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Volume 69, 1940
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Key TO Species OF Odostomia B.

Shell 3.5 mm. in height or over.
 Whorls flat or nearly so, overhanging suture, angulated; plait small but distinct.
  Periphery distinctly angled; outlines straight; whorls quite flat castlecliffensia.
  Periphery sub-angled; outline rather pupoid towards summit; whorls very lightly convex; peripheral thread present. haurakiensis
 Whorls convex, not flat; plait not distinct.
  Whorls strongly convex, roundly shouldered, spire gradate murdochi
  Whorls not strongly convex, little or not shouldered; spire not gradate.
   Height of aperture less than one-third that of shell.
    Whorls moderately convex, little shouldered; no spiral sculpture whatever; protoconch considerably exsert. vestalis
    Height of aperture greater than one-third that of shell.
    Whorls lightly convex; not shouldered.
     Microscopic striae present; one incised spiral line below suture pedica
     Microscopic striae absent; two incised spiral lines bordering each side of suture. aucklandica
Shell less than 3.5 mm. high.
 Microscopic spiral striae present.
  Incised spiral sculpture absent.
   Columella with a strong twisted plait; whorls very flatly convex; suture not much impressed; periphery sub-angled. gorensis
   Columella-fold weak or almost obsolete.
    Suture moderately impressed.
    Height of aperture nearly one-third that of shell, which is well elevated.
     Body-whorl high; no angulation; aperture narrowly ovate. cryptodon
     Body-whorl wide, sub-angled at periphery; aperture broadly and roundly ovate.
      Peripheral cord present. acutangula
      Peripheral cord absent. parvacutangular
     Height of aperture nearly half that of shell, which is distinctly squat. taumakiensis
    Suture strongly impressed; whorls flatly rounded; body-whorl angulated at periphery, with a weaker angulation just below. biangulata
   Incised spiral sculpture present.
    Two incised spiral lines bordering each side of suture; shell not stout; whorls flatly convex; protoconch not depressed but erect. manukauensis.
  Microscopic spiral striae absent.
   Incised spiral sculpture absent; shell very small with a strong twisted plait and heavy embryo. graviapicalis
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Odostomia castlecliffensis n.sp. (Fig. 21.)

Shell of moderate size, high conic; height of spire 2 ½ times that of aperture, outlines straight. Post-nuclear whorls 5 ¾ in number, flat, the periphery, which is low down near suture, distinctly angled and bearing a low spiral thread; suture strongly incised. Protoconch heterostrophic, planispiral; nucleus almost wholly immersed. Spiral sculpture confined to the weak thread on periphery. Body-whorl in height a little less than half that of entire shell, flat above, strongly angled on periphery, convex on base; aperture rhomboidal; columella set vertically, arcuate, its plait very weak and inconspicuous and situated at insertion. There is the very slightest indication of a narrow, almost linear umbilical chink; inner lip narrowly callused; outer lip straight, but angulated at periphery of whorl.

Height, 4.1 mm.; width, 2.0 mm. (holotype).

Locality: Castlecliff, Wanganui (Castlecliffian).

Type in Auckland Museum (ex writer's collection).

This species simulates O. obstinata, common in Awamoan beds at North Otago localities, but the divergent apical characters provide a ready means of separation.

Odostomia haurakiensis n.sp. (Fig. 24.)

Shell small, narrowly umbilicate, high conic; height of spire 2 ½ times that of aperture, outlines pupoid. Post-nuclear whorls 5 ½ in number, convex, overhanging; the low periphery distinctly angled; suture deeply incised. Protoconch heterostrophic, planispiral, low; nucleus almost completely immersed. Spiral sculpture confined to a fine thread on angulated periphery. Body-whorl in height a little less than half that of shell, flatly convex above and on base, sub-angled at periphery; aperture broken, but probably originally rhomboidal; columella set vertically, arcuate, its plait moderately conspicuous and situated almost at insertion; inner lip narrowly callused; outer lip broken.

Height, 3.9 mm.; width, 1.7 mm. (holotype).

Locality: Hauraki Gulf, in 28 fathoms.

Type(unique) in Auckland Museum (ex Finlay collection).

The present species is a very close relative of O. castlecliffensis, and they both possess the same general style of shell. A ready means of separation is to be had, however, in the rather pupoid outline, heavier plait, more open umbilicus and more convex whorl of haurakiensis.

Odostomia murdochi Suter. (Fig. 32.)

1904. Odostomia proxima Murdoch, Trans. N.Z. Inst., vol. 37, p. 226, pl. 8, fig. 19; not of de Folin, 1872.

1913. Odostomia murdochi Suter, Man. N.Z. Moll., p. 340, pl. fig. fig. 24 (Atlas).

The type material of this species consists of four shells, no particular one of which was designated as holotype by either Murdoch or Suter. None of the specimens agrees exactly with the dimensions given by Murdoch and copied by Suter, so that the writer now selects as lectotype the shell which most nearly approaches the

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recorded dimensions. Suter's figure in the Atlas is a reproduction of Murdoch's drawing, and does not do justice to any one of the four shells. It makes the habit rather less stout than is typical, and does not portray the true shape of whorl. The outline of whorl is not evenly convex, as shown in Murdoch's illustration, but is flatly convex over centre and drawn in rather rapidly to sutures, especially to the posterior one, where there is almost a tabulation, so that the spire appears somewhat staged.

The protoconch is heterostrophic, apparently planispiral, the nucleus almost completely hidden in succeeding whorl.

Height, 4.0 mm.; width, 1.8 mm. (lectotype).

Localities: Whangaroa Harbour (type); Banks Peninsula, recorded by Iredale, fide Suter, Manual; but in the light of other distributions shown by the present paper this may be questioned.

Type in Dominion Museum, Wellington.

Odostomia vestalis Murdoch. (Fig. 33.)

1905. Odostomia vestalis Murdoch, Trans. N.Z. Inst., vol. 37, p. 227, pl. 8, fig. 20.

1913. Odostomia vestalis Murdoch. Suter, Man. N.Z. Moll., p. 342, pl. 17, fig. 4 (Atlas).

The figure in the Atlas shows the whorls too flat and not sufficiently shouldered. The protoconch is more exsert than is usual for Odostomia, and is large, rather globose and planispiral. The general build and features of body-whorl and aperture remind one of Gumina, but the spire is much too high and the apex different.

Height, 4.4 mm.; width, 1.5 mm. (holotype).

Locality: Whangaroa Harbour.

Type in Dominion Museum, Wellington.

Odostomia pedica n.sp. (Fig. 34.)

Shell small, high conic; height of spire about 1 ½ times that of aperture, outlines straight. Post-nuclear whorls 4 ½ in number, convex; suture impressed. Protoconch heterostrophic, planispiral; nucleus about one-half immersed. Fine microscopic striae present here and there on all whorls; one incised spiral line margins suture below. Body-whorl in height slightly over half that of shell, broadly convex in one sweep from suture to base; aperture subovate, angled behind, fairly widely rounded in front; columella set vertically, arcuate, its plait not visible externally, but seen as a low fold within aperture; inner lip narrowly callused; outer lip thin, straight.

Height, 3.8 mm.; width, 1.6 mm.

Locality: Snares Islands.

Type (unique) in Wanganui Museum.

This species has been separated from among Suter's paratypes of O. inornata, which has an apex of helicoid type, and from which it is also distinct in having the suture margined.

Odostomia aucklandica n.sp. (Fig. 28.)

Shell of moderate size, conic, moderately elevated; height of spire twice that of aperture, outlines straight. Post-nuclear whorls

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4 ½ in number, convex, closely shouldered above; suture well impressed. Protoconch heterostrophic, apparently planispiral, low; nucleus almost completely immersed. Pine microscopic spirals present; two fine incised spiral lines present bordering both anterior and posterior sutures; spiral sculpture seen only on good specimens. Body-whorl in height a little greater than half height of shell, its outline convex throughout in one broad sweep from suture to base; aperture broadly ovate, angled behind, effuse in front; columella about vertical, arcuate, its fold exceedingly weak, almost obsolete; inner lip distinctly narrowly callused; outer lip thin, straight.

Height, 4.1 mm.; width, 1.95 mm. (holotype).

Locality: Faith Harbour, Auckland Islands.

Type in Auckland Museum (ex Finlay collection).

O. aucklandica combines the characters of both O. cryptodon and O. murdochi. It resembles the former in possession of a weak plait and in apertural shape, and the latter it simulates very closely in general build of whorl and character of suture. The Faith Harbour shells, however, show some variability in the depth of incision of suture, and not all have the suture so deeply impressed and the shoulder so strong as murdochi. Cryptodon is a much smaller species-with suture less incised and the body-whorl higher, and it lacks, incised spiral lines.

Odostomia gorensis n.sp. (Fig. 26.)

Shell very small, conic; height of spire 1 ½ times that of aperture, its outlines straight. Post-nuclear whorls 3 ¾ in number, very lightly convex; suture not strongly impressed. Protoconch large for size of shell, rounded over summit, planorboid. Sculpture consists of excessively fine, rather wavy, microscopic spiral striae. Body-whorl in height equal to half that of shell, flatly convex above, sub-angled at periphery, convex on base; aperture rhomboidal, angled behind, moderately widely rounded in front; columella thick, a little oblique, arcuate, its plait conspicuous, blunt and situated some distance below insertion; inner lip not callused; outer lip straight.

Height, 2.2 mm.; width, 1.1 mm.

Locality: Chatton, near Gore, Southland (Ototaran).

Type (unique) in writer's collection.

Odostomia, cryptodon Suter. (Fig. 29.)

1908. Odostomia cryptodon Suter, Trans. N.Z. Inst., vol. 40, p. 365, pl. 29, fig 12.

1913. Odostomia. cryptodon Suter, Man. N.Z. Moll., p. 335, pl. 16, fig. 16 (Atlas).

Suter gives a very fair illustration of this species, but he shows the pillar insufficiently excavated and the body-whorl contracted to base too low down and too sharply. The planispiral apex is considerably immersed.

Height, 2.7 mm.; width, 1.5 mm. (holotype). Corresponding dimensions of shell from Dunedin Harbour: 4.0 mm.; 1.9 mm.

Localities: Te Onepoto Bay, near Lyttelton (type); Dunedin Harbour, under stones between tide marks; Moeraki; Taieri Beach; Chatham Islands (a worn shell); off Otago Heads, in 40 fathoms.

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Fig. 1. Odostomia bartrumi n.sp.; paratype, × 13. Fig. 2.—Odostomia inornata (Marwick); holotype, × 13. Fig. 3.—Odostomia pudica suter; holotype × 13. Fig. 4.—Odostomia wantakiensis n.sp.; holotype, × 13. Fig. 5—Odostomia takapunaensis suter; holotye, × 13. Fig. 6.—Odostomia ototarana n.sp.; holotype, × 13. Fig. 7.—Odostomia zecorpulenta n.sp.; holotype. × 13. Fig. 8.—Odostomia incidata Suter; holotype, × 13. Fig. 9.—Odostomia waipaoa (Marwick); holotype, × 13.

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Fig. 10.—Odostomia chordata Suter; holotype, × 13. Fig. 11.—Odostomia obstinate n.sp.; holotype. × 13. Fig. 12.—Odostomia arowhana (Marwick); holotype, × 13. Fig. 13—Odostomia acatnmida n.sp; holotype, × 13. Fig. 14.—Odostomia pervaga n.sp.; holotype. × 13. Fig. 15.—Odostomia mahoenuica n.sp; holotype, × 13. Fig. 16.—odostomia twneri n.sp.; holotype × 13. Fig. 17.—Odostomia sherriffi Hutton; holotype, × 13 Fig. 18.—Odostomia vaga n.sp.; holotyupe, × 13. Fig 19.—Odostomia ancisa (Marwick); holotype, × 13. Fig. 20.—Odostomia chattonesis n.sp.; holotype × 13.

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Fig. 21. Odostomia castlecliffensis n.sp.; holotype, × 13. Fig. 22.—Odostomia grariapicalis n.sp.; holotype, × 13. Fig. 23.—Odostomia biangulata n.sp.; holotype × 13. Fig. 24.—Odostomia haurakiensis n.sp; holotype, × 13. Fig. 25—Odostomia paracutangula n.sp; holotyope, × 13. Fig. 26.—Odostomia gorensis n.sp.; holotype × 13. Fig. 27.—Odostomia taumakiensis Suter; holotype × 13. Fig. 28.—Odostomia aucklandica n.sp.; holotype × 13. Fig. 29.—Odostomia cryptodon Suter; holotype × 13. Fig. 30—Odostomia manukauensis n.sp; holotype, × 13. Fig. 31.—Odostomia acutangula Suter; holotype, × 13 Fig. 32.—Odostomia murdochi Suter; holotype × 13 Fig. 33.—Odostomia vestalis Murdoch; holotype × 13 Fig. 34—Odostomia pedica n.sp.; holotype × 13.

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Type in Wanganui Museum.

The Dunedin Harbour shells provide a very perfect match of the type, which is an immature specimen, the surface of which is not well preserved, so that Suter was led to record the absence of spiral sculpture for this species. The shells from Dunedin Harbour, however, have irregularly spaced, wavy microscopic striae.

Odostomia acutangula Suter. (Fig. 31.)

1908. Odostomia acutangula Suter, Trans. N.Z. Inst., vol. 40, p. 368, pl. 29, fig. 13.

1913. Odostomia acutangula Suter, Man. N.Z. Moll., p. 334, pl. 16, fig. 14 (Atlas).

Acutangula is known from a single specimen. The periphery is angled and the angulation carries a low spiral cord. The figure in the Atlas gives a fair representation of the whorls of the spire, but it shows the body-whorl hardly enough distended, the angulation too sharp, the columella straight instead of arcuate, and the aperture too narrow and not broadly enough rounded in front.

The protoconch is heterostrophic and apparently planorboid, of about one volution, and well tilted. Most of the nucleus is submerged.

Height, 2.4 mm.; width, 1.2 mm.

Locality: Port Pegasus, Stewart Island, in 18 fathoms.

Type in Wanganui Museum.

Odostomia parvacutangula n.sp. (Fig. 25.)

Shell very small, conic, moderately elevated; height of spire 1 ½ times that of aperture, outlines straight. Post-nuclear whorls 3 ¾ in number, lightly and evenly convex; suture impressed. Protoconch heterostrophic, planispiral, large, bulbous, exsert; nucleus about two-thirds immersed. Spiral sculpture not definite, traces of fine microscopic striae here and there. Body-whorl in height equal to about half that of shell, lightly convex above, flattish on base, periphery roundly angulated; aperture broadly oval, angled behind, effuse in front; columella thin, set vertically, arcuate, plait practically absent and indicated only by a broad, low swelling at insertion; inner lip thinly and narrowly callused; outer lip thin, straight; basal lip very broadly convex.

Height, 2.0 mm.; width, 1.0 mm (holotype).

Locality: Auckland Islands, in 95 fathoms.

Type in Auckland Museum (ex Finlay collection).

These Auckland Island shells are in build, shell-form, and apical characters very close to acutangula, from Stewart Island, but they are smaller and have less convex whorls and almost obsolete pillar-plait.

Odostomia taumakiensis Suter. (Fig. 27.)

1908. Odostomia taumakiensis Suter, Trans. N.Z. Inst., vol. 40, p. 363, pl. 28, fig. 7.

1913. Odostomia taumakiensis Suter, Man. N.Z. Moll., p. 341, pl. 17, fig. 3 (Atlas).

Suter's figure in the Atlas shows the whorls too convex; they should be moderately and evenly rounded. His comparison with O. murdochi is faulty in part. The salient separative features he

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does not comment on, viz., larger size and stouter habit of murdochi, which also has sutures much more strongly cut in and the spire somewhat stepped. The type of murdochi is imperforate, so that his comparison in reference to this character is of little value.

The protoconch is considerably immersed.

Height, 3.0 mm.; width, 1.5 mm. (holotype).

Localities: Taumaki Island, west coast of South Island, in 10 fathoms (type); Petane (Nukumaruan).

Type in Wanganui Museum.

Odostomia biangulata n.sp. (Fig. 23.)

Shell very small, conic; height of spire a little over twice that of aperture, outlines straight. Post-nuclear whorls 3 ½ in number, lightly convex, shouldered above, cut in rapidly close to suture below; suture strongly incised. Protoconch heterostrophic, apparently planispiral, large, bulbous; nucleus considerably immersed (partly broken). Fine, microscopic striae over whole surface of last whorl and on base; a spiral angulation on periphery of body-whorl, and another, not so strong, on base not far below peripheral one. Body-whorl in height about equal to half that of shell, shouldered at summit, flatly convex above the sub-angled periphery, convex on base; aperture sub-ovate, angled behind, rather effuse in front; columella about vertical, arcuate, its plait small, high up and seen only within aperture; inner lip as a narrow callus; outer lip straight.

Height, 2.0 mm.; width, 0.9 mm. (holotype).

Locality: Shell-bed, Target Gully, Oamaru (Awamoan).

Type in Auckland Museum (ex writer's collection).

The biangulate character of the last whorl is a good means of separation.

Odostomia manukauensis n.sp. (Fig. 30.)

Shell small, conic; height of spire 1 ½ times that of aperture, outlines straight. Post-nuclear whorls 4 ¼ in number, increasing rather rapidly in size, very lightly convex; suture distinct. Protoconch heterostrophic, low, rounded, apparently planispiral; nucleus almost completely immersed. Ill-defined microscopic spirals over surface of later whorls; in addition there are two strongly incised spiral lines situated respectively close to anterior and posterior margins of whorl. Body-whorl capacious, lightly convex above and on base, somewhat sharply convex at periphery; aperture broadly subovate; columella set vertically, arcuate, a small, low swelling near its insertion; inner lip thinly and narrowly callused; outer lip effuse, thin, straight; basal lip drawn down and inwards towards axis of shell.

Height 2.3 mm.; width, 1.3 mm. (holotype).

Locality: Orua Bay, Manukau Harbour (specimens from the collection of the Rev. W. H. Webster).

Type and several paratypes in War Memorial Museum, Auckland.

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Odostomia graviapicalis n.sp. (Fig. 22.)

Shell very small, conic; height of spire nearly twice that of aperture, its outlines straight. Post-nuclear whorls 4 in number, lightly convex; suture moderately impressed. Protoconch large, rather bulbous, planispiral; nucleus large, not central, a little immersed. Spiral sculpture entirely absent. Body-whorl in height equal to half that of shell, flatly convex above, sub-angled at periphery, flattish on base; aperture narrowly rhomboidal, acutely angled behind, produced downwards and inwards in front; columella a little oblique, arcuate, its plait distinct and a little below insertion; inner lip narrowly callused; outer lip straight.

Height, 2 1 mm.; width, 0.95 mm. (holotype).

Locality: Clifden, Southland, band 6A (Hutchinsonian).

Type in Auckland Museum (ex writer's collection).

Easily distinguished from gorensis by its lack of spiral sculpture, less stumpy habit, rather differently shaped aperture, and less marked angulation of body-whorl.