
(A.) Waihaoia n. subgen.
Shell rather small, fusiform. Spire nearly as high as aperture. Nucleus scaphelloid, of 2 or 3 smooth whorls. Body-whorl contracting gradually;

fasciole absent. Sculpture of axial ribs indistinctly angled to tubercular. Aperture with a very shallow anterior sinus. Outer lip simple, thin or slightly thickened. Columella of type with 5 strong plaits on a convex pad; some species have only 4.
| A. Axial ribs only. | |
| 1. Narrowly fusiform | allani. |
| 2. Broadly fusiform, spire gradate | rugosa. |
| Broadly fusiform, spire conic | regularis. |
| B. Axial ribs on spire, but later disappearing | scitula. |
| C. Axial ribs on spire, but later becoming slightly tubercular. | |
| 1. Shell very narrow, anterior end long, 9 axials | thomsoni. |
| 2. Anterior end short, tubercles sharp, 12 axials | bathgatei. |
| 3. Tubercles blunt, 8–12 | superstes. |
| D. Axial ribs on spire, but later becoming strongly tubercular. | |
| 1. Anterior end long, tubercles moderate | suteri. |
| 2. Anterior end short, tubercles very strong | phymatias. |
| E. Strongly tubercular on both spire and body. | |
| 1. Sharp tubercles nearer anterior suture | aculeata. |
| 2. Rather blunt tubercles nearer posterior suture | confusa. |
| 3. Shell very broad, nucleus flat | dyscrita. |
Waihaoia allani n. sp. (Plate 61, fig. 5.)
Shell small, fusiform. Spire conic, about as high as aperture Nucleus scaphelloid, of about 3 smooth whorls. Post-embryonic whorls 4 ½, at first convex, then bluntly angled about middle; body-whorl with short steep shoulder, below which it is straightened, then moderately contracted; fasciole not marked. Sculpture: spire-whorls with 12, body-whorl with 14, strong axial ribs, bluntly angled on shoulder and twisted backward, interstices equal or slightly wider; on spire, ribs extend from suture to suture and well down on body-whorl. Aperture moderate, angled above with extremely shallow sinus below. Outer lip thick, not reflexed, slightly convex, a little ascending. Columella with pad bearing 5 strong folds, anterior weakest. Outer lip with thin glaze.
Holotype in Dominion Museum.
Height, 45.5 mm.; diameter, 16 mm.
Locality.—Greensand, McCullough's Bridge, Waihao River.
The nucleus of the holotype is not normal, having the appearance of being unrolled.
Waihaoia regularis (Finlay). (Plate 61, fig. 6.)
| 1926. |
Alcithoe regularis Finlay, Trans. N.Z. Inst., vol. 56, p. 247, pl. 55, fig. 4. |
In general appearance this shell resembles M. clifdenensis; but if any reliance can be placed on recapitulation as shown by ornamentation, &c., the relationship is not so close as at first glance appears. The post-embryonic whorls are strongly axially ribbed from suture to suture right from the neanic stage, whereas ribs do not appear in M. clifdenensis until the penultimate whorl. Also, the anterior notch in W. regularis is not so deep nor the fasciole so well defined. Further, although the nucleus is damaged, enough remains to show, in the writer's opinion, that it had a flattened top, as in the W. bathgatei group. The shallow anterior notch agrees with this systematic position. W. regularis represents a broadened offshoot from W. bathgatei, or earlier, for the sculpture is of simple axial ribs throughout.
Locality.—Clifden (band 8).

Waihaoia bathgatei (Finlay). (Plate 61, fig. 7.)
| 1926. |
Alcithoe bathgatei Finlay, Trans. N.Z. Inst., vol. 56, p. 248, pl. 56, figs. 4, 5. |
The spire is about equal in height to aperture, and the whorls have each about 16 narrow ribs with wide concave interspaces. The ribs soon become angled, and finally short sharp tubercles are developed, but the ribs still extend to the suture above and well down over the body-whorl. The anterior notch is rather shallow, and on some specimens the fasciole is quite convex, it is not rough nor bounded by a ridge; the columella has 4 folds, decreasing anteriorly; sometimes a fifth weak fold appears posteriorly. The apex of the nucleus is generally flattened, but occasionally somewhat domed.
This species seems to be a development from such forms as W. suteri, the anterior end of the shell having shortened considerably and the notch become slightly deeper.
Locality.—Clifden, Southland (bands 4, 6a).
Waihaoia phymatias (Finlay). (Plate 61, fig. 11.)
| 1926. |
Alcithoe phymatias Finlay, Trans. N.Z. Inst., vol. 56, p. 248, pl. 56, figs. 1–3. |
This shell is a descendant of W. bathgatei in which the tubercles on the shoulder have greatly enlarged, with a consequent loss of the axial ribs, the bathgatei stage being represented about the fourth conch-whorl. The anterior notch is still shallow, and the fasciole is not marked off from the base of the body-whorl. The nucleus is somewhat larger, and is a little more flattened on the summit.
Locality.—Clifden (band 6b).
Waihaoia dyscrita (Finlay). (Plate 61, fig. 12.)
| 1926. |
Alcithoe dyscrita Finlay, Trans. N.Z. Inst., vol. 56, p. 249, pl. 55, fig. 9 |
An accelerated offshoot from W. phymatias in which the height has greatly decreased compared with the diameter. The top of the nucleus is broken in the single specimen found, but there is little doubt that it was the same as that of A. phymatias. Further points of agreement are the columella with 4 mitriform folds, and the shallow anterior notch not forming a well-defined fasciole. Only the first 2 or 3 axials are without a tubercle, so that the ancestral ornamentation, occupying 2 or 3 whorls in W. bathgatei and W. suteri and 1 whorl in W. phymatias, has almost disappeared.
Locality.—Clifden (band 6c).
Waihaoia confusa n. sp. (Plate 61, fig. 3.)
| 1917. |
Lapparia hebes (Hutton): Suter, N.Z. Geol. Surv. Pal. Bull. 5, p. 40 (in part, but not of Hutton), pl. 12, fig. 12. |
Shell somewhat small, elongate fusiform. Spire strongly gradate, about two-thirds height of aperture. Diameter of spire-whorls about twice their height, and with a concave shoulder; body-whorl slightly convex, very gradually contracting, with a well-marked fasciole not bounded above by a ridge. Suture undulating. Sculpture of 7 sharp tubercles on the shoulder, not produced into definite axial ribs. Aperture oblong, angled above, deeply notched below. Columella with 4 plaits, the upper 3 strong, more than equal to the interspaces, the lowest very weak.
Holotype in collection of New Zealand Geological Survey.
Height, 33 mm.; diameter, 15 mm.
Locality.—164, greensand, Kakahu.

Hutton's lost type of Pleurotoma hebes has now been recovered, and it shows how far Suter went astray in his selection of plesiotypes. The original shell is certainly not a Volute, but is congeneric with the shell described by Suter as Ptychatractus nodosoliratus. Of the two specimens figured by him as plesiotypes, one (pl. 12, fig. 11) is a Mauia biconica, the other (pl. 12, fig. 12) belongs to the species described above. Many specimens of another shell, Mauia angusta (Suter), were also identified by Suter as Lapparia hebes.
Lapparia, a genus from the North American Eocene, has strong spiral ornamentation reticulated by the growth-lines, and a deep anterior notch.
Waihaoia thomsoni n. sp. (Plate 61, fig. 1.)
Shell rather small, narrowly fusiform. Spire turreted, in height equal to or a little greater than aperture. Whorls 6 remaining, at first probably convex, but soon developing a sloping shoulder, on later whorls shoulder-angle gets lower until finally it is in middle of whorl, above it whorl is slightly concave and very steep; body-whorl subcylindrical for some distance below shoulder, then contracting quickly, finally slowly to a rather long neck, which is bent outwards in type but is straightened in a paratype, and bears an inconspicuous fasciole. Sculpture: spire-whorls with 9 narrowly rounded axial ribs with wide interstices, on later whorls these decrease in number and relative length and strength until on body-whorl there are only 6 obsolete knobs. Aperture long and narrow, with sub-parallel sides contracting to angle above; anterior sinus shallow, the effect rather increased by outward twist to canal. Outer lip broken but apparently straight, thin, not reflexed or ascending. Columella long, projecting slightly, bent, outwards below, bearing 4 well-spaced folds, anterior and posterior weakest. Inner lip with well-defined outer edge.
Holotype in Dominion Museum.
Height, 52 mm.; diameter, 15 mm.
Locality.—Greensand, Waihao Downs.
Distinguished from W. suteri by the fewer and much weaker ribs, also by the disposition of the columellar folds.
A crushed shell from Hampden in Dr. Marshall's collection at Wanganui Museum perhaps is related to this species. It has ribs on the spire-whorls, but they are obsolete on the body. The length is about 73 mm., and the columella bears 5 plaits.
Waihaoia suteri n. sp. (Plate 61, fig. 2.)
Shell rather small, narrowly fusiform. Spire turreted, equal in height to the aperture. Nucleus damaged but apparently scaphelloid. Post-embryonic whorls 6, early ones convex but soon developing a shoulder which later becomes concave and prominent; body-whorl for a short distance below shoulder almost cylindrical, then contracting rather quickly at first but afterwards more slowly so as to produce a rather long neck which has practically no fasciole. Sculpture: early whorls with 12 narrow high axial ribs extending from suture to suture; on third whorl these ribs become angled and have a short concave shoulder; on later whorls their number is reduced to 11 and finally to 10; angulation increases in strength, and above it ribs are much weaker though they persist to suture; on body-whorl ribs extend down on to contracting base but do not reach neck. Aperture long, with subparallel sides, angled above, scarcely notched below. Outer lip broken, but apparently thin and straight, not ascending or

reflexed. Columella with 4 narrow, well-spaced plaits decreasing in strength anteriorly. Inner lip thin.
Holotype in Canterbury Museum.
Height, 50 mm.; diameter, 17 mm. (circa).
Locality.—Greensand, Waihao Forks.
By some mistake this specimen was figured by Suter (1915, pl. 4, fig. 14) as the holotype of Fulguraria aculeata (Hutton), and a wrong magnification given.
Waihaoia superstes n. sp. (Plate 61, fig. 10.)
Shell rather small, narrowly fusiform. Spire turreted, slightly less in height than aperture. Whorls 5 remaining, angled above mid-point, and with a sloping slightly concave shoulder; body-whorl straightened below shoulder, then contracting rather quickly, finally again straightening out to an inconspicuous though convex fasciole which is not definitely bounded above. Sculpture: early whorls with 14 or 15 low axial ribs which scarcely reach sutures, and which are developed into low tubercles on shoulder-angle; on later whorls axials become shorter and fewer, and tubercles stronger; on penultimate there are 12 and on body-whorl 8. Aperture long, sides subparallel, anterior sinus very shallow. Outer lip slightly convex and ascending. Columella with 4 rather weak spaced plaits. Inner lip moderate.
Holotype in collection of New Zealand Geological Survey.
Height, 44 mm.; diameter, 16 mm.
Locality.—894, Mokau River.
This shell is very like W. thomsoni except that the anterior end is shorter. It probably belongs to a northern stock that has descended from W. thomsoni independently of the Southland species W. bathgatei, &c., for the anterior sinus shows no sign of deepening.
This is the Fulgoraria arabica (Hutt.) of Suter (Henderson and Ongley, 1923, p. 36).
Waihaoia aculeata (Hutton). (Plate 64, fig. 6.)
| 1885. |
Voluta aculeata Hutton, Trans. N.Z. Inst., vol. 17, p. 325. |
| 1915. |
Fulgoraria (Alcithoe) aculeata (Hutton): Suter, N.Z. Geol. Surv. Pal. Bull. 3, p. 31, pl. 4, fig. 14. |
The shell figured by Suter is not the holotype. The statement to this effect in the explanation of plates (Suter, 1915, p. 63) errs also in stating that the figure is “enlarged 1 ½ diameters.” The original specimen from which the photograph was taken is in Canterbury Museum, and comes from Waihao Forks. It is described above under the name Waihaoia suteri n. sp. Suter's description of W. aculeata is, however, correct, being based on the true type specimens. Important features of the shell are the small size; straight, sharp outer lip; sharp spines on last whorls developed from strong angled axial ribs on the early whorls; shallow anterior notch which makes an inconspicuous fasciole.
The locality was given by Hutton as White Rock River, but the specimens are not from the bed usually collected from. Their matrix has a large amount of a greenish mineral that appears to be glauconite, so further collecting is needed to clear up a doubt as to the age of this species.
Waihaoia rugosa n. sp. (Plate 70, fig. 4.)
Shell of moderate size, fusiform. Spire almost as high as aperture. Whorls bluntly angled with a long sloping shoulder; body-whorl inflated,

contracted relatively quickly to the large fasciole. Suture undulating. Sculpture of 20 strong axial ribs with equal interstices, ribs twisted and extending from suture to suture on spire and across body-whorl almost to fasciole; on spire-whorls are numerous somewhat irregular spiral lirae crossed by fine close growth-lines. Aperture with wide but shallow anterior sinus. Columella with 4 strong folds with interspaces twice their width. Inner lip extending from aperture as a thick callus.
Holotype in collection of New Zealand Geological Survey.
Height, 54 mm.; diameter, 25 mm.
Locality.—1038, Flat Trig., North Shore, Palliser Bay.
A. rugosa is probably related to Voluta corrugata var. B Hutton. The latter, however, is much larger, and has 5 strong folds and 1 weak one on the columella. W. rugosa differs from the typical Waihaoia in having a well-developed fasciole. The sinus which forms it, however, is fairly shallow.
Waihaoia scitula n. sp. (Plate 64, fig. 5.)
Shell small, thin, broadly fusiform. Spire turreted, one-half height of aperture. Nucleus missing. Post-embryonic whorls prominently angled about the middle, with a fairly broad concave shoulder; body-whorl for some distance cylindrical, then contracted to a short neck bearing an insignificant fasciole not bounded above. Sculpture of about 20 broad axials on early whorls reaching from suture to suture but very weak on shoulder, on penultimate whorl these become obsolete, and body-whorl is quite smooth. Aperture triangular, rather wide, narrowly channelled above, with a shallow anterior sinus. Outer lip thin, straight, not ascending or reflexed. Columella with 4 strong spaced plaits. Inner lip thin, restrained.
Holotype in Canterbury Museum.
Height, 31 mm.; diameter, 14 mm.
Locality.—Porter River, Trelissick Basin.
“Scaphella gracilis (Swainson),” according to Hutton's tablet. Easily distinguished from that species by the general shape, concave shoulder, and shallow anterior notch. It is not closely related to any other species so far collected.
