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Volume 70, 1940-41
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Sub-family Tomocerinae Börner, 1906.

Tribe Neocerini nov.

It is necessary to erect this new tribe for the reception of the following genus containing two species which are related partly to the Lepidophorellini and partly to the Tomocerinae, but which, owing to their peculiar characteristics, do not fall comfortably into either of the foregoing groups.

Genus Neocerus nov.

Genotype: Neocerus spinosus sp. nov.

The following characters serve to distinguish this genus:—Scales are present, Abd. III longer than Abd. IV; dens lightly annulated and corrugated, two or three-segmented and spined. Mucro haired. Third antennal segment shorter than fourth. Eight ocelli to each side. Tenent hairs much reduced. For explanation of formula for dental spines see under Tomocerus, p. 346.

Key to the Species of Neocerus.
1. Creamy-white to orange-coloured species with two short, non-clavate tenent hairs to each foot. N. spinosus sp. nov.
2. Greyish-violet coloured species with one short, clavate tenent hair to each foot. N. insolitatus sp. nov.

Neocerus spinosus sp. nov. Plate 55, figs. 269–272; Plate 56, fig. 273.

Colour: In life, cream to orange mottled with black; mounted and with scales present, greyish-orange; denuded of scales, cream to orange. Irregular black markings on sides of body, particularly in anterior region. First three antennal segments orange to brownish, Ant. IV mauve. Legs and furcula pale cream to brown, tibio-tarsi mauve.

Clothing: Heavy clothing of large, longitudinally-ribbed and transversally-striated scales. Occasional plain setae mainly around posterior region and on legs.

Body: Length up to 5·2 mm. Antennae twice as long as head, the four segments related as 10: 16: 17: 23. Ocelli, eight to each side, all large and arranged in two rows of 5 and 3 respectively. No post-antennal organ. Abd. III longer than Abd. IV as 20: 12. Ventral tube long cylindrical, with terminal knob-like swelling.

Legs: Claw with one very long inner tooth one-quarter down from claw base. Two long exterior lateral basal teeth. Empodial appendage simple, lanceolate, and passing into a long wavy filament which reaches beyond tip of claw. Two short slender non-clavate tenent hairs to each foot.

Furcula: Scaled and with occasional plain setae. Manubrium to mucrodens as 37: 55. Dens two-segmented, segments related to mucro as 19: 27: 9. Mucro with many long setae and four teeth, a large, recurved apical tooth, two small ventral teeth, and one extremely large ventral tooth supplied with stout basal spine. From seven to ten prominent corrugations at distal end of dens. At distal end of first segment of dens two very large tridentate spines. Remaining dental spines simple and arranged in

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two series, one of equal spines and the other of unequal spines. Equal spines arranged according to formula 14/20–25, and unequal spines according to formula 9, 0–I, 0–1, 0–III/2, I, 1–2, I, 1, I, 1, I, 1, I, 1, III, 2. In addition, numerous scattered small spines.

Localities: Lake Brunner, in bush soil and debris in forest; South Westland, in forest near the Karangarua River, amongst bush debris and soil; Lake Kanieri, in leaf debris in kahikatea forest.

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Type: Slide 3/630 and Figured Paratype: Slide 3/631, Dominion. Museum Collection.

Neocerus insolitatus sp. nov. Plate 56, figs. 274–275.

Colour: In life and mounted, body, head, legs, and furcula strong greyish-violet.

Clothing: Of typical scales and occasional plain setae.

Body: Length up to 2·5 mm. Antennae twice as long as head, the four segments related as 7: 10: 11: 16. Mesothoracic terga completely covers prothorax and overlies slightly hind portion of head. Segmentation of body rather indistinct. Abd. III longer than Abd. IV as 19: 12. Ventral tube short and cylindrical. Ocelli, eight to each side, all large, arranged in two rows of five and three, respectively.

Legs: Claw with one large inner basal tooth, two exterior lateral basal teeth, and basally with up to eleven folds. Empodial appendage lanceolate and passing into a long wavy filament which reaches beyond end of claw. A single unusually short clavate tenent hair, about one-quarter length of claw, to each foot. A long prominent basal spine on each side of claw.

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Furcula: Manubrium to mucrodens as 27: 49. Dens three-segmented, last segment with five prominent corrugations. Dens faintly annulated and coarsely corrugated. Mucro similar to N. spinosus. Dental spines in two series, formula being 2, 11/8–12 and 0/12 with numerous irregularly-placed very small spines.

Locality: Waiho Gorge, amongst leaf debris in forest.

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Type: Slide 3/637, Dominion Museum Collection.

Tribe Tomocerini nov.

Owing to the erection of the new tribe Neocerini in the Tomocerinae it now becomes necessary to propose this new tribe Tomocerini to contain the old genus Tomocerus.

The Tomocerini have six ocelli to each side and Ant. III much longer than Ant. IV.

Genus Tomocerus Nicolet, 1841.

Ocelli, six on each side. Clavate tenent hairs well developed. Dentes spined, mucrons long and clothed with setae.

The occurrence and arrangement of the dental spines is of great importance in the diagnosis of species, and is expressed as a formula wherein large spines are shown as a number in bold type and small spines in lighter type. The dens usually is segmented and where the suture of a segment interrupts the series of spines an oblique line is shown. Variations in the numbers of spines are shown by hyphens.

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Two species occur in New Zealand, one of which is a well-known European form.

Key to the New Zealand Species of Tomocerus.
1. Claw with 5–7 equal inner teeth and single clavate, spined tenent hair. Dental spines 4–8/2–8, I, 1–2, I. T. minor Lubbock
2. Claw with one large and five smaller inner teeth, one clavate and spined tenent hair, one non-clavate tenent hair, and dental spines 2–3, I, 2–3, I, 1, I/o. T. setoserratus sp. nov.

Tomocerus minor Lubbock, 1862. Plate 56, figs. 281–282.

Colour: Lead coloured with scales, but without scales varying from yellow to almost black. Yellow forms often with lateral black mottling.

Clothing: Heavily clothed with typical ribbed and striated scales.

Body: Length to 4 mm. Antennae shorter than body, with basal segment yellow and the others purple.

Legs: Claw with 5–7 small inner teeth, and two lateral external basal teeth. Empodial appendage half as long as claw with single small tooth. A single clavate and spined tenent hair shorter than claw to each foot.

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Furcula: Dental spines tridentate. 4–8/2–8, I, 1–2, I. Mucro with apical, longer sub-apical, paired unequal basal teeth and from 5–10 small intermediate teeth.

Locality: Ross, South Westland, in and under old logs.

Tomocerus setoserratus sp. nov. Plate 56, figs. 276–280.

Colour: In life, grey; mounted and with scales, grey over a basal colour of yellow; denuded of scales, yellow with black mottling along sides of body. Legs and furcula dirty yellow, tibio-tarsi pale violet. Antennae, I yellow, II grey, III and IV bluish-grey.

Clothing: Heavily clothed with longitudinally-ribbed, transversely-striated scales and occasional plain setae, which are most prevalent around posterior region of abdomen and on legs and furcula.

Body: Length up to 3·5 mm. Antennae shorter than body, four-segmented, the segments related as 8: 12: 42: 19. Ant. III and IV annulated. Ocelli six to each side, all large. Tergum of mesothorax completely covers prothorax. Ventral tube short and swollen distally. Abd. III longer than Abd. IV as 11: 7.

Legs: Claw with six inner teeth, one of which is large and situated at basal end, and five others of medium size arranged equally spaced in line; two outer lateral basal teeth and six basal folds. Empodial appendage lanceolate, about half as long as claw, and with long, thin inner tooth at about one-third from base. A single strong clavate and spined tenent hair to each foot, and immediately above it an equally strong non-clavate tenent hair.

Furcula: Manubrium to dens to mucro as 23: 28: 6. Dens three-segmented, but sutures very ill-defined. Dental spines large and tridentate, the formula being 2–3, I, 2–3, I, 1. I/o. In addition, last segment of dens bears along its whole length two rows of strongly serrated setae down one side, exactly similar to serrated setae of Acanthomrus Wom. Mucro heavily clothed with setae, with an

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apical tooth, a very large sub-apical tooth, 5–7 smaller intermediate teeth, and large paired basal teeth. The numbers of intermediate teeth may vary between the two mucrones of the same individual.

Locality: Lake Brunner, in old logs.

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Type: Slide, 3/639, Dominion Museum Collection.

This species is somewhat similar to T. minor Lubb., from which it differs in the formula for the dental spines, the presence of serrated setae on the dens, in the dentition of the claw and empodial appendage, and in the arrangement of tenent hairs.