant in other similar localities in the same district. The males appear to be rare: I have only one male among some twenty-eight females; but as my specimens were nearly all collected early in October it is quite possible that the females had appeared first and the males had not reached their maximum of abundance.
As has been noticed in connection with Paradorydium westwoodi and Cephalelus leptocarpi n. sp., the insect is frequently (always, in my experience) confined to a very small patch, though there may be large areas of the food-plant, which grows often in pure associations, in the immediate vicinity.
The adults are rather sluggish, but are capable of leaping some distance, though the posterior legs are apparently very little specialized for this purpose.
The jointed rush (Leptocarpus simplex) has smooth green or reddish internodes, while the fairly frequent nodes are surmounted and partly surrounded by rougher greyish or brownish sheaths. It is to these sheaths the leaf-hoppers bear such an extraordinary resemblance. The insect always rests with its long axis parallel to that of the stem—a position necessary for “purposes” of mimicry, but probably assumed for the sake of comfort.
Occasional examples are parasitized by large, long-legged scarlet mites with long hairs. In one case three of these mites were fastened tightly to the base of one of the tegmina, which, apparently as a result of their sucking, had curled up and twisted like the half of a dried pea-pod. More commonly only one mite is found, and that tucked beneath the tegmina, the abdomen of the Cephalelus being then often slightly deformed, but no other effects apparent.
The nymphs of all ages are found on the food-plant with the adults. They are almost uniform pale brown, with a very indistinct pallid median dorsal line, and a head much less produced than that of the adult. I have not yet succeeded in finding the eggs.
Holotype and allotype in Myers collection, Biology Laboratory.

Fig. 9.—Paradorydium philpotti n. sp.: male genitalia, ventral view.
Fig. 10.—Cephalelus hudsoni n. sp.: male genitalia, ventral view.
Fig. 11.—C. hudsoni n. sp.: nymph, early instar; legs omitted.
Fig. 12.—C. hudsoni n. sp.: nymph, late instar.
Fig. 13.—C. leptocarpi n. sp.: male genitalia, ventral view
Fig. 14.—C. hudsoni n. sp.: head of female.
Fig. 15.—C. leptocarpi n. sp.: head of female.
Fig. 16.—Dikraneura maorica n. sp.: tegmen.
Fig. 17.—D. maorica n. sp.: wing.
Fig. 18.—D. maorica n. sp.: female genitalia.
Fig. 19.—Typhlocyba australis (Frogg.): tegmen.
Fig. 20.—T. australis (Frogg.): wing.
Fig. 21.—T. australis (Frogg.): male genitalia.
Fig. 22.—T. australis (Frogg.): female genitalia.
Fig. 23.—T. australis (Frogg.): imago (after Froggatt).
Fig. 24.—Erythroneura zealandica n. sp.: tegmen. a, costal plaque.
Fig. 25.—E. zealandica n. sp.: wing.
Fig. 26.—E. zealandica n. sp.: male genitalia, lateral view.
Fig. 27.—E. zealandica n. sp.: male genitalia, ventral view.
Fig. 28.—E. zealandica n. sp.: female subgenital plate.
Fig. 29.—E. zealandica n. sp.: wing (venational aberration not present on opposite wing).

Cephalelus leptocarpi n. sp. (Figs. 13, 15.)
♂♀ Same colour and markings as C. hudsoni n. sp., from which it differs in its smaller size, genitalia of male (fig. 13), and, above all, in the very distinctly spatulate head (fig. 15). Examination of a large series of both species has shown these differences to be constant.
Length: Male, total length, 8·5–9 mm.; head, 2–2·5 mm.: female, 8·5–10 mm.; head, 3 mm.
Described from twenty males and twenty-two females.
Holotype (male) and allotype in Myers collection, Biology Laboratory.
There are two specimens in the Hutton collection at Canterbury Museum from Whangarei (no date), where I myself took the material on which I have founded the species. The locality was the edge of a mangrove-swamp, and the food-plant Leptocarpus simplex, as in the previous species. It is interesting that although there were acres of almost pure Leptocarpus association, yet this insect was found, after a large area had been fruitlessly swept with the net, rather plentifully on a patch not a yard square, and apparently nowhere else in the vicinity.
The nymphs, which are dark brown in colour, are found on the food-plant with the adults. The distinctly spatulate head is noticeable at a very early stage.
This species is parasitized by apparently the same red long-legged mite as C. hudsoni n. sp.
The sexes were present in approximately equal numbers. December-January.
Subfamily 3. Typhlocybinae.
Syn. Eupteryginae auctt.
Probably every one is vaguely familiar with the minute whitish active insects which leap vigorously in all directions whenever the grass of lawn or meadow or alpine slope is disturbed by one's passage. Their precipitate flight may even carry them into the picnicker's tea, whence I have occasionally rescued useful specimens. These extremely minute leaf-hoppers, the smallest and most delicate of the Cicadellidae, are members of the subfamily Typhlocybinae. Grass-land is by no means their only habitat; in fact, their distribution wherever vegetation occurs is so wide, and their numbers so great, that there can be few commoner insects in New Zealand. In spite of this fact, the Typhlocybinae have been so neglected that until 1906, when Kirkaldy described fourteen species from Australia, not a single species had been recorded from Australasia; and the same reproach holds in New Zealand to the present date.
As indicated above, the neglect the Typhlocybinae have suffered at the hands of collectors has been such that I have not seen a single specimen in any New Zealand collection other than my own. This paper consequently deals only with material of my own collecting, and its distributional records are, therefore, as inadequate as the area in which I have collected has been circumscribed. For the guidance of future collectors who may wish to gather in a portion of the harvest of new species which assuredly await description, a few notes on collecting and preserving these leaf-hoppers may be useful. Practically the only method of taking those species which occur on bushes and long herbage is that of beating and sweeping. A deep net should be used and the catch examined frequently, otherwise it is impossible to prevent the escape of some while others are

being taken from the net. The species frequenting ground-herbage and short grass are with difficulty obtained thus, and then only by very close sweeping. Such species apparently live very near the ground, well below the free tops of the plants which the sweeping-net reaches. The only certain way of securing these is by capturing them individually with a small net about 3 in. in diameter. This method, necessitating kneeling, as it does, has its disadvantages, but, on the other hand, much more can be observed of the habits of the insects than in the random processes of beating and sweeping. In swampy country, with pools of water, many leaf-hoppers are to be found hopping on the surface of the water, on to which they have come from their adjacent host-plants, and many more can be sent to join them by shaking and kicking the herbage surrounding the water, from which the insects are collected with comparative ease. It is of the utmost importance that records of the food-plants be kept wherever possible.
For killing Typhlocybines and all other leaf-hoppers a cyanide-bottle or small tube, with a smaller tube or quill through the cork, in coleopterist-fashion, is most convenient. It is highly essential that the inside of the bottle be kept dry, otherwise the insects will stick to the sides and to one another and suffer irreparable damage. If the cyanide-bottle has been prepared by the plaster-of-paris method, practically no care will maintain a perfectly dry interior. Pieces of dry potassium cyanide should be placed in the bottom of a tube, covered by several layers of blotting-paper, and then by a tightly-fitting cork, slightly perforated to allow the fumes to pass. Above this cork, which should be forced tightly down on the cyanide, place a piece of blotting-paper and a little crumpled tissue-paper. Several such tubes should be at hand, and if they are kept only for leaf-hoppers no moisture will collect in them.
For permanent preservation Typhlocybines should be gummed across the apex of card triangles, so that the underside of at least the head and genitalia may be examined, or they may be pinned through the scutellum with the finest pins used by micro-lepidopterists. For correct determination a large series is convenient, some of which should be mounted to display the tegmina and wings. On collecting-excursions of long duration they may be kept indefinitely in small screws of tissue-paper; but specimens to be mounted with extended wings must be set as soon as possible after death.
Except in the case of such economically important species as the apple leaf-hopper (Empoasca mali Le Baron), the rose leaf-hopper (Typhlocyba rosae Linn.) and the grape leaf-hopper (Erythroneura comes Say) the biology of the Typhlocybinae is but little known. The first of the above-mentioned species, the American apple leaf-hopper, not only inflicts serious damage on the foliage of apple, potato, bean, and lucerne, but also actually transmits two diseases, fireblight (Bacillus amylovorus) of pomaceous fruit-trees and hopper-burn of potatoes. Even in the cold winters of North America, specimens of the apple leaf-hopper frequently winter in the adult state. In New Zealand, specimens of Erythroneura zealandica n. sp. have been taken after midwinter, and, considering our milder climate, it is quite possible that many of our species are carried over the winter as imagines. The evergreen nature of our bush would help such species as find their host-plant there.
In New Zealand we have one species ranking as a serious apple pest, and probably as a carrier of fireblight; another of very wide occurrence

on a variety of field-crops and pasture-plants, but not yet convicted of serious damage; and a third species, an endemic one, inflicting the same damage on Coprosma grandifolia as the introduced Typhlocyba australis (Frogg.) on apple.
The Typhlocybinae are among those leaf-hoppers parasitized by minute wasps of the family Dryinidae. None, however, have yet been successfully reared from New Zealand species, nor have I found affected leaf-hoppers very numerous. Bird enemies probably account for very few, although in America these small leaf-hoppers are said to form a certain proportion of the food of humming-birds. Finally, Mr. Muir informs me that egg parasites are generally common.
The structural characters of the Typhlocybinae render them probably the most homogeneous and easily recognized subfamily of the Cicadellidae. In the tegmen the three longitudinal veins of the corium—the radius, media, and cubitus—run without branching to the apical cells, so that no ante-apical cells are formed. The 1st A corresponds almost exactly with the claval suture (Cu2 not present) and the third with the claval margin, leaving only a single vein on the disc of the clavus—a character shared (according to McAtee) by no other Auchenorrhyncha. The ocelli are almost always difficult to distinguish, and are often lacking. Long elliptical areas about the middle of each costa bear a thick circumscribed coating of a wax-like substance. The detachable pruinose flakes are known as the “costal plaques.” Some species—e.g., Erythroneura cyathea n. sp.— possess them well developed; but they are by no means a specific character, being subject to individual variation.
Important generic distinctions are found in the venation of tegmina and wings, the main characters being very constant; but the details vary with the individual. Good specific characters are found in the genitalia.
The Typhlocybinae are to be considered among the most highly evolved of the Cicadellidae, their specialization consisting in a simplified venation probably derived from a much more complex type.
The New Zealand species at present collected include six species, in three genera. Of these six species, five are new, and one apparently introduced from Australia, although Mr. Muir informs me that it is by no means certainly endemic there. I am indebted to Mr. Muir for kindly reading the manuscript of this subfamily and for valuable advice and help.
I have followed McAtee in the generic nomenclature. Typhlocyba is often used by other authors where McAtee uses Eupteryx, of which Typhlocyba is in part a synonym. Under such a system my Typhlocyba would be Empoa.
| 1. | Wing with submarginal vein | Dikraneura. |
| Wing without submarginal vein | 2 | |
| 2. | Second apical cell of tegmen triangular and stalked; fourth apical vein of tegmen curving, to end in radial margin | Typhlocyba. |
| Second apical cell of tegmen nearly oblong, based on cross-vein; fourth apical vein of tegmen running parallel to radial margin and ending in apical margin | Erythroneura. |
In connection with this key it should be noticed that, according to Osborn, Gillette, and Woodworth, all those species in which the wing has no submarginal vein and the first two veins unite before reaching the margin, so that only three veins attain the margin, fall into the genus Typhlocyba Germar. Only the African genus Molopopterus Jacobi, which is easily distinguished by other characters, is excepted.

The synonymy of Typhlocyba Germar would, according to these authors, stand as follows: Erythroneura Fitch, Idia Fieber, Zygina Fieber, Anomia Fieber, Aidola Melichar, Empoa Fitch (part), Zyginella Low. The splitting of this unweildy genus becomes, therefore, an obvious desideratum, and Kirkaldy and McAtee proposed to effect this on the old distinction, denounced by Woodworth as untrustworthy and inconstant, that in Typhlocyba (sens. strict. and as used by me) the fourth apical vein of tegmen curves to the radial margin, and the second apical cell is triangular and usually stalked, while in Erythroneura (sens. strict.) the fourth apical vein of tegmen runs parallel to the radial margin and ends in the apical margin, the second apical cell being more or less oblong and based directly on the cross-vein. Examination of very large series of New Zealand specimens has shown this to be a perfectly constant character so far as our species are concerned, and it has consequently been adopted in this paper. The figures of Typhlocyba and Erythroneura venation will show this difference more clearly than a printed description. Should further study demonstrate instability in this venational character, all the species I have described under Erythroneura must be placed in Typhlocyba, which will then have the complete synonymy given above. In all the species of both genera which I have examined the male genitalia, and especially the aedeagus, differ fundamentally, and thus afford a sounder distinction between the two genera than the venation alone could supply. The figures will show this clearly.
Genus 1. Dikraneura Hardy.
An almost cosmopolitan genus.
Dikraneura maorica n. sp. (Figs. 16–18.)
♂. Vertex, pronotum, and scutellum sulphur-yellow. Tegmina paler. Eyes black. Wings hyaline, somewhat opalescent. Underside and legs uniform yellow. Claws black. Tarsi often green. Rostrum tipped with crimson. Vertex rather more than twice as wide as medianly long. Fore-border rounded. Pronotum rounded anteriorly; posterior angles somewhat produced but truncate; posterior border moderately excavated. Genitalia concolorous with rest of body, except ovipositor, which is shining brown.
Length (including, in this and subsequent species, the folded tegmina): Female, 3 mm.
Male unknown.
This beautiful little species is near to D. flavipennis (Zett.), which it resembles also in habitat (among sedges in damp places). It is distinguished by the fore-border of vertex, which in our species is rounded instead of angular, and by the absence of black points on the hind tibiae.
Among carex and grass in bush-clearing (swamp-forest), Long Acre, Wanganui, December.
Described from nine females.
Holotype in Myers collection, Biology Laboratory.
Genus 2. Typhlocyba Germar. (Figs. 19–23, 35, 36.)
Typhlocyba australis (Froggatt). Australian Apple Leaf-hopper.
Empoasca australis Frogg., Agric. Gaz. N.S.W., vol. 29, p. 568, 1918.
♂♀. Vertex and pronotum bright sulphur - yellow; eyes usually blackish. Scutellum bright yellow, often with deeper-yellow patch on each side of base. Tegmina of the same bright-yellow tint, membrane iridescent

and hyaline. Wings hyaline; underside uniform yellow. Frons yellow. Vertex approximately twice as broad as medianly long. Fore-border rounded. Pronotum trapeziform, rounded anteriorly and slightly excavated posteriorly. Scutellum slightly narrower at base than base of head including eyes. Ovipositor shining mahogany-brown.
Total length, 4 mm. (male and female).
November to March.
Nelson, Auckland, and Hawke's Bay orchard districts, where it inflicts considerable damage on the foliage of apple and hawthorn. This species is probably introduced from Australia, and must not be confused with the well-known Empoasca mali (Le Baron) of America, which has not yet been introduced, and from which it differs not only structurally but also biologically, in that the results of its attack are rusty spots and patches on both sides of young and old leaves, instead of a curling of the younger leaves only, as in the American species. Its method of attack is very similar to that of Typhlocyba (Empoa) rosae (Linn.), which it resembles in some other respects, but from which it is quite distinct.
Genus 3. Erythroneura Fitch.
| 1. | Colour dark, with numerous markings | E. ansonae. |
| Colour pale and practically uniform | 2 | |
| 2. | Colour bright golden-yellow | E. kiekie. |
| Colour pale-yellowish, greenish, or whitish | 3 | |
| 3. | Under 3 mm. long. Colour green or greenish-yellow. Dorsal surface of abdomen black | E. zealandica. |
| Over 3 mm. long. Colour whitish. Dorsal surface of abdomen white | E. cyathea. |
Erythroneura zealandica n. sp. (Figs. 24–29, 37, 39, 41, 42.)
Vertex dirty greenish-yellow, with sometimes an indistinct greyish smudge on each side. Eyes greyish-black or grey, edged with paler. Pronotum and scutellum yellowish-green, the latter sometimes with darker-coloured area on each side of base. Scutellum divided by transverse middle line into a plane anterior portion, with indications of three median longitudinal stripes, and an elevated posterior portion including the acute apex. Dorsal surface of abdomen black, showing through tegmina and wings. Tegmina yellow to yellowish-green. Membrane and wings hyaline. Underside uniform pale-yellowish, the tibial spines and tip of rostrum darker. Vertex about twice as wide as medianly long. Pronotum broadly rounded anteriorly and slightly excavated posteriorly, with a very faint and occasional indication of a median longitudinal keel. Base of scutellum about

Fig. 30.—Erythroneura kiekie n. sp.: male genitalia.
Fig. 31.—E. cyathea n. sp.: male genitalia.
Fig. 32.—E. cyathea n. sp.: tegmen. a, costal plaque.
Fig. 33.—E. cyathea n. sp.: wing.
Fig. 34.—E. ansonae n. sp.: female subgenital plate.
Fig. 35.—Typhlocyba australis (Frogg.): aedeagus, lateral view (after Muir).
Fig. 36—T. australis (Frogg.): apex of aedeagus (after Muir).
Fig. 37.—Erythroneura zealandica n. sp.: aedeagus, ventral view.
Fig. 38.—E. cyathea n. sp.: aedeagus, ventral view.
Fig. 39.—E. zealandica n. sp.: caudal view of pygophor (eighth sternite removed).
Fig. 40.—E. kiekie n. sp.: caudal view of pygophor; anal segment not shown.
Fig. 41.—E. zealandica n. sp.: aedeagus, lateral view.
Fig. 42.—E. zealandica n. sp.: male genital style.
Fig. 43.—E. kiekie n. sp.: male genital style.
Fig. 44.—E. cyathea n. sp.: male genital style.
Fig. 45.—Thanatodictya tillyardi n. sp.: male.
Fig. 46.—T. tillyardi n. sp.: lateral view of male genitalia.
Fig. 47.—T. tillyardi n. sp.: male pygophor.

as wide as base of vertex between eyes. Apex of scutellum acute. Male genitalia short and abrupt; claspers bent abruptly, tips almost reaching the dorsal line of abdomen in lateral view. Female genitalia—ovipositor short, tipped with fuscous, and furnished with seven or eight very stout short bristles. Ninth or last unmodified abdominal tergite abruptly truncate.
Total length, 2·7 mm. (male and female).
This is the common grass-frequenting species, found in large numbers in any grassy places, whether the grass be introduced pasturage or native tussock. It is also common in all stages on tauhinu, or cottonwood (Cassinia leptophylla). There is considerable colour-variation, those from Wanganui being yellowish, the common Wellington specimens greenish, and those taken on tauhinu in various localities of the Wellington district a peculiar greyish-green resembling that of the food-plant. I can find no accompanying structural differences. The various forms seem to grade into one another. A somewhat dilapidated female taken on mangrove at Whangarei appears to belong to this species. Typical specimens have been collected from manuka (Leptospermum scoparium). While making the above description wide enough to include all the forms here enumerated, I have retained the Cassinia forms as types.
Tararua Range, Wellington, 4,000–5,000 ft.; Wanganui; October to June.
Described from eighty-eight males and eighty-four females.
Holotype, allotype, and paratypes in Myers collection, Biology Laboratory.
Erythroneura kiekie n. sp. (Figs. 30, 40, 43.)
♂♀. Whole upper surface bright yellow, thoracic nota often orange. Eyes blackish (eye-colour is of little taxonomic value, as it may vary from whitish to black in the two eyes of the same individual). Tegmina yellow, the membrane a little paler, but not hyaline. Wings hyaline. Underside uniform pale yellow. Claws, except posterior ones, black. Ovipositor pale-brownish. Rostrum tipped with brownish. Upper surface of abdomen blackish. Vertex very broadly rounded anteriorly; not twice as wide as medianly long. Median longitudinal suture distinct. Pronotum rounded anteriorly, and considerably excavated posteriorly—the anterior portion paler and somewhat depressed, posterior half shining and darker yellow. Scutellum a little wider at base than base of vertex between eyes; transverse line about half-way, to which the sides gradually slope from base, beyond which transverse line (caudad) they rapidly narrow to the very acute and somewhat elevated apex.
Total length, 3·7 mm. (male and female).
This handsome species has been taken only on kiekie (Freycinetia Banksii), to which it may possibly be confined, and on which it occurs in considerable numbers.
York Bay, Day's Bay, Wellington; Whangarei; November to March.
Described from six males and six females.
Holotype, allotype, and paratypes in Myers collection, Biology Laboratory.
Erythroneura cyathea n. sp. (Figs. 31–33, 38, 44.)
♂♀. General colour pale yellowish- or greenish-white or practically colourless. Vertex pallid, tumid, produced, punctate, with occasionally two brownish patches varying from faint smudges to distinct spots. Eyes variegate. Pronotum with an anterior smooth, collar-like portion;

remainder finely transversely striate. Humeral angles produced, truncate, covering base of tegmina. Pronotum and tegmina very pale yellowish-or greenish-white or practically colourless. Pronotum narrowly rounded anteriorly, and very slightly excavated posteriorly. Wings milky. Base of scutellum considerably wider than posterior border of vertex. Scutellum with depressed anterior portion and slightly elevated acute apex. Triangular area on each side of base of scutellum often brownish. Under - surface pale creamy-white. Claws fuscous. Frons very tumid, with dull frosted surface. Genitalia of both sexes pale. Upper surface of abdomen white.
Total length, 3·4–4 mm. (male and female).
In one well-marked variety (No. 175) the colour is a somewhat brighter yellowish, and the hind border of pronotum, disc of scutellum, extreme edge of radial margin of tegmina, and to a less extent the membrane of tegmina are suffused with fuscous.
In a second variety (No. 185, one specimen) the brown colour is confined to the two basal triangular spots of the scutellum and to its sides, forming a very distinct dorsal V. As I can find no accompanying structural peculiarities, and as the faint brown markings sometimes present in the more typical specimens vary greatly in intensity, I am constrained to regard this as nothing more than a colour variety comparable to those of the common Cercopid Philaenus trimaculatus (Walker).
This variable and common species occurs among mixed undergrowth in ordinary rain-forest. With specimens collected by beating and sweeping it is difficult to be sure of the food-plants, but one is the tree-fern Cyathea dealbata, and another, on which all stages can be collected, is Coprosma grandifolia. Specimens from the tree-ferns are uniformly and constantly more whitish, and the types have been selected from these.
Tararua Range, Wellington district, 2,000–3,000 ft., November to June.
Described from twenty-six males and forty females.
Holotype, allotype, and paratypes in Myers collection, Biology Laboratory.
Erythroneura ansonae n. sp. (Fig. 34.)
♀. General colour yellowish marked and suffused with greenish and olivaceous. Vertex fairly considerably produced, about twice as wide as medianly long. Eyes black, edged with yellow. Median suture of vertex distinct, with or without obscure dusky blotch on each side. Pronotum— disc dark olive, anterior and lateral borders pale yellow. Just caudad of anterior yellow edge the olive area is edged with blackish; a few other small irregular black marks. Pronotum sloping gradually from rounded anterior point to posterior border, which is considerably but gradually excavated. Scutellum at base much wider than base of vertex, but not quite so wide as eyes. Transverse line at ½ divides the olivaceous anterior portion, with its two shining brown basal triangles, from the elevated bright-yellow apex. Lateral angles of scutellum paler. Dorsal surface of abdomen dark, showing through closed tegmina and wings. Tegmina semitransparent, olivaceous; membrane infuscated. Ventral surface olivaceous or stone-coloured. Rostrum tipped with reddish-brown. Leg-spines darker. Ovipositor brownish, with a row of short stout bristles along each side. Subgenital plate notched at tip (fig. 34), a very distinct character. Frons and clypeus deep dirty-yellow. Clypeus with some indications of transverse rugosities. Genae darker.
Total length (female), 3·5-3.7 mm. Male unknown.
The markings superficially resemble those described for E. (Zygina) parvula (Boh.), but are much less clearly defined. A comparison of the

two insects shows, moreover, that the whole habitus of the present species is different, the tegmina particularly being one and a half times as long as those of E. parvula.
Damp grassy and sedgy clearings in rain-forest, Wellington, Wanganui December-April.
I dedicate this distinct and handsome species to Mrs. H. J. D. Stowe (née Anson), whose co-operation in collecting has been invaluable.
Described from two females.
Holotype in Myers collection, Biology Laboratory.
II. Fulgoroidea (Plant-hoppers).
Some sixteen species of this interesting superfamily have now been described from New Zealand. This number probably represents a mere fraction of the total. It includes several introduced forms from Australia, notably Siphanta acuta Walker, Scolypopa australis Walker (common in tremendous numbers on a great variety of plants in the Waikato and Auckland districts), and Sephena cinerea Kirkaldy (also common in the Auckland Province). The latter has been proved to carry fireblight (Bacillus amylovorus) of apple-trees, Mr. R. Waters, Bacteriologist of the Biology Division of the Department of Agriculture, having artificially induced the disease by means of cultures from infected leaf-hoppers. An effort was made to ascertain whether the bacilli were transferred by the sucking of the plant-hopper, and with this end in view, at Dr. Tillyard's suggestion, cultures were prepared from the salivary glands of this insect, but were unsuccessful owing to the presence in overwhelming numbers of other bacteria.
A considerable number of endemic Fulgoroids now await description They will not be published until a thorough study of the genitalia has been made. Mr. F. Muir, whose assistance in this superfamily has been especially valuable, is examining next year some of Walker's Cixiid types in the British Museum, so that a complete revision must await the results of this examination. In the meantime I am describing here a new and striking species which is the unique representative of its family in New Zealand, and is not therefore likely to be confused with any other species.
Family Dictyophoridae.
Well represented in the Australian and Oriental regions.
Genus Thanatodictya Kirkaldy.
Bull. 1, Div. Ent., H.S.P.A. Expt. Sta., p. 392, 1906.
A typically Australian genus.
Head very elongate, narrow, porrect. Vertex bordered on each side by a keel. Tegmina hyaline: apical third subreticulate. Legs simple; posterior tibiae four-spined. Vertex nodulose at base, and stigma dark (this last sentence characterizes the subgenus Niculda Kirk.).
Thanatodictya tillyardi n. sp. (Figs. 45-47.)
♂. General colour fulvous clouded with fuscous. Vertex bright yellowish-brown, except nodulosities at base, which are darker. Immediately anterior to these, vertex is finely transversely striate; a dark-brown stripe between longitudinal ridges of vertex. Pronotum concolorous with

base of vertex;- median pronotal keel ivory-white. A median ivory-white stripe on mesonotum expanding caudad, and ending in bluish-white posterior angle of scutellum. Mesonotum darker brown, with longitudinal keel on each side. Basal abdominal terga marked heavily with fuscous; remainder of dorsal surface of abdomen rich brown clouded with fuscous, segmental margins relieved sparingly and medianly with pallid. Tegmina and wings hyaline, colourless. Veins brown, darker distally. Stigma blackish, with normally two internal transverse veins; but the right tegmen of one specimen has only one transverse vein in stigma. A blackish smudge extends from apex of tegmen to apex of suture, and farther as an ill-defined line along dorsum of tegmen. Head slightly ascending; about as long as remainder of body. Legs longitudinally angular; stramineous striped with piceous; sparingly setose. Ocelli clear yellow ringed with scarlet. One specimen is much paler than the other.
Total length (male), 7–8 mm.; tegmen, 6–7 mm.
By its nodulose base to vertex it belongs to subgenus Niculda Kirkaldy. In its somewhat ascending head it resembles T. (Niculda) anadyomene Kirkaldy; but differs in its larger size and in the presence of only two transverse veins in the stigma of tegmen, both well marked.
Holotype (male), Myers collection, Biology Laboratory; paratype (male), Tillyard collection, Cawthron Institute, Nelson.
Described from two males taken by Dr. R. J. Tillyard at a height of 3,000 ft on the Dun Mountain, Nelson, in February. I have much pleasure in naming it after the discoverer.
Since the above was written Mr. A. Philpott and M. André Tounoit have taken the species in considerable numbers by sweeping bracken-fern (Pteridium esculentum) near the same locality. Some of the specimens show a strong and beautiful green suffusion.
List of References.
Cogan, E. S., Contribution towards our Knowledge of the Homoptera of South Africa, Ohio Journ. Sci., vol. 16, p. 161, 1916.
Edwards, J., The Hemiptera-Homoptera of the British Islands, 1896.
Froggatt, W. W., The Apple-leaf Jassid (Empoasca australis), Agric. Gaz. N.S.W., vol. 29, p. 569, 1918.
Gillette, C. P., American Leaf-hoppers of the Subfamily Typhlocybinae, Proc. U.S. Nat. Mus., vol. 20, p. 709, 1898.
Hartzell, F. Z., The Grape Leaf-hopper, New York Agric. Expt. Sta. Bull. 359 et al., 1913.
Hutton, F. W., Synopsis of Hemiptera of New Zealand, Trans. N.Z. Inst., vol. 30, p. 167, 1898.
Kirby, W. F., Notes on Dorydium (?) westwoodi Buchanan White, Trans. Ent. Soc. Lond., 1894, p. 411.
Kirkaldy, G. W., Leaf-hoppers and their Natural Enemies, Ent. Bull. 1, H.S.P.A., 1906.
—— Leaf-hoppers: Supplement, Ent. Bull. 3, H.S.P.A., 1907.
—— Entomologist, vol. 34, p. 339, 1901.
—— List of Hemiptera of Maorian Subregion, Trans, N.Z. Inst., vol. 41, p. 22, 1909.
Lathrop, F. H., Leaf-hoppers injurious to Apple-trees, New York Agric. Expt. Sta. Bull. 451, 1918.
McAtee, W. L., Genera of the Eupterygidae (Homoptera-Jassoidea), Proc. Biol. Soc. Washington, vol. 31, p. 109, 1918.
Metcaf, Z. P., The Wing-venation of the Jassidae, Ann. Ent. Soc. America, vol. 6, p. 103, 1913.
Osborn, H., Leaf-hoppers of Maine, Maine Agric. Expt. Sta. Bull. 238, 1915.
—— Life-histories of Leaf-hoppers of Maine, Maine Agric. Expt. Sta. Bull. 248, 1916.
—— Leaf-hoppers affecting Cereals, Grasses, and Forage Crops, U.S. Dept. Agric. Bur. Ent. Bull. 108, 1912.
Rand, F. V., and Pierce, W. D., A Co-ordination of our Knowledge of Insect Transmission in Plant and Animal Diseases, Phytopathology, vol. 10, p. 189, 1920.
Signoret, V., Ann. Soc. Ent. France (5), vol. 10, p. 43, 1880.
White, F. B., List of Hemiptera of New Zealand, Ent. Mo. Mag., vol. 15, 1879.
