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Volume 54, 1923
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– 452 –

Genus Hilara Meigen.

Anterior protarsi of male usually enlarged; proboscis not longer than but about as long as head-height; vein Sc meeting costa; R4 and R5 branching; metapleurae bare.

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Table of Species.
1 Thorax tawny, with indistinct darker stripes; abdomen brownish-yellow, the posterior margin of each segment tawny; genital appendage of ♂ nearly as long as body; length of body, 6 mm. flavinceris n. sp.
Thorax not tawny; greyish or blackish species 2
2 Altogether cinereous or greyish species with yellow legs 3
Blackish species 5
3 Altogether a cinereous littoral species; length, 2·25 mm. philpotti.
Body greyish, legs yellow 4
4 Wings tinged with yellow smithii.
Wings clear; a long genital appendage vector n. sp.
5 Wings tinged with brown 6
Wings hyaline 7
6 A narrow-bodied fly; ♂ without genital appendage; length, 6 mm. fumosa.
A small fly; ♂ with a short genital appendage; length, 4 mm. fossalis n. sp.
7 Genitalia of ♂ distinct, recurved over apex of abdomen, but without genital appendage; length, 3·75 mm. littoralis n. sp.
Genitalia of ♂ with a distinct appendage; length not less than 5 mm. 8
8 Genital appendage short; length of body, 5·5 mm. hudsoni.
Genital appendage nearly as long as body; length of body, 6 mm. dracophylli n. sp.
Provisional Table based on Chaetotaxy.*
1 Two pairs of scutellars 2
At least three pairs of scutellars 3
2 Acrostichals biserial and strong; dorsocentrals uniserial and strong (fig. 52); occipitals in two rows, the two vertical ones of inner row outstanding (fig. 51) fumosa.
Acrostichals absent; dorsocentrals biserial (?), the inner row regular and weak, the outer irregular in size (fig. 55); occipitals weak, in two rows, three on each side of vertex strong (fig. 54) fossalis n. sp.
3 Three pairs of scutellars; acrostichals weak, forming at least four rows; dorsocentrals uniserial, terminal ones outstanding, remainder weak (fig. 57) smithii.
More than three pairs of scutellars 4
4 Five pairs of scutellars; acrostichals weak, in at least four rows; dorsocentrals weak, biserial (fig. 49); occipitals in three rows, the outer triserial, weak but for two strong orbitals (one of which is a vertical) and one strong postvertical on each side (fig. 48) vector n. sp.
Four pairs of scutellars and sometimes a pair of discals 5
5 Acrostichals absent; dorsocentrals uniserial, weak but ending posteriorly in a strong bristle; the outer and pre-apical scutellars outstanding (fig. 46); occipitals weak, in four rows littoralis n. sp.
Acrostichals present 6
6 Apical scutellars weak, sometimes a pair of discals present 7
Scutellars uniform, no discals present (fig. 43) flavinceris n. sp.
7 A pair of scutellar discals sometimes present; marginal scutellars, except weak apical pair, uniform; dorsocentrals ending posteriorly in a pair of large bristles before scutellum (fig. 50) dracophylli n. sp.
Apical scutellars weak, the pre-apicals outstanding; no discals present; dorsocentrals ending in two pairs of outstanding bristles posteriorly hudsoni.

[Footnote] * H. philpotti is not included.

[Footnote] † It is doubtful if H. fumosa belongs to this genus.

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H. philpotti Miller.

H. philpotti Miller, Trans. N.Z. Inst., vol. 45, p. 202, figs. 9 and 10, 1913.

In fig. 10 the vein Sc should not be evanescent, but should meet the costa, though indistinctly. The cone-shaped protuberance referred to in the original description is really the very short genital appendage.

H. flavinceris n. sp.

♂. Eyes emarginate at antennae, narrowly approximated for a distance on front below ocellar triangle, which is prominent and tawny, but greyish-brown pruinose to black in some lights; ocelli orange; a pair of short ocellar bristles and several minute ones behind (fig. 42). Front very narrow at approximation of eyes but broadening below to antennae, gently descending along eye approximation but almost vertically from thence to antennae, thus angulate in profile; tawny and greyish-brown pruinose; a double row of delicate and short hair-like orbital bristles, not extending to line of antennae, which are inserted about the middle line of head; first and second joints tawny and greyish-pruinose, with delicate bristles, first somewhat longer than second; third joint of both antennae lost. Face bare, coloured as front, slightly longer than broad, oral margin not extending very close to antennae. Cheeks very much restricted and inconspicuous, clothed with tawny hairs. Proboscis orange-yellow to brown, equal to or a little less than height of head; palpi tawny, clothed with delicate tawny hairs basally and with a few long bristle-like tawny hairs below apically. Occiput convex, greyish-yellow, with darker reflections due to minute pubescence; a broad area along each orbit clothed with short black bristles merging into long golden stiff hairs below; in bristly area toward vertex is a regular system of chaetae (fig. 42).

Thorax tawny, greyish-yellow pruinose, pleurae black in some lights; dorsum somewhat arched, testaceous in some lights, with four darker longitudinal stripes seen when light falls from in front; chaetotaxy as in fig. 43, there being very few strong bristles; pleurae bare except propleurae, which are clothed with delicate yellow hairs extending on to humeral areas. Scutellum with triangular testaceous area at base, otherwise tawny, with four pairs of erect marginal bristles.

Wings faintly tinged with tawny; veins tawny; vein Sc not evanescent but meeting costa; R4 strongly curved forward after leaving R5 Halteres tawny.

Legs tawny, darker at apex of tarsi, not strongly bristled but bristles more distinct at apices of joints; anterior protarsi somewhat thickened; pulvilli tawny; claws black on distal half but tawny on basal half, owing to tawny pubescence.

Abdomen rich brownish-yellow, but tawny across posterior margin of each segment and at sides; clothed with long delicate tawny hairs, which become bristle-like on apical segment and laterally on posterior margin of each segment. Along sides of abdomen are clear pit-like spots. Genital appendage almost as long as body, tawny in colour but fuscous along lower edge.

♂. Length, 6 mm; + genital appendage, 11 mm.

Holotype: 1208, D. M.

Habitat.—George's Stream, Wainuiomata (G. V. Hudson).

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H. littoralis n. sp.

♂. Eyes dichoptic, emarginate at antennae; front brownish-black, with cinereous reflections and a row of delicate bristles on each side, ocellar triangle brownish-black, with pair of strong ocellar bristles. Face short and cinereous. Antennae (fig. 44) large, black with silvery reflection, third joint clavate with short style. Proboscis a little longer than head-height, directed downwards, blackish-brown; palpi short and black, but brownish basally. Occiput cinereous and bristly (fig. 45).

Thorax blackish-brown in ground-colour, pleurae cinereous to silvery; pteropleurae with transverse row of delicate grey bristles; dorsum (when viewed with light falling from in front) narrowly margined on each side with blackish-brown; adjoining this margin is a silvery area and centrally a large reddish-brown area with a pair of indistinct darker stripes more conspicuous anteriorly behind head, where the reddish-brown merges into a cinereous reflection (fig. 46); the whole surface distinctly bristly, among which is a system of chaetae as in fig. 46. Metanotum cinereous.

Wings not coloured, veins tawny; Sc ending in costa; anal angle prominent. Halteres brown.

Legs brown, tarsi black; bristly; anterior protarsi somewhat enlarged.

Abdomen bristly; black with cinereous reflections. Genitalia more or less almond-shaped when viewed from above; in profile (fig. 47) lying forward over dorsal surface of apical abdominal segments.

♂. Length, 3·75 mm.

Holotype: No. 1165, D. M.

Habitat.—Occurring in large numbers on sandy sea-beach at the mouth of the sluggish Hokio Stream.

H. vector n. sp.

♂. Eyes as in H. flavinceris; ocellar triangle black with brownishgrey tomentum; the pair of ocellar bristles easily distinguished from the smaller posterior series; ocelli amber-yellow; front black with grey tomentum and row of delicate bristles along orbits. First and second

Figs. 38–41.—Brachystoma hamiltoni n. sp.

  • 38. Antenna.

  • 39. Dorsum of thorax, showing chaetotaxy.

  • 40. Wing.

  • 41. Genitalia, ♂.

  • 42. Occiput, ♂, showing chaetotaxy.

  • 43. Dorsum of thorax, showing chaetotaxy.

Figs. 44–47.—Hilara littoralis n. sp.

  • 44. Antenna.

  • 45. Occiput, ♂, showing chaetotaxy.

  • 46. Dorsum of thorax, showing colour-pattern and chaetotaxy.

  • 47. Genitalia, ♂.

Figs. 48, 49.—Hilara vector n. sp.

  • 48. Occiput, ♂, showing chaetotaxy.

  • 49. Dorsum of thorax, ♂, showing chaetotaxy.

Fig. 50.—Hilaria dracophylli n. sp.: dorsum of thorax, ♂, showing chaetotaxy.
Figs. 51–53.—Hilara fumosa.

  • 51. Occiput, ♂, showing chaetotaxy.

  • 52. Dorsum of thorax, ♂, showing chaetotaxy.

  • 53. Wing.

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– 456 –

joints of antennae black but covered with grey tomentum, bristly; third joint and style shaped as in H. flavinceris, blackish-brown with greyish tomentum. Face black but clothed with greyish tomentum, three short delicate bristles on each side beneath each antenna. Proboscis slightly longer than head-height, dark-brown; palpi densely clothed with greyish tomentum, bristles and hairs grey. Cheeks cinereous with greyish hairs; occiput cinereous, hairy below, bristly above, among which only three pairs of bristles are distinct (fig. 48).

Thorax slightly humped, black, but clothed with cinereous tomentum except on a bare testaceous area at posterior angle of each humerus; dorsum with four longitudinal blackish stripes. Chaetotaxy weak except for alar, notopleural, humeral, and post-humeral bristles, which are outstanding; acrostichals and dorsocentrals represented by rows of numerous weak bristles (fig. 49). On scutellum is marginal row of ten erect bristles.

Wings slightly greyish, veins testaceous but paler basally, stigma slightly tawny; Sc ending in costa. Halteres tawny.

Legs testaceous; apex of tibiae and tarsi darker; anterior protarsi blackish and distinctly thickened.

Abdomen with spot-like pits on each segment; black, with cinereous tomentum; posterior margin of each segment except the first pale greyish-tawny. Genital appendage elongate, blackish-brown, but tawny along middle on each side, this colour extending over upper edge basally.

♂. Length, 6 mm.; + gential appendage, 10 mm.

Holotype: No. 954, D. M.

Habitat.—Lake Wakatipu (W. G. Howes).

H. dracophylli n. sp.

♂. Head black, but greyish-pruinose; eyes and front as in H. flavinceris, the latter with a row of delicate black orbital bristles; front black, with indistinct brownish area just above antennae, which are inserted in middle height of head and prominent; first and second joints shaped as in H. flavinceris, tawny, delicately bristled; third joint black, greyish-pruinose, elongate, clavate, terminated by a style about two-thirds its length, from the end of which projects a short, brownish chitinous spine. Face shaped as in H. flavinceris, black, with greyish pubescence. Proboscis about as long as head-height or a little longer, dark testaceous to dark brown, blackish at apex; palpi black with black bristles. Cheeks inconspicuous, black, with a greyish pubescence and clothed with a few delicate dark-brown hairs. Occiput convex, black but greyish-pruinose. hairy below, bristly above the system as in H. flavinceris.

Thorax somewhat humped, black, humeri and notopleurae testaceous with grey tomentum; alar regions testaceous; scutellum testaceous but for a small black area at base; pleurae testaceous immediately beneath wing-articulation. Chaetotaxy as in fig. 50, acrostichals bring represented by several rows of very short delicate bristles, and dorsocentrals, which are uniserial, by somewhat stronger ones. On scutellum are three pairs of strong marginal bristles, a pair of less prominent discal ones, and a pair of apical, convergent delicate bristles; in some specimens discal and apical bristles are absent. Dorsum greyish-pruinose with three indistinct longitudinal black stripes seen in some lights.

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Wings hyaline except for testaceous articulation and brownish cell 2nd C and stigma; veins brown, paler basally; venation as in H. flavinceris. Halteres pale yellow but brownish at base.

Legs testaceous, clothed with delicate hair-like bristles; anterior protarsi slightly thickened; claws rather elongate, dark testaceous basally, black distally; pulvilli testaceous.

Abdomen dark testaceous, with testaceous markings, more or less distinct, across posterior margin of each segment except first, upon which is a triangular testaceous spot along anterior and another along posterior margin, their apices meeting in centre of segment. The whole clothed as in H. flavinceris. Genital appendage (fig. 2a) shaped as in H. flavinceris, dark-brown, with narrow tawny median stripe on each side broadening basally.

♂. Length, 6 mm.; + gential appendage, 10·5 mm.

Holotype: No. 1207, D. M.

Habitat.—New Zealand (locality and collector unknown).

H. fumosa Hutton.

Clinocera fumosa Hutton, Trans. N.Z. Inst., vol. 33, p. 31, 1901.

Owing to the well-developed anal angle of the wing and the shape of the antennae, &c., this species cannot be placed in Clinocera, and it is doubtful if it really belongs to Hilara, although I have placed it therein for the present.

♂. Eyes contiguous over greater part of front, which is confined to a triangular black area with greyish reflection above antennae and to small area in front of ocellar triangle; eyes deeply emarginate; ocellar triangle black, with cinereous reflection; ocellar bristles strong but not long. Antennae black, first two joints short, bristly and yellowish-brown in some lights; third joint clavate, with greyish pubescence; terminal style slender, rather reddish brown, minutely pubescent, two-jointed, first joint very small, the whole a little longer than third antennal joint. Face short but broad, black, but greyish-white in some lights. Proboscis shorter than head-height, rather stout, blackish-brown; palpi not quite as long as proboscis, black, but greyish in some lights, and clothed with black bristle-like hairs. Occiput black, greyish in some lights, convex below, but flat above and concave medianly; among the bristles (fig. 51) is a pair of outstanding rather divergent verticals.

Dorsum of thorax black, with three longitudinal grey stripes; prothorax, incisions of transverse suture, notopleurae, humeri, and alar regions grey, this pattern being indistinct in some lights; pleurae black, with greyish reflections; metapleurae bare; a tuft of bristles above insertion of anterior coxae. Chaetotaxy of dorsum as in fig. 52. Scutellum black, but greyish in some lights, with three pairs of erect marginal bristles, pre-apical ones being weak.

Wings (fig. 53) slightly tinged with brown, veins and stigma brown; anal angle prominent; Sc meeting costa; Cu1 and Cu2 in one line, cell Cu being thus somewhat shorter than cell M.

Legs black, bristly, slender; tarsi normal; anterior coxae not unusually lengthened.

Abdomen elongate, narrow, cylindrical, cinereous but black in groundcolour, distinctly bristly, bristles for the most part hair-like but marginal ones stronger and larger. Genitalia inconspicuous.

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♀. Eyes broadly diclioptic, head rather flat on top; front rather convex, blackish-grey; face rectangular and cinereous; proboscis shiny black, stout, curved backwards, just under twice length of head-height; a distinct pair of long, divergent ocellar bristles. Legs more or less tawny; tarsi and upper sides and middle of femora fuscous; coxae, except apex of anterior pair, cinereous-black.

♂ and ♀. Length, 6 mm.

Holotype: ♂, Hutton's collection, Canterbury Museum; ♀, No. 1212, D. M.

Habitat.—Wellington (G. V. Hudson); Otira (J. W. Campbell).

H. hudsoni Hutton.

Empis hudsoni Hutton, Trans. N.Z. Inst., vol. 33, p. 30, 1901.

♂. Eyes dichoptic, deeply emarginate; occiput, front, and face black, but with cinereous tomentum brownish in some lights; fronto-orbital and ocellar bristles present. Antennae shaped as in H. fossalis; black, but first and second joints brownish. Proboscis a little longer than head-height, black; palpi brown. Occiput bristly, bristles arranged irregularly on each side of centre, which is bare.

Dorsum of thorax black, with a pair of indistinct brownish stripes; silvery spot on each side between humeri and wings; humeri silvery above; scutellum and alar regions along sides of dorsum to sutural incisions brownish; pleurae cinereous; metanotum brownish. Acrostichal bristles represented by four rows of short, weak bristles; dorsocentrals also weak and short but ending in two pairs of stronger bristles before the scutellum; a strong humeral bristle and numerous small ones; a pair of strong notopleurals, anterior to which are some weak ones; a strong supra-alar, posterior to which is a second one and two weak short ones; a strong post-alar; metapleurae bare; scutellar bristles consisting of four pairs, apical pair weak, pre-apical strong, remainder somewhat weaker.

Wings clear, stigma clouded; halteres brown.

Figs. 54–56.—Hilara fossalis n. sp.

  • 54. Occiput, ♂, showing chaetotaxy.

  • 55. Dorsum of thorax, showing chaetotaxy.

  • 56. Genitalia, ♂.

Fig. 57.—Hilara smithii n. sp.: dorsum, ♂, showing chaetotaxy.
Figs. 58–62.—Hilarempis fulvipes.

  • 58. Antenna.

  • 59. Dorsum of thorax, showing chaetotaxy.

  • 60. Wing.

  • 61. Anterior leg, ♂.

  • 62. Genitalia, ♂.

Figs. 63–64.—Hilarempis nigra n. sp.

  • 63. Dorsum of thorax, ♂, showing chaetotaxy.

  • 64. Genitalia, ♂.

Figs. 65–68.—Empimorpha torrentalis n. sp., ♂.

  • 65. Head, front view.

  • 66. Occiput of head, showing chaetotaxy.

  • 67. Dorsum of thorax, showing chaetotaxy.

  • 68. Genitalia.

– 459 –
– 460 –

Anterior and middle legs brownish, posterior cinereous brown to black; tarsi black; coxae cinereous; anterior protarsi enlarged.

Abdomen brownish to greyish black, clothed with very weak, short bristles; genital appendage much shorter than the abdomen, brown, the genitalia brownish-black.

♀. As male, but the abdomen conical and terminated by a pair of stylets.

♂. Length, 5·5 mm. ♀. Length, 4 mm.

Holotype: Hutton's collection, Canterbury Museum.

Habitat.—Wainuiomata; Otira Gorge (G. V. Hudson); Okere.

H. fossalis n. sp.

♂. Eyes dichoptic, deeply emarginate; ocellar triangle and front black; a pair of conspicuous ocellar bristles, behind each of which is row of small delicate ones; row of bristles on each side of front. Antennae black, inserted about middle height of head; first and second joints bristly; third joint clavate and terminated by a comparatively short style, which ends in a bristle-like appendage. Face black, with greyish reflection. Proboscis as long as head-height, brownish, but tawny along lower side; palpi black. Occiput convex at orbits, but depressed medianly; chaetotaxy, as in fig. 54.

Thorax black; the humeri, pleurae, two indistinct stripes on dorsum, scutellum, and a transverse band in front of scutellar suture greyish in some lights; metapleurae bare; chaetotaxy of dorsum distinct (fig. 55).

Wings smoky, rather short and blunt, stigma clouded; halteres brownish, heads somewhat darker.

Legs bristly inclined to tawny, upper sides of femora and tarsi black; anterior protarsi enlarged.

Abdomen black, slightly greyish in some lights, with distinct bristles towards posterior margin of each segment; genitalia prominent, brownish-black, appendage broad and not quite length of abdomen (fig. 56).

♂. Length, 4 mm.

Holotype: No. 1199, D. M.

Habitat.—Makerua Swamp, skimming over water; Ohau beach, frequenting damp sand bordering a sluggish stream.

H. smithii Hutton.

Empis smithii Hutton, Trans. N.Z. Inst., vol. 33, p. 30, 1901.

♀. Head lost. Dorsum of thorax brownish-grey, with three golden stripes; chaetotaxy distinct (fig. 57); six erect marginal bristles on scutellum, three on each side, apical ones convergent; pleurae, metanotum, and scutellum cinereous; metaplurae bare. Legs tawny and bristly. Wings ochreous, inclined to translucency; R4 branching from R5 at an acute angle; halteres tawny.

Abdomen cinereous, ending in pair of stylets; posterior margin of each segment rather paler; the whole clothed with minute black bristles.

The type specimen, which is the only one of this species yet secured, is badly damaged, and a fuller account is not possible in consequence.

♀. Length, 5 mm.

Holotype: Hutton's collection, Canterbury Museum.

Habitat.—Ashburton (Smith).