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Volume 59, 1928
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Vegetation of the Upper Bealey River Basin, with a List of the Species.

[Read before the Philosophical Institute Of Canterbury, April 4th, 1928; received by Editor, 4th October, 1028; issued separately, 25th March, 1929.]

This paper proposes to give the names of the flowering plants and ferns collected in the Basin of the Bealey between the mouth of the Mingha and the Western Divide. It is strange that hitherto no comprehensive list of the plants of this area has been attempted. Many botanists have visited the district, and no portion of the Southern Alps is better known to visitors and tourists, many thousands of whom annually cross Arthurs Pass. The authors therefore desire to provide such with a reliable list of the species likely to be met with, and to furnish a basis for the further study of the florula of the district. As much of the area is practically inaccessible the list cannot be complete, but it probably contains about ninety per cent. of the species. A few records of plants not seen by the authors from the writings of earlier investigators have been included.

Characteristics of the Flora.

The Upper Mingha and many of the higher valleys and ridges have not been visited by us. The region is indeed inexhaustible, and though the vegetation at similar elevations is fairly uniform, there are doubtless a good many rare and local species that have not yet been collected. For the present also, the more critical forms, some hybrids and novelties, are omitted. Some of the genera, particularly Celmisia, Olearia, Coriaria, Acaena, and possibly Epilobium, swarm with hybrids.

A collection has been made including nearly all the species listed, and may be viewed at Arthurs Pass by those interested. It is hoped that thus some, who have only had a casual knowledge of the subject, may be roused to take an interest in the scientific side of it. Unfortunately at present the roadsides and neighbouring hills and streams are rapidly being despoiled of their characteristic vegetation by tourists who gather armfuls of the blossoms of Metrosideros, Elytranthe, Ranunculus Lyalli, Celmisia, Senecio and of other genera, in many cases only to leave them, to die by the roadside. If this depredation can be reduced by a fuller understanding of the beauty and uniqueness of our native flora, there may be some hope of preventing the harm that is being done at present to the plants on the sides of the main road over the pass and on the most travelled tracks.

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Some remarks are here offered as to the general characteristics of the vegetation of the district. The florula is comparatively limited in species, no doubt for the following main reasons:—

(1)

High evaporation rate.

(2)

Lack of soil.

(3)

Low winter temperature.

(4)

Frequent heavy gales.

(5)

Prejudicial effect of beech forest on the germination of seedlings.

A few remarks will be offered under each head:

(1) High evaporation rate.—In spite of the huge rainfall, averaging about 180 inches in the year, the atmosphere is a dry one, and the plants are those of a comparatively dry area. This somewhat paradoxical fact is due to the effect of the mountain range on the moisture contained in the prevailing northwest wind. As the wind-descends the Southern Alps it becomes warmer, and rises higher and higher in temperature above the dew point, so that finally the rate of evaporation becomes extremely high. Unfortunately no records-are available to illustrate this, but there can be no doubt that in spite of the high precipitation the atmosphere is usually dry. The rainfall as recorded at the Arthurs Pass Railway Station from 1924-1927 is as follows: 1924, 160 in.; 1925, 201 in.; 1926, 205 in.; 1927, 171 in. As a rule the rain, which comes mostly from the northwest, falls heavily and disappears quickly. On one occasion 17 in. fell in twenty-four hours, but the very heavy rainfall does not preclude occasional periods of drought. Thus in January 1928 the rainfall was considerably under an inch, and for the first three months of 1928 it was only 14 in. During January the vegetation was in many places parched or even killed, and the rain of March and February fell in short periods of time. During the whole period the streams were unprecedentedly low. On the other hand, in the first three months of 1927, 58 in. of rain fell. An illustration of abnormal evaporation may be given. Mr. S. Page informs us that he has seen light rain falling from a low cloud at Arthurs Pass to within ten or twelve feet of the ground and then evaporating. This unusual phenomenon was doubtless exceptional, but it, and the slow rusting of iron that is apparent, obviously could only occur with an exceptionally high rate of evaporation. The complete absence of miro, matai, rimu, and kamahi, and almost complete absence of rata, though all of these are abundant in the Otira Valley, are no doubt in part due to this cause. Similarly the larger and softer-leaved coprosmas and olearias are chiefly confined to the western side of the range, or the higher valleys on the eastern aide; e.g. Olearia Colensoi is almost entirely a plant of the west, Coprosma lucida is abundant in the Otira Valley, but so far only one plant has been found on the eastern side. Generally speaking the forest and sub-alpine scrub of the eastern side consists of small-leaved plants.

(2) Lack of Soil—As the hillsides and valleys are up to about 4,000 ft. or 4,500 ft. everywhere covered with loose shingle, in places overlain by two or three inches of soil, the rain rapidly sinks through

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below the surface-level, and much of it passes away in under-ground streams. The lack of soil tends to limitation of species, and the limitation of species tends m turn to lack of soil. It is only in flat places, therefore, which soon turn into bog, that the ground is always damp, though bog may also be found on hillsides near stream courses, near under-ground streams, or where there is no surf ace shingle. The result is that the vegetation of the hillsides is generally xerophytic. Such plants of dry districts as Exocarpus Bidwillii, Hebe Lycopodioides, Discaria toumatou, are abundant in certain situations within the valley. Here and there on the river-valleys there is a comparatively deep rich soil, no doubt formed in succession from some old sphagnum bog, but the steepness of the hills and the presence of boulders, angular rocks, and shingle slopes apparently prevent the accumulation of humus elsewhere. Another contributing factor is the almost complete absence of dust from the atmosphere, so that even the river-bed raoulias do not develop the thick cushions that, they do in the lower plains, where the abundant dust blown before the northwest wind-storms tends to build them up into ever higher and higher cushions. Raoulia australis and R. tenuicaulis here lie flat on the ground, though R. Haastii forms cushions up to 3 or 4 inches in thickness. Where the river-bed is not subject to floods, under the thin layer of humus may be found four or six inches of clay. Only in the older bogs or in soil consolidated from these bogs is there any depth of humus.

(3) Low winter temperatures.—Another factor limiting the number of species present is the extreme winter cold, due in part to altitude, in part to the fact that there are here narrow valleys shut in on all sides by mountain ridges, and in part to the proximity of high snow-covered mountain peaks. The area dealt with lies roughly between 2,200ft. altitude and 6,000 ft., the limit of vegetation, though several plants have been found at 7,000 ft. on Mount Rolleston. This, which is the dominating peak of the district, is roughly 7,400 ft. in height. Unfortunately again no figures are available for winter temperatures, though a minimum thermometer has now been placed in the district; but there can be no doubt that in winter frosts the thermometer falls to zero Fahrenheit. In the winter of 1925 snow fell to the depth of 2 ft. 6 ins. in the settlement (2,400 ft. elevation), and of 3 to 5 ft. in the col of the pass (3,000 ft. elevation). This was undoubtedly exceptional, for as a result a snow-field was formed in the head of the Bealey river, which as it slipped downwards bore away the Nothofagus forest, trees being destroyed which had been growing from twenty-five to forty years.

(4) Wind.—For about nine months in the year the area is subject to strong, sometimes violent, gales from the northwest. The months of May, June and July, are usually fairly calm. The only other wind that is at all prevalent is the southwester which alternates with the northwest; but probably there is only one day southwest for every three days of northwest wind. Of course the direction of the winds is largely controlled by the direction of the valleys, so that in Bealey valley the southwester of the plains is approximately southeast in direction. Though there is little evidence in storm-

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twisted or brush-back trees, of the general direction of the wind, it is undoubtedly one of the factors in limiting forest growth; and the absence of forest from the sides of the hills at the top of the pass in evidently in part due to the strength of the winds there. Usually the forest line lies between 4,000 and 4,500 ft.; but here it is only 3, 000, ft. The winds must of course immensely increase the rate of evaporation and so add to the dry ness of atmosphere.

(5) The effect of the Nothofagus forest on undergrowth.—It is well known that the Nothofagus forests are freer from undergrowth than any other type of New Zealand forest. In the lower Mingha valley the forest is nearly pure Nothofagus cliffortioides. In the Bealey proper there is a much greater intermixture of species. Where the dry hard beech-leaves fall and accumulate and there is a heavy canopy of beech branches overhead, little undergrowth apparently can exist. The beech-leaves only slowly turn into humus, and no good seed-bed is formed. Though not so prejudicial to growth as pine needles, they seem to have a similar effect. Owing to the ravages of man in the Bealey valley, and in more recent years to those of a bud-burrowing larva, the beech forest has suffered much, resulting in a greater and greater development of undergrowth. At present indeed insect larvae are doing much harm to many different species, those especially attacked are Nothofagus cliffortioides, Phyllocladus alpinus, species of Coriaria and Ranunculus Lyallii. Hoheria glabrata is also subject to pests, but comes up in large quantities where the beech has been cut down. It is not intended to discuss the problems of succession here, but merely to indicate that the forest is by no means stable. Obviously under conditions such as those described, the ordinary coastal vegetation of such an area as Banks Peninsula cannot exist, and consequently we find that the forest trees and shrubs of the coastal area are for the most part absent here. Thus tree-ferns are unrepresented except by the low-growing Alsophila Colensoi, and only three species of filmy fern have so far been discovered, though many species are abundant at Otira. Hedycarya, Carpodetus, Pittosporum eugenioides, species of Sophora, Pennantia corymbosa, Melicytus ramiflorus, Hoheria angustifolia, Plagianthus betulinus, Schefflera digitata, Nothopanax arboreum, and other common trees and shrubs of the coastal area are missing here. Indeed the only conspicuous forest trees of Banks Peninsula to be found in the Bealey watershed are Griselinia littoralis, Aristotelia serrata (chiefly in sheltered gullies), and Fuchsia excorticata. The two latter no doubt owe their presence to the fact that one is partly and the other totally deciduous in winter. Of course these differences in the lowland and sub-alpine forests are known to all observers, but we think it well to call special attention to them in order to emphasize the existence of the conditions that we are dealing with. Mention should also be made of the almost total absence of lianes and phanerogamic epiphytes; Clematis indivisa for example occurs in the Otira Gorge but not in the upper Bealey Valley; Muehlenbeckia australis is also absent, and the only lianes or scramblers common are two species of Rubus. There are also so far as noted no flowering epiphytes here, the forest in consequence having none of that sub-tropical appearance so frequently noted in the New Zealand

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bush. The absence of lianes and epiphytes, which are mostly plants of warmer regions, is doubtless due to the low winter temperature. However, the total number of species present is apparently somewhat “higher than in the Mt. Cook region, where the flora has been listed by Professor A. Wall. There the higher mountains and the absence of a low pass probably account for the smaller number of species.

One other point already referred to in these introductory notes should be further emphasized, and that is the remarkable contrast in the vegetation of the eastern and western sides of the range; of this ~we can give no complete explanation, nor do we propose to consider it at length here, but the least observant cannot fail to notice the comparative richness of the Otira plant life compared with that in Arthurs Pass. The township of Otira is some 1,250 ft. lower than the Arthurs Pass settlement, very much warmer, has doubtless a much moister atmosphere and a richer soil, and to these conditions much of the greater wealth of species must be due; but further explanations are probably needed to account for the absence of certain spceies from the colder area. These, however, can only be dealt with when the vegetation of the western side of the range is more fully known.

The Plant Formations.

The vegetative covering of the Bealey Valley basin includes n fairly representative sample of the sub-alpine and alpine plant-formations of New Zealand. The series extends from mountain beech-forest through bog, scrub and tussock to fell-field at the upper limit of vegetation. In the list of species appended to this paper the main plant-formations in which each is found is given in a general way. What these formations consist of will now briefly be indicated.

Forest.—Forest covers the mountain sides up to an altitude of about 4,500 ft. except in the pass itself where it is quite absent. Its limits correspond precisely to the area exposed to the full force of the northwest wind. The dominant tree, and for the most part the only tall tree in the forest, is the mountain beech, Nothofagus cliffortioides. Rarely is a tree of the pahautea, Libocedrus Bidwillii, met with. Other trees present, such as Nothopanax simplex and Griselinia littoralis belong to a lower tier of foliage. The undergrowth, consisting of shrubs with a few ferns and herbaceous plants, is not dense except in places where a valley bottom somewhat permits the soil to retain its water.

A subassociation of the forest is that formed where Dacrydium biforme mixes with the stunted trees of the beech. This occurs in situations unfavourable to tree-growth, namely in boggy ground and on exposed ridges.

Scrub.—Three distinct scrub-formations are present in the Bealey Basin. On the river bed is Discaria scrub, but only a fragment is left, and of this the original species are almost all exterminated except the Discaria and a Coprasma. In the forest-belt on exposed ridges is a low scrub in which the principal members are Leptospermum scoparium and Exocarpus Bidwillii. Above the forest-belt is sub-alpine scrub, in which species of Dracophyllum, Phyllocladus, and

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various composite shrubs play a conspicuous part. Several associations can be defined.

Tussock.—In situations inimical to shrub-growth either on account of exposure, or of deficient drainage, tussock-formation occurs. Danthonia Raoulii is dominant over most of the area, but along the higher levels D. crassiuscula is the principal species.

Bog occurs on all fairly level stretches of land either in the forest zone or above it. Its dominant life-forms are cushion-plants, of which Donatia novae-zealandiae and Gaimardia setacea are perhaps the most conspicuous.

Cushion-plant associations other than bog include the shingle-bed of the Bealey River, in which species of Raoulia are most conspicuous, and some open associations above the scrub-line. These alpine cushionplant associations are generally situated on seepage-slopes, and hence the habitat differs from that of bogs in being well drained. Phyllachne clavigera and Anisotome imbricata are the dominant species.

Meadow.—The alpine meadows are among the most wonderful sights to be seen on the mountains. They border streams at altitudes of from 3,000 to 5,000 ft. The dominant plants are showy species of Senecio (S. Lyallii and S. scorzoneroides) and Ranunculus Lyallii.

Fell-field.—Under this term is included the open rocky portions of the mountain from about 5,000 ft. to the upper limit of vegetation. The plants occur in clumps or singly in crevices among the rocks. They include cushion-plants like Hectorella and Pygmea, herbs like Senecio scorzoneroides, and shingle-plants like Hebe Haastii.

Shingle-slips are common on the mountains but only a few true shingle-slip plants are present. Haastia Sinclairii is the most common shingle plant. These slips, however, are not so much consolidated as in the mountains to the eastward, and are composed of larger and perhaps more angular fragments.

Reed swamp is characterized by the presence of species of Carex, Heleocharis and Juncus. It occurs on the Bealey river bed where Heleocharis acula is common, and within the forest-zone where Carex ternaria is the most prominent species.

Water formations occur by streams in the main river-bed and on the mountains. Two formations at least may be distinguished, one in which Potamogeton and Myriophyllum occur, and another with Montia and Epilobium macropus as the most conspicuous members.

List of Species of Pteridophytes and Spermophytes.

*Species not observed by the authors.

Lycopodiaceae.
Lycopodium Selago Linn. Scrub.
L. novae-zealandicum Col. Forest.

We have given this form the rank of species as it appears to us to be as much entitled to it as is L. varium. It is possible that the three forms—Billardieri, varium, and novae-zealandicum are but habitat forms of one polymorphic species. In any case we consider them each of equal status.

L. fastigiatum R. Br. Bog. tussock.
L. scariosum Forst. Forest.
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Hymenophyllaceae.
Hymenophyllum flabellatum Lab. Forest.
H. villosum Col. Forest.
H. multifidum (Forst.) Sw. Forest, scrub.
Gleicheniaceae.
Gleichenia dicarpa R. Br. var. alpina (R. Br.) Hook. f. Bog.
G. Cunninghamii Heward. Forest on ridge.
Osmundaceae.
Leptopteris superba (Col) Presl. Forest.
Schizaeaceae.
Schizaea fistulosa Lab. Bog.
Cyatheaceae.
Alsophila Colensoi Hook. f. Forest.
Polypodiaceae.
Cystopteris fragilis Bernh. Forest.
Polystichum vestitum (Sw) Presl. Forest, scrub.
P. cystotegium (Hook) J. B. Armstr. Shingle slips.
Asplenium Richardi Hook. f. Forest.
A. trichomanes Linn. Forest (Halpins Creek).
Blechnum (Lomaria) procerum (Forst.) Forest.
B. (L) fluviatile (R. Br.) Salom. Forest.
B. (L) penna-marina (Poir.) Kuhn. Forest.
B. (L) lanceolatum. (Spreng.) Sturm By side of creek in forest.
B. (L) vulcanicum (Blume) Kuhn. Forest.
Hypolepis millefolium Hook. f. Forest.
Histiopteris incisa (Thunb.) J. Sm. In second-growth forest and scrub.
Pteridium esculentum (Forst.) Ckne. Forest on ridge.
Paesia scaberula (A. Rich) Kuhn. Forest on ridge.
Polypodium Billardieri (Willd.) C. Chr. Forest.
P. pumilum (Armstr.) Ckne. Cushion plant fornation.
Pinaceae.
Libocedrus Bidwillii Hook. f. Forest.
Podocarpaceae.
Podocarpus Hallii Kirk. Forest.
P. nivalis Hook. Forest, scrub, tussock.
P. Hallii X nivalis Forest.
Dacrydium biforme (Hook) Pilg. Bog and dry forests.
D. laxifolium Hook. f. Bog, scrub.
Phyllocladus alpinus Hook. f. Scrub.
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Naiadaceae.
Potamogeton Cheesmanii Benn. Water in river-bed.
Gramineae.
Hierochloe redoleus R. Br. Forest, scrub.
Ehrharta Colensoi Hook. f. Shingle slips, tussock.
Deyeuxia Forsteri (Roem et Schult). Kunth Forest.
Agrostis subulata Hook. f. Fellfield, Senecio meadow.
A. Dyeri Petrie. Tussock.
Deschampsia Chapmani Petrie. (Recorded by Cheeseman).
Trisetum Youngii Hook. f. Tussock.
Danthonia Raoulii Steud. Tussock.
D. flavescens. Hook. f. Forest, tussock, swamp.
D. crassiuscula Kirk. Tussock.
D. oreophila Petrie. Forest, tussock, fellfield.
D. pilosa R. Br. Tussock.
D. Cunninghamii Hook. f. Forest edge.
Poa Lindsayi Hook. f. River-bed.
P. novae-zealandiae Hack. Forest, fellfield. Mt. Rolleston, 7,000 ft. (Miss E. Washbourn).
P. Colensoi Hook. f. var. breviligulata Petrie. Scrub, tussock, fellfield.
P. Cockayneana Petrie. Scrub.
Festuca novae-zealandiae (Hack) Ckne. Tussock.
*Agropyrum aristatum (Petrie) Cheesm. (Recorded by Cheeseman).
Cypekaceae.
Heleocharis acuta R. Br. Swamp in river-bed.
Scirpus cerrnuus Vahl. Ranunculus meadow.
Carpha alpina R. Br. Bog.
Schoenus pauciflorus Hook. f. Forest, bog, Senecio meadow, tussock.
Oroeobolus strictus Berggr. Bog.
O. pumilio R. Br. Bog, tussock.
O. pectinatus Hook. f. Bog.
Uncinia compacta R. Br. River-bed.
U. caespitosa Boott. Forest, scrub.
U. filiformis Boott. Forest.
U. riparia R. Br. Forest.
U uncinata (L) Kukenth. Forest.

We have accepted uncinata as the correct specific name, as was done by Kukenthal, because we consider that the name uncinata is sufficiently distinct from Uncinia. Furthermore it is probable that article 55 of the International Rules will be amended at the Botanical Congress of 1930.

U. leptosiachya Raoul. Forest.
Carex pyrenaica Wahl. Tussock.
C. virgata Sol. Swamp.
C. stellulata Good. Bog.
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C. leporina Linn. River-bed, swamp.
C. ternaria Forst. Swamp.
*C. Berggreni Petrie (Recorded by Cheeseman).
C. inversa R. Br. River-bed.
C. acicularis Boott. Cushion plant formation.
C. dissita Sol. var. monticola Kuk. Swamp, river-bed.
C. lagopina Wahl. Senecio meadow.
C. Buchanani Berggr. Scrub.
C. Sinclairii Boott. Swamp.
Centrolepidaceae.
Gaimardia setacea Hook. f. Bog.
G. ciliata Hook. f. Bog.
Juncaceae.
Rostkovia (Marsippospermum) gracilis Hook. f. Cushion plant formation, and fellfield.
Juncus polyanthemos Buchen. Swamp.
J. lampocarpus Ehr. Swamp.
J. tenuis Willd. Swamp.
J. novae zealandiae Hook. f. Bog, swamp.
Luzula Colensoi Hook. f. Cushion plant formation.
L. pumila Hook. f. Cushion plant formation.
L. campestris D.C. var. Petriana Buchen. Scrub, tussock, river-bed, Senecio meadow.
L. campestris D.C. var. picta (A. Rich) Hook. f. Forest.
L. leptophylla Buchen. River-bed.
Liliaceae.
Enargea parviflora (Kunth.) Skott. Forest (Halpins Creek) and near top of pass.
Astelia Cockaynei Cheesem. Forest, scrub.
A. Petriei Ckne. Fellfield, tussock.
A. linearis Hook. f. Bog.
Phormium Colensoi Hook. f. Scrub.
Orchidaceae.
Prasophyllum Colensoi Hook. f. Bog, forest.
Pterostylis areolata Petrie. Forest.
P. Oliveri Petrie. Forest.
Lyperanthus antarcticus Hook. f. Bog.
Caladenia bifolia Hook. f. C. Lyallii Hook. f. Bog, forest.
Corysanthes rotundifolia Hook. f. Forest.
C. triloba Hook. f. Forest.
Thelymitra uniflora Hook. f. Scrub (Blimit Peak)
Gastrodia Cunninghamii Hook. f. Forest.
Chiloglottis cornuta Hook. f. Damp rock in forest.
Fagaceae.
Nothofagus cliffortioides (Hook. f.) Oerst. Forest.
Santalaceae.
Exocarpus Bidwillii Hook. f. Scrub in forest belt.
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Loranthaceae.
Elytranthe tetrapetala (Forst.) Engl. Forest, on Nothofagus cliffortioides and in one case on Dracophyllum longifolium.
E. flavida (Hook. f.) Engl. Forest, on N. cliffortioides.
Polygonaceae.
Rumex flexuosus Sol. Wet ground in river-bed, and bog on pass.
Muehlenbeckia axillaris Walp. River-bed, forest.
Portulacaceae.
Claytonia australasica Hook. f. Shingle slip, Senecio meadow, cushion plant formation.
Montia fontana Linn. Running water.
Caryophyllaceae.
Stellaria gracilenta Hook f. River-bed.
Stellaria parviflora. Banks et Sol. ex Hook. f. Forest.
Colobanthus crassifolius Hook. f. Senecio meadow.
C. acicularis Hook. f. Cushion plant formation.

Some of the specimens collected possess five sepals, others four sepals. Whether the number of sepals is variable in this species or the specimens with four sepals should be referred to C. monticola Petrie we leave for future investigation.

Hectorella caespitosa Hook. f. Fellfield.
Ranunculaceae.
Clematis australis Kirk. Scrub, here only a scrambling plant.
Ranunculus Lyallii Hook. f. Scrub, tussock, Senecio meadow.
R. Lyallii var. Traversii (Hook. f.) Cheeseman. (Upper end of Rough Creek).
R. sericophyllus Hook. f. Fellfield, Mt. Rolleston, 7000 ft. (Miss E. Washbourn).
R. hirtus Forst. Wet ground.
R. rivularis Forst. Swamp.
R. foliosus Kirk. Senecio meadow.
R. tenuicaulis Cheesem. Sub-alpine scrub.
Caltha obtusa Cheesem. Senecio meadow.
Cruciferae.
Cardamine heterophylla (Forst.) O. E. Schulz. Forest.
C. heterophylla var. uniflora Hook. f. Senecio meadow.
Droseraceae.
Drosera Arcturi Hook. Bog.
D. spathulata Lab. Bog.
D. stenopetala Hook. f. Bog.
Pittosporaceae.
Pittosporum tenuifolium Banks & Sol. Forest. (Halpins Creek).
P. divaricatum Ckne. Forest.
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Rosaceae.
Rubus australis Forst. Forest.
R. schmidelioides A. Cunn. var. coloratus Kirk. Forest.
Geum parviflorum Sm. Forest, Senecio meadow.
G. uniflorum Buch. Cushion plant formation.
Acaena microphylla Hook. f. var. inermis Kirk. Forest edge.
A. Sanguisorbae Vahl var. pilosa T. Kirk. Forest edge.
A. microphylla X sanguisorbae. Forest edge.
A. saccaticupula Bitter. Scrub, forest.
Potentilla anserina Linn. var. anserinoides (Raoul) Cheesem.
Leguminosae.
Carmichaelia uniflora Kirk. River bank by mouth of Graham's Creek, and mouth of Mingha.
Geraniaceae.
Geranium microphyllum Hook. f. River-bed.
G. sessiliflorum Cav. River-bed.
Oxalidaceae.
Oxalis lactea Hook. Waterfall.
Coriaraceae.
Coriaria sarmentosa Forst. River-bed.
C. angustissima Hook. f. River-bed.
C. sarmentosa X angustissima (C. lurida) River-bed.
Rhamnaceae.
Discaria toumatou Raoul. Scrub on river-bed.
Elaeocarpaceae.
Aristotelia serrata (Forst.) W. R. Oliv. Forest.
A. fruticosa Hook. f. Forest, scrub.
A. serrata X fruticosa (A. Colensoi) Forest.
Malvaceae.
Hoheria glabrata Sprague & Summerhayes. Forest, scrub.
Violaceae.
Hymenanthera alpine Kirk. Scrub.
Viola Cunninghamii Hook. f. Scrub.
V. filicaulis Hook. f. Forest.
Thymeleaceae.
Pimelea prostrata (Forst.) Willd. Scrub, river-bed.
Drapetes Dieffenbachii Hook. Scrub, tussock,
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Myrtaceae.
Myrtus pedunculatus Hook. Forest on ridge.
Metrosideros umbellata Cav. Scrub in forest belt.
Leptospermum scoparium Forst. Scrub in forest belt.
Ongagraceae.
Fuchsia excorticata Linn. f. Forest.
Epilobium erectum Petrie. Forest.
E. pubens A. Rich. Forest.
E. alsinoides A. Cunn. Forest, tussock.
E. pedunculare A. Cunn. Tussock; rocky ground, 5000 ft.
E. gracilipes Kirk. River-bed.
E. crassum Hook. f. Shingle slips.
E. pycnostachyum Haussk. Shingle slips.
E. rubromarginatum Ckne. Fellfield.
E. insulare Haussk. Bog.
E. nummularifolium R. Cunn. Damp scrub and bog.
E. novae-zealandiae Haussk. (Specimens in Petrie Herbarium).
E. melanocaulon Hook. River-bed.
E. microphyllum A. Rich. River-bed.
E. glabellum Forst. River-bed.
E. macropus Hook. River-bed, Senecio meadow.
E. pictum Petrie. Scrub.
E. chloraefolium Haussk. Bog.
Haloragidaceae.
Haloragis depressa (A. Cunn.) Walp. River-bed.
Gunnera dentata Kirk. Forest.
G. prorepens Hook. f. Swamp (McGraths Creek)
Myriophyllum propinquum A. Cunn In water on river-bed.
Araliaceae.
Nothopanax Colensoi (Hook. f.) Seem. Forest, scrub.
N. simplex (Forst.) Seem. Forest.
Pseudopanax lineare (Hook. f.) Laing & W. R. Oliv. n. Comb. Forest.
(Panax Lineare Hook. f. Fl. Nov. Zel. 1, 93, 1853).
We consider this species on account of its life History, best placed in the genus Pseudoptmax.
P. crassifolium (Sol.) C. Koch. var. unifoliolatum T. Kirk. Forest.
Umbelliferae.
Hydrocotyle novae zealandiae DC. Fellfield.
Hydrocotyle sp. Forest.

We are unable to refer this form which is fairly abundant to any described species and refrain from describing it as new, until the whole of the New Zealand species are revised.

– 727 –
Schizelema pallidum (Kirk) Domin. Swamp.
S. Haastii (Hook) Domin. and var. cynopetalum Domin. Fellfield, forest, Senecio meadow.
Oreomyrrhis andicola Endl. Senecio meadow.
Aciphylla crenulata J. B. Armstr. Scrub, tussock.
A. similis Cheeseman. Tussock, forest.
A. colensoi Hook. f. var. maxima Kirk. Scrub, tussock.
Anisotome pilifera (Hook. f.) Ckne. & Laing. Fellfield, tussock.
A. Haastii (F. Muell.) Ckne. & Laing. Forest, scrub.
A. pilifera X Haastii.
A. aromatica Hook. f. Fellfield, tussock, scrub, forest.
A. imbricata (Hook. f.) Ckne. Cushion plant formation.
Angelica montana (Forst.) Ckne. Forest.
Cornaceae.
Griselinia littoralis Raoul. Forest.
Ericaceae.
Gaultheria rupestris (Forst) R. Br. Forest, scrub, tussock.
G. antipoda Forst. Forest, scrub.
G. depressa Hook. f. Cushion plant formation.
Pernettya nana Col. By side of creek, 2,500 ft.
Epacridaceae.
Pentachondra pumila (Forst) R. Br. Bog, scrub, tussock, forest.
Cyathodes pumila Hook. f. Bog.
C. empetrifolia Hook. f. Forest.
Leucopogon Fraseri A. Cunn. Scrub in forest zone, and river-bed.
Archeria Traversii Hook. f. Forest.
Dracophyllum Traversii Hook. f. Forest, scrub.
D. longifolium (Forst) R. Br. Forest, scrub.
D. rosmarinifolium (Forst) R. Br. (D. uniflorum Hook. f.) Scrub, bog, forest.
D. longifolium X rosmarinifolium. (D. acicularifolium (Cheesem.) Ckn.) Scrub, tussock.
D. Kirkii Berggr. Scrub.
D. pronum W. R. Oliv. Fellfield, scrub, tussock.
D. Kirkii X pronum (D. saxicolum) Scrub, tussock.
Myrsinaceae.
Suttonia divaricata (A. Cunn.) Hook. f. Forest, scrub.
S. nummularia Hook. f. Forest, bog, scrub.
– 728 –
Gentianaceae.
Gentiana patula (Kirk) Cheesem. Senecio meadow, scrub.
G. bellidifolia Hook f. Bog, tussock, Senecio meadow, fellfield; Mt. Rolleston, 7,000 ft. (A. Anderson).
G. bellidifolia Hook. f. Near top of pass by wayside.
Boraginaceae.
Myosotis pygmaea Col. Senecio meadow.
M. Forsteri Lehm. Damp forest and scrub.
M. explanta Cheesem. Damp sub-alpine rocks.
Scophulariaceae.
Mazus radicans (Hook. f.) Cheesem. Forest.
Hebe salicifolia (Forst.) Pennell. Forest edge.
H. subalpina (Ckne) Ckne & Allan Scrub, forest.
H. vernicosa (Hook. f.) Ckne & Allan Forest, tussock, scrub.
H. ciliolata (Hook. f.) Ckne & Allan Tussock.
H. lycopodioides (Hook. f.) Ckne Allan Tussock.
H. Haastii (Hook. f.) Ckne & Allan var. macrocalyx (J. B. Armstr.) Fellfield; Mt. Rolleston, 7,000 ft. (Miss E. Washbourn).
H. epacridea (Hook. f.) Ckne & Allan. Forest by creek.
H. macrantha (Hook. f.) Ckne & Allan. Scrub, tussock.
H. buxifolia (Benth) Ckne & Allan. Tussock, scrub, bog.
H. buxifolia var. pauciramosa Ckn. & Allan. Senecio meadow.
Pygmaea pulvinaris Hook. f. Fellfield.
Veronica Lyallii Hook. f. River-bed.
V. Bidwillii Hook. River-bed.
V. linifolia Hook. f. River-bed on wet rocks.
Ourisia macrocarpa Hook. f. var. calycina (Col) Ckne. Scrub.
O. Crosbyi Ckne. Forest.
O. caespitosa Hook. f. Scrub, creek bank in forest.
O. sessilifolia Hook. f. Tussock; Mt. Rolleston, 7,000 ft. (A. Anderson).
Euphrasia revoluta Hook. f. Tussock.
E. Laingii Petrie. River-bed in forest.
E. zealandica Wettst. Bog.
E. Cockayniana Petrie. Bog.
Plantaginaceae.
Plantago Brounii Rapin. Senecio meadow.
P. lanigera Hook. f. Scrub.
P. triandra Berrgr. Wet roadside and swamp.
– 729 –
Lentibulariaceae.
Utricularia monanthos Hook. f. Bog.
Rubiaceae.
Coprosma serrulata Hook. f. Forest, scrub.
C. lucida Forst. Forest (Halpins Creek).
C. rugosa Cheesem. Forest, scrub.
C. brunnea (Kirk) Ckne. River-bed.
C. ramulosa Petrie. Forest.
C. linariifolia Hook. f. Forest.
C. pseudocuneata W. R. Oliv. ined (= C. cuneata Auct. not Hook. f.) Forest, scrub.
C. crenulata W. R. Oliv. (= C. retusa Petrie. See Trans. N.Z. Inst., Vol. 49, p. 153, 1917).
C. parviflora Hook. f. Forest, scrub.
C. propinqua A. Cunn. Forest.
C. ciliata Hook. f. Forest, scrub.
C. foetidissima Forst. Forest.
C. depressa Col. Scrub.
C. repens Hook. f. Scrub.
Nertera setulosa Hook. f. Rocks in forest.
Asperula perpusilla Hook. f. River-bed.
Galium umbrosum Sol. Wet ground in river-bed.
Campanulaceae.
Pratia angulata (Forst) Hook. f. Creek-bed in forest.
P. macrodon Hook. f. Fellfield, tussock.
Wahlenbergia albomarginata Hook. Tussock, river-bed.
Stylidiaceae.
Donatia novae zealandiae Hook. f. Bog.
Phyllachne Colensoi (Hook. f.) Berggr. Cushion plant formation.
Forstera sedifolia Linn. f. Bog and rock.
F. tenella Hook. f. Forest.
Compositae.
Lagenophora petiolata Hook, f. Forest, tussock.
Brachycome Sinclairii Hook. f. Tussock, scrub, Senecio meadow.
Olearia Colensoi Hook. f. Scrub.
O. arborescens (Forst) Ckne & Laing. Forest.
O. ilicifoia Hook. f. Scrub.
O. arborescens X ilicifolia (O. macrodonta) Scrub.
O. avicenniaefolia (Raoul) Hook. f. Forest.
O. lacunosa Hook. f. Forest.
O. lacunosa X ilicifolia. Scrub.
O. Haastii Hook. f. Forest by river.
O. nummularifolia (Hook. f.) Forest, scrub.
Celmisia Walkeri Kirk. Tussock.
– 730 –
C. incana Hook. f. Tussock, scrub.
C. novae-zealandiae (Buch) Cheesem. Forest, bog, tussock, scrub.
C. discolor Hook. f. Forest, tussock, bog.
C. coriacea Hook. f. Scrub, tussock.
C. Armstrongii Petrie. Scrub, tussock, bog.
C. spectabilis Hook. f. Scrub, tussock.
O. petiolata Hook. f. Scrub, tussock, Senecio meadow.
C. Haastii Hook. f. Scrub, tussock, Senecio meadow.
C. gracilenta Hook. f. Swamp, forest.
C. alpina (Kirk) Cheesem. Bog.
C. gracilenta X spectabilis. Forest.
C. glandulosa Hook. f. Bog.
C. bellidioides Hook. f. By creek in forest.
C. sessiliflora Hook. f. Tussock.
C. laricifolia Hook. f. Fellfield, tussock.
C. linearis J. B. Armstr. Bog.
Haastia Sinclairii Hook. f. Shingle slips.
Gnaphalium luteo-album Linn. By creek in forest.
Raoulia subulata Hook. f. Fellfield.
R. australis Raoul. River-bed.
R. Haastii Hook. f. River-bed.
R. tenuicaulis Hook. f. River-bed.
R. glabra Hook. f. River-bed.
R. grandiflora Hook. f. Fellfield, tussock.
Helichrysum bellidioides (Forst) Willd Senecio meadow, river-bed.
H. filicaule Hook. f. By side of creek and dry forest.
H. depressum (Hook. f.) Benth & Hook. f. River-bed.
H. microphyllum (Hook. f.) Benth & Hook. f. Wet cliffs in forest.
Cassinia Vauvilliersii (H. & J.) & Hook. f. Forest, scrub.
Craspedia uniflora Forst. River-bed, scrub, Senecio meadow.
Cotula pyrethrifolia Hook. f. Forest, tussock, fellfield.
C. squalida Hook. f. Forest.
Leucogenes grandiceps (Hook. f.) Beauv. Fellfield, tussock, shingle slips.
Erechtites glabrescens Kirk. Forest.
Abrotanella linearis Berggr. Fellfield.
Senecio lagopus Raoul. River-bed.
S. bellidioides Hook. f. Forest, bog, scrub.
S. Lyallii Hook. f. Senecio meadow, tussock.
S. scorzonerioides Hook. f. Tussock.
S. Lyallii X scorzonerioides. Tussock, Senecio meadow.
S. elaeagnifolius Hook. f. Forest, scrub.
S. Bidwillii Hook. f. Scrub.
S. elaeagnifolius X Bidwillii. Forest.
Microseris scapigera (Forst) Hoffm. Swamp, wet rocks.
Taraxacum magellanicum Comm. Fellfield.