
Art. 32.—A New Tachinid Genus and Two New Species.
[Read before the Wellington Philosophical Society, 25th October, 1921; received, by Editor, 31st December, 1921; issued separately, 12th May, 1923.]
Plate 37.
Amongst a large collection of Diptera recently presented to me by Mr. J. W. Campbell, of Christchurch, is a most interesting new species of Tachinid, which I describe below in a new genus under the name of Campbellia campbelli. Not only has Mr. Campbell secured large numbers of valuable Diptera, but he is also an accomplished observer of the structure and habits of these insects, as is amply shown in his recent work on the adult and pre-adult stages of the Blepharoceridae.*
In 1911 Mr. E. H. Atkinson captured a second species, which was sent to the British Museum, and through the courtesy of Mr. F. W. Edwards the specimen has been submitted to me, and is herewith recorded under the specific name of cockaynei n. sp., named after Mr. A. H. Cockayne, Government Biologist.
The extraordinary wing-development in these insects also occurs, only to a much greater extent, in the New Zealand Anthomyid Exsul singularis Hutton. A tendency to this development to a less extent occurs in several other species of mountain Diptera.
Subfamily Gymnosominae.
Genus Campbellia n. g.
Wing very broad, more than half as long as broad, the proximal half of posterior margin parallel to costa, distal part abruptly ascending to apex; apical cell closed and petiolate; third vein meeting costa very slightly below apex of wing; last section of fourth vein abruptly but evenly rounded upward to meet the third at right angle; posterior crossvein about one and a third times its own length from the small cross-vein which is a little beyond the middle of the discal cell; first vein meeting costa almost immediately above small cross-vein; veins bare, stoutly developed; no sign of a costal spine.
Head much broader than high; eyes bare, dichoptic in male but convergent at ocellar triangle, extending below vibrissal angles, which are well above oral margin; the latter produced forward snout-like. Antennae well separated basally, extending close to forward curvature of face; first joint very short, third short and rather oval, a little more than one and a half the length of the second; arista pubescent, stout, three-jointed, first two joints short. Facialia bare, a few short weak bristles above vibrissae, frontal bristles weak and short, extending to base of second antennal joint; otherwise parafrontals bare; post-orbital bristles weak, not extending below middle of posterior orbits. Proboscis rather slender, elongate; palpi well developed, slender but swollen apically.
[Footnote] *Trans. N.Z. Inst., vol. 53, pp. 258–88, 1921.

Thorax quadrate, scutellum narrow and broad; metanotum conspicuous behind scutellum; few outstanding chaetae, spatulate spines on wing costa at articulation and on meso- and noto-pleurae being unique (at least in male).
Legs stout and bristly; posterior tibiae with row of strong spines, becoming shorter basally and not reaching beyond middle, on outside of joint; protarsus not quite half length of whole tarsus; claws large; pulvilli large and narrow.
Abdomen (in male) transversely rugose, broader than thorax, broadly ovate, flattened; four visible segments, the first, second, and third of equal length, the fourth one and a half as long as preceding. Genitalia more or less prominent, curved beneath fourth segment. No abdominal chaetae, but among vestiture are distinct marginal bristles on third segment and lateral ones on third and fourth segments.
| Abdomen tawny, but fuscous apically; scutellum black, with a golden apex | campbelli n. sp. |
| Abdomen deep purplish-blue on first two segments, the remainder rather cupreous and dusted with brownish pollen; scutellum unicolorous, bronzy-black | cockaynei n. sp. |
C. campbelli n. sp.
♂. Ptilinal suture (fig. 1) rather narrower above but evenly rounded, its termini situated in line with vibrissal angles well above line of lower eye-margins. Facial plate without carina, broad, concave, black, with cinereous tomentum, each side abruptly produced on to facialia and forward below to broad and prominent tawny epistome, which is minutely tomentose. Facialia prominent, black, bare except for a few short bristles just before vibrissae, which are large and widely separated, and situated well above oral margin. Peristomalia ascending, black, but with golden or whitish down and strong marginal bristles, behind which are a few short weaker ones. Oral cavity very large, elongate and rectangular; proboscis long and slender, rigid, geniculated, brownish in colour, the labella small; palpi slender, elongate, clavate, orange-red in colour. Lower margin of head longer than longitudinal axis at antennae; head, in profile, produced abruptly forward at oral margin, which is ascending (fig. 2).
Antennae (fig. 3) black, with a greyish reflection, comparatively short, extending to produced epistome; inserted above middle line of head and eye; broadly separated at insertion by globular projection of the shiny black lunula. Second joint normal, clothed with short greyish pubescence and short black bristles, a large conspicuous one above; third joint orbicular and clothed with minute greyish pubescence; arista stout, tapering to a point, longer than third joint, black, with stiff pubescence, three-jointed, the first very short, second a little longer.
Eyes bare, convergent in front of ocellar triangle and reaching well toward oral margin below line of vibrissal angles (fig. 1). Vertex narrow, space between eyes being occupied mostly by vertical triangle. Chaetotaxy of head consisting of delicate bristles; inner vertical bristles longer than others, reclinate and convergent; outer verticals absent; post-orbitals extending almost to inner verticles and not reaching to halfway along orbits, but merging into and ending in very short bristles; a pair of well-separated post-verticals.

Front dull black in colour, but brownish in some lights; depressed throughout its length in the form of a shallow V-shaped depression; broad at antennae but narrowing abruptly to vertical triangle; deeply notched behind antennae, and between this and lunula is a deep, crescentic, black depression. Vertical triangle large and broadly triangular, black in colour, black ocelli prominent and well separated. Ocellar bristles delicate, convergent, proclinate; post-ocellars delicate; to each
Campbellia campbelli n. sp.
Fig. 1.—Ptilinal suture, &c.
Fig. 2.—Profile of head.
Fig. 3.—Antenna.
Fig. 4.—Bristle of mesopleurae.
side of ocellars is a short delicate bristle, while between ocellars post-ocellars are brownish hair-like bristles; pre-ocellars absent. Parafrontal areas black in ground-colour, but altogether silvery in some lights; projecting above plane of eye and front, demarcation being abrupt; transversely rugose between front and eye, much narrowed at vertex, broad at antennae, below which they descend along ptilinal suture to run abruptly to a point on orbits in line with vibrissal angles; depressed between ptilinal suture and orbits Frontal bristles in two rows descending

a little below antennae, otherwise parafrontals are bare. Parafacials without bristles; restricted to narrow area extending from terminii of ptilinal suture to a point on orbits; black but with golden to greyish pubescence around ptilinal terminii. Orbits sinuated to point on parafrontals below antennae. Cheeks silvery-white and clothed with delicate white hairs; narrow below parafacials but broadening below eyes and extending upwards on to occiput. Occiput black in ground-colour but with silvery grey tomentum; concave; bare but for a few delicate hairs and some short delicate bristles, into which merge orbital bristles from above; orbital area broad.
Thorax quadrate, black in ground-colour, dorsum dusted with silvery and golden pollen; a shiny black, broad area on each side extending from angles of transverse suture to alar calli. Scutellum narrow but broad; black but golden-yellow across apex, where are three deeper yellow spots, a narrow longitudinal median one and an oval one on each side. Metanotum conspicuous, developed as in nemocera; black, cinereous in some lights, bare. Sternopleurae clothed with delicate black bristles, among which are stronger ones above, there being in particular a large very strong spine near upper posterior angle. Hypopleural bristles consisting of a vertical row of strong bristles and a pair of smaller ones at upper posterior angle. Mesopleurae clothed with numerous large black bristles, many of which are spatulate in shape (fig. 4); they are arranged over upper part of mesopleurae, but extend below along posterior margin. Pteropleurae with a clump of large bristles above and a single spine below. Notopleural bristles consisting of a dense clump of black spatulate spines, there being also a large pointed spine among a few delicate bristles in front. Humeral bristles consisting above of a large stout spine, a long delicate bristle strongly recurved distally, some short delicate black bristles and a few golden ones; below is a clump of golden bristles and spatulate spines. Post-humerals consisting of some small black bristles, from among which projects a larger distinct one. The dorsum is uniformly clothed with short delicate bristles, among which the following may be seen with difficulty: a pair of presuturals; two pairs of dorsocentrals, near scutellar suture; one pair of acrostichals, near scutellar suture. A conspicuous pair of post-alars, large spine on alar angle outstanding. Scutellum clothed with delicate bristles, apical margin bare.
Wings (Plate 37, fig. 1) tawny at base, anterior portion clouded with brown, apex and remainder clear; veins tawny at base but otherwise blackish-brown; lower squamae large, rather hood-like, brownish, translucent, surface punctured; upper squamae brownish on one half, whitish on other, much smaller than lower squamae and fringed with long golden hairs. Petiole of apical cell about one and a half length of small cross-vein; a short blunt stump from fourth vein projecting into discal cell on distal side of small cross-vein; posterior cross-vein slightly sinuate; first vein very broad distally; halteres tawny and mushroom-shaped.
Legs stout and bristly; posterior femora laterally flattened, golden-yellow except for the blackish-brown apical third; clothed with numerous golden hair-like bristles basally and strong black bristles above, fewer below; three pre-apical bristles below to the outside. Middle femora blackish-brown but golden-yellow on basal third, anterior femora blackish-brown dusted with greyish pollen above and basally. Tibiae blackish-brown, each with a ridge on which is a row of short spines down outside, more distinct on posterior tibiae, which have a well-marked ridge running

down outer surface but ending before apex; on inside of this ridge proximally are three delicate bristles, and opposite these, on outside, is a row of five strong spines, being smaller to the base but not extending halfway down the joint. On side of tibia, almost in line with third most distal spine, is a distinct spatulate bristle. On upper side is a delicate pre-apical bristle, and below a stout apical one. Middle tibiae with stout apical spine below and delicate pre-apical one above; towards centre, but more distally, is a stout bristle toward lower side. Anterior tibiae with delicate apical bristles below and a pre-apical one above.
Tarsi normal in shape, bristly, blackish-brown; the protarsi pale brown, not quite half length of whole tarsi. Onychotarsi longer than preceding joint and with pair of delicate elongate bristles above; claws large, black; pulvilli large, narrow, pale brown.
Abdomen broadly ovate, flattened, four visible segments transversely rugose; tawny, apical segment fuscous in middle and apex, with greyish pollen; posterior margin of each segment darker. Dorsal plates curved on to ventral surface; ventral plate of first segment large, extending to margin of dorsal plate; ventral plates of other segments small, that of the fourth deeply notched on posterior margin to receive globular end of the black genitalia which are curved beneath apex of abdomen; ventral membrane conspicuous.
Abdomen diaphanous, dusted with yellowish-white pollen on each side of third and fourth segments; a blackish elongate spot on each side of centre margin of first segment. Surface of each segment sparsely clothed with short delicate bristles, being denser along sides. Distinct marginal bristles on third segment, and distinct bristles on each side of third and fourth segment.
♂. Length, 7·5 mm.
Genotype: No. 1259, D. M.
Habitat.—Blackball. Captured by Mr. J. W. Campbell; a single specimen.
C. cockaynei n. sp. (Plate 37, fig. 2.)
♂. Palpi tawny; front velvet - black and longitudinally striated. Dorsum of thorax shiny, rather blue-black; a golden tomentose area between scutellar and transverse sutures; dorsum margined with silvery tomentum, giving a striped appearance to thorax; three areas of silvery tomentum in front of transverse suture, arranged indistinctly as three short stripes, central one broadest; in side lights these merge to form a broad area continuous with posterior golden area. Pleurae black but with silvery to brownish tomentum in continuation with that margining dorsum; metanotum clothed with brownish tomentum. Petiole of apical cell about twice as long as small cross-vein. Legs blackish, anterior femora silvery and posterior somewhat tawny at base; posterior tibiae, protarsi, and epitarsi with golden hair along inner side. Abdomen deep purplish-blue, but the two apical segments rather cupreous in most lights; brownish pollen at posterior angles of second segment, along sides of third segment, and over most of apical. Genitalia prominent, projecting knoblike beneath apex of abdomen.
♂. Length, 7 mm.
Holotype in British Museum.
Habitat.—Day's Bay, Wellington Only one specimen, captured by Mr. E. H. Atkinson on flowering veronica.
