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Volume 72, 1942-43
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Scrophulariaceae.

Hebe menziesii (Benth.) Ckn. et Allan.

A much branched, erect, glabrous shrub to 2 m. or more high; older branches stout, erect, yellow-green, rough by old leaf scars and swellings, leafy toward the tips; young wood at the tips yellowish or yellow-green, conspicuously bifariously pubescent; pubescence

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widest above the leaves, narrowing upward; internodes with opposite and alternate shield-like inflations below the leaves, the inflations rounded and bossed at the base. Leaves openly spaced, wide spreading, 2–2·5 cm. long, 6 mm. broad, obovate-oblong, subacute, rounded at the base, shortly petiolate, coriaceous, green or yellow-green and concave above, lighter green below, convex, keeled; margins entire; petiole narrow, 2 mm. broad over a suddenly widened and thickened boss-like base. Racemes simple 2–6 (4) towards the ends of the branches, 3–4 cm. long, finely bifariously pubescent; bracts lanceolate, subacute, membranous or ciliolate at the margins, rounded or keeled at the back, about equalling the calyx. Flowers 1 cm. long, 1 cm. wide with the lobes spread, white; calyx 5 mm. long, 4 partite, light green, segments as the bracts, corolla tube exceeding the calyx about 2 mm., slender, 2 mm. diam., widest at the throat; limb 4-lobed, wide spreading; lobes lanceolate, rounded at the tip; stamens 2, exserted to half the length of the lobes; style longer than the lobes; ovary 2 mm. long, ovoid, slightly compressed. Capsule slightly exceeding the calyx, 6 mm. long, 3 mm. broad, obovoid, subacute, compressed, narrowed to the base.

H. menziesii was described from specimens of a plant discovered by Menzies at Dusky Sound, but it has since ben confused with Nelson and Marlborough plants not nearly related. To the species we refer our specimens from subalpine areas on the western side of McKinnon Pass and the summit of the Longwood Range, and Mr. W. A. Thomson has plants in his garden at Dunedin collected from the hills above Lake Alabaster and the Pyke River, in the watershed of the Hollyford River. Our description is drawn from specimens—now in the Herbarium, Plant Research Bureau, Wellington—collected in fruit at McKinnon Pass and in flower from plants in cultivation.

Hebe buxifolia (Benth.) Ckn. et Allan var. pauciramosa Ckn. et Allan × propinqua (Cheesem.) Ckn. et Allan.

A low, closely-branched, yellow-green, rounded shrub to about 5 m. high. Branches ± 4 cm. lng, 4 mm. diam. with the leaves on. Leaves 3 mm. long, 2 mm. broad, loosely imbricating, decussate, coriaceous, obovate, subacute, convex above, keeled beneath, sessile by a wide base, smooth and glabrous. Flowers in 2–4 short dense spikes at the ends of the branches and a terminal one, forming a rounded head; bracts 3 mm. × 2 mm. diam., similar to the leaves; calyx 4 mm. long, 2 mm. diam.; segments obovate, obtuse or subacute, unequal. Corolla 1 cm. long, white; tube equalling the calyx, 1 mm. diam.; limb 1 cm. diam. across the spreading lobes; lobes 4 mm. long, ovate to almost orbicular. Stamens 2, exceeding the tube. Capsule 3 mm. long, ovate, compressed.

This is a smaller and more compact plant than Petrie's Veronica cassinioides, which Cockayne and Allan (1934, p. 40) regard as H. buxifolia × lycopodiodes, and it was found it some quantity where the parent species intermingle on a flat ridge at 900 m. altitude on the Garvie Mountains.

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Hebe willcoxii (Petrie) Ckn. et Allan.

This species, which Cheeseman (1925, p. 810) considered “apparently rare in the wild state,” is a simple species characteristic of subalpine shrub associations and grass land at McKinnon Pass, at the Upper Cleddau Valley, and the mountains above the Homer and Eglinton Valleys. In lesser amount it occurs in the upper basins of the Routeburn River where the type specimens were collected, but it is again common in the upper basins of the Matukituki River at Hector's Col, Mount French, and other mountains near Mount Aspiring. Apparently its range in the Fiord Botanical District is wide. Wall (1927, p. 253) records it from Lewis Pass, where we did not find it, but where a glaucous plant, probably H. glaucophylla Ckn. is common on Mount Technical with H. traversii and a bewildering hybrid mixture. H. willcoxii is here separated from the earlier described H. cockayniana, which species we restrict to the North-Western Botanical District.

Hebe brockiei sp. nov.

Frutex parvus, H. willcoxii affinis; rami annotini fusci, hornotini virides, bifariam pubescentes. Folia conqesta, 1–3 cm. longa, 8 mm. lata, late obovata, subacuta, coriacea, nitentes, supra profunde concava, subtus obscure carinata. Racemi oppositi ex axillis foliorum superiorum orti; bracteae ovato-lanceolatae, acutae, 1 mm. longae, pedicelli breves, calyx 4-partitus, 2 mm. longus, tubum corollae excedens, segmentis acutis, corolla 5 mm. longa, tubus brevis, lobis late ovate-lanceolatis. Capsula late ovata, acuta, 4 mm. longa.

A small, much-branched, spreading, glabrous shrub 20–30 cm. high; older branches dark brown, rough by old leaf scars, leafy towards the tips; tips green, ringed with dark brown at the nodes, bifariously pubescent above the leaves. Leaves closely placed, spreading, 1–1·5 cm. long, 8 mm. broad, broadly obovate, obtuse, abruptly sub-apiculate, narrowed to a sessile base, coriaceous, light shining green and deeply concave above, lighter green, rounded and obscurely keeled beneath; margins yellowish, entire. Flowers in 2 opposite racemes near the tips of the branches; rhachis 2–4 cm. long, slender, purplish, pubescent, with few or many flowers crowded on the upper portion; pedicels short; bracts ovate lanceolate, acute, half the length of the calyx, membranous at the margins; calyx 4 partite, green, 2 mm. long, exceeding the corolla tube; segments ovate lanceolate, acute, with membraneous margins; corolla white, 5 mm. long, tube short; lobes broadly ovate lanceolate, obtuse, stamens½ the length of the lobes; style longer than the lobes. Capsule broadly ovate acute, 4 mm. long, compressed. Allied to H. willcoxii, but a much smaller plant.

Habitat: Grassland on hills between Amuri Pass and Lake Man, at the head of the Doubtful River, Canterbury, at 1200–1500 m. altitude.

Type in Herbarium, Plant Research Bureau, Wellington.

This small shrub is plentiful at the type habitat, where it was discovered by Mr. W. B. Brockie, the enthusiastic collector in charge of the Native Plant Section at the Botanic Gardens, Christchurch.

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Hebe ramosissima sp. nov.

Suffrutex, H. petriei affinis, sed multo parvior, confertim ramosus, ramulis adscendentibus, foliis tectis; folia basi ferc connata, recurvata, 5 mm. longa, 2 mm. lata, late obovata, obtusa, supra concava, subtus obscure carinata, carnosa; spicae ramulis 2–4, terminales, 12 floribus; bracteae 4 mm. longae, obtusae; flores 8 mm. longi; 6 mm. diam.; calyx profunde 4-partitus; corollae tubus 5 mm. longus; lobis ovatis, 3 mm. longis. Capsula ovata, 5 mm. longa.

A prostrate, closely-branched, glabrous, softly woody plant forming closely-matted patches 20–30 cm. diam. Stems 10–20 cm. long, ± 2 mm. diam., heavily marked by old leaf scars, somewhat tortuous, rooting sparingly. Branches ascending, closely uniformly leafy, 7–8 mm. diam. with the leaves on. Leaves closely quadrifariously imbricating, opposite pairs almost connate at the base, wide spreading and recurved, 5 mm. long, 2 mm. broad, broadly obovate, obtuse, narrowed to a sessile base, concave and dark green above, lighter green and obscurely keeled below, fleshy; upper margins dark red, obscurely and coarsely toothed; lower margins finely ciliolate. Flowering heads terminal, about 12 flowered, loose, about 2 cm. long, 1–5 cm. diam., formed of reduced 2–4 flowered spikes. Flowers white, very shortly pediceled, ± 8 mm. long with the lobes spread, 6 mm. diam. across the lobes; bracts 4 mm. long, linear, obtuse, concave above, rounded below, red tipped; calyx 4 partite to the base, 4 mm. long; segments as the bracts; corolla tube longer, 5 mm. long, 2 mm. diam.; lobes 4, ovate, rounded or subacute, 3 mm. long, 1·5–2·5 mm. broad; stamens 2; slightly exceeding the tube; style shorter than or equalling the stamens. Ovary linear, 2 mm. long. Capsule ovate, obtuse, 5 mm. long, 2·5 mm. wide, laterally compressed, didymous.

Habitat: Moist debris on Mount Tapuaenuku, Inner Clarence Basin, Marlborough, 2150 m. altitude.

This small alpine species has some affinities with H. haastii and H. epacridea, but it appears to be still more closely related to H. petriei, which is, however, a very much larger spreading species.

Type specimens in the Herbarium, Plant Research Bureau, Wellington.

Veronica dasyphylla T. Kirk.

As stated by Cockayne and Allan (1927, p. 42) in their discussion on Hebe dasyphylla, “This is a distinct but compound species,” and in various forms it has a considerable range. Plants of fellfield on the Garvie Mountains and of moist hollows on the Rock and Pillar Range are very similar to those represented by specimens in the Dominion Museum Herbarium collected by Buchanan from Black Peak and Mount Alta and by Speden from the Old Man Range, and minor differences, due to habitat conditions, are to be expected in specimens from widely different localities. The flowers are numerous, solitary and sessile in the axils of the upper leaves, not terminal as in all the descriptions, and they almost conceal the branches when in full bloom. The corolla is large 1–1·5 cm. diam. when spread, sometimes larger, diurnal. The varieties hereunder described are distinct forms, easily separated from those comparable in leaf size and shape with those of the type. Cheeseman erred in including it

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in the Hebe section in Veronica, and Cockayne and Allan (loc. cit.), then unaware of its peculiar dehiscence and following his placing, included it in the genus Hebe. In Pygmaea also the flowers are axillary and solitary near to the ultimate tips of the branchlets, not terminal.

—–var. minor nov.

Typo multo minor, ramulis gracilibus, sparse foliatis; folia 3 mm. longa, 2 mm. lata, ovata; corolla 1 cm. diam., tubus 3 mm. longus, lobis ovato-spathulatis.

Much smaller in all its parts as compared with the type. Branches laxly leafy, 4 mm. diam. with the leaves on, purplish, everywhere dotted with stiff, white, retrose hairs. Leaves small, 3 mm. long, 2 mm. broad, obovate, rounded at the tips, finely pubescent on both surfaces. Corolla 1 cm. diam. when spread; tube 3 mm. long; lobes of the limb ovate spathulate, rounded at the tips.

Habitat: Turfy hollows amongst low grasses and other vegetation on the summit of Mount St. Mary, Kurow, Waitaki Valley.

Type specimens in the Herbarium, Plant Research Bureau, Wellington.

The slender, open, and laxly leafy habit, the small, rounded leaves, and smaller flowers of this plant separate it from all other forms near to the type.

—–var. subacuta. nov.

Typo minor, ramulis dense foliatis; folia 3 mm. longa, 1 mm. lata, lineari-lanceolata; flores utin var. minore.

Smaller than the type. Branches closely leafy, 4 mm. diam. with the leaves on, purplish, covered with stiff, spreading, or retrose hairs. Leaves smaller, 3 mm. long, 1 mm. broad, linear lanceolate, subacute, slightly spreading, minutely pubescent on both surfaces. Flowers as in var. minor.

Habitat: Peaty ridges at Rough Peaks, Lake Wakatipu.

Type specimens in the Herbarium, Plant Research Bureau, Wellington.

Easily separated by its smaller size and by its small linear lanceolate leaves.

We are acquainted with this variety in other parts of the Lake Wakatipu District, and Wall's specimens in the Dominion Museum Herbarium, from Cecil Peak, belong here.

Veronica uniflora T. Kirk.

The small specimens in Buchanan's Herbarium at the Otago University Museum compare quite well with his description and drawing (1882, p. 347) of it as Logania armstrongii, and the calyx and ovary are pilose as figured. The type habitat, “Hector's Col, Mount Aspiring, 5000 ft. alt.” (Matukituki Saddle of the maps), and the slopes in its near vicinity have been visited by Petrie and others, and by the authors on different occasions, but the plant collected in 1881 has not been rediscovered. Much is still unknown of the vegetation on the steep faces rising to Mount Bevan and Mount Barff from the snow-filled ravine leading up to Hector's Col. and records of the occurrence of the plant at other stations must in the meantime be regarded as doubtful. Buchanan's specimens may

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be those of an epharmonic form, but they do not match, even nearly, any specimens or plants of V. dasyphylla we have seen. Young leaves have a few stiff pointed hairs standing up from their tips, the flowers are comparatively large, solitary, and sessile, and they appear to be axillary as in V. dasyphylla and other small species.

Veronica muelleri Buch.

A slender, prostrate, closely branched, softly woody plant, spreading and rooting to form dense patches 20 cm. or more in diam. Branches short, 2–4 cm. long, leafy tetragonous, 1·5 cm. diam. with the leaves on, ascending; stems slender, purplish, glabrous or with a few thick hairs. Leaves spreading, ·5–1 cm. long, 2–3 mm. broad, oblong or ovate or obovate spathulate, rounded or obtuse at the tip, narrowed to the petiole, almost or quite flat, entire or with 1–3 teeth on each side, glabrous, shining green on both surfaces; leaf tips and teeth with short stout apicals; petioles about½ the length of the blade, almost connate at the base. Flowers axillary, solitary or branched in pairs; pedicels short, about 1 mm., elongating to about 5 mm. in fruit, glabrous, purplish; bracts at the base of the pedicels, shorter and narrower than the leaves, entire; calyx 4–5 mm. long, divided to the base into 4 equal and open segments, or with 4 equal segments and a short narrow linear segment; equal segments obovate-spathulate, glabrous; corolla pink, 6–7 mm. long, exceeding the calyx 2 mm. or more, ephemeral, 4 lobed, slightly spreading, 4 mm. diam., patent to 8 mm. diam. when deciduous; lobes 2 mm. broad, broadly lanceolate obtuse, the anterior lobe orbicularspathulate and emarginate, 4 mm. broad and frequently split into 2 narrow segments; tube shorter than the calyx segments, 3 mm. long, 2 mm. diam., whitish; stamens 2, exceeding the tube,½ the length of the lobes; anthers large, pink, 2 lobed; style slender, equalling the stamens. Ovary 1 mm. diam., conical, slightly compressed, seated on a ringed disk. Capsule obcordate included in the calyx, 4 mm. long and broad, laterally much compressed, grooved at the septum, splitting simultaneously both loculicidally and septumically into 4 segments in dehiscence.

Habitat: Consolidated moraine, fans and debris slopes in upper basins of the western branch of the Matukituki River at Hector's Col. Altitude 700–1400 m. Specimens have been deposited in the Herbarium, Plant Research Bureau, Wellington. This species is described as a straggling plant with a terminal inflorescence, and the plate shows an elongated corolla tube, 4 lobed wide-spreading limb and a carpel with a didymous dehiscence, but the straggling habit is not apparent, and the stems and branches intermingle closely. The flowers, apparently terminal, are axillary, the corolla tube is distinctly shorter than the calyx, and the lobes spread widely only when deciduous. Buchanan no doubt figured a corolla raised from the receptacle, and assumed the method of dehiscence.

Veronica plano-petiolata sp. nov.

V. muelleri affinis, sed differt ramulis rubicundis pilosis, foliis atris vel rubicundis, petiolis planis, floribus albis, calyce breve piloso.

A slender, prostrate, closely branched softly woody plant, spreading and rooting to form dense patches 20 cm. or more in diam.;

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branches short 2–4 cm. long, leafy tetragonous, about 1 cm. diam. with the leaves on; stems ascending, slender, purple, pubescent. Leaves spreading, ± 6 mm. long, 2 mm. broad, ovate-spathulate, subacute, narrowed to the petiole, almost or quite flat, entire or with a stout tooth on each side, somewhat fleshy, glabrous and shining, dark green changing to purple, petiole equalling or shorter than the blade, flat, 1 mm. broad, widening at the almost connate base, reddish.

Flowers axillary, solitary or 2 together; pedicels short, 1 mm. long, slightly elongating in fruit, sparingly pilose, purplish; bracts unequal, the larger one 4 mm. long, oblong-spathulate, entire, green, ciliolate on the margins, the smaller one 1 mm. long, linear, obtuse; calyx 4 mm. long, 4 partite to the base, openly divided; segments linear spathulate, obtuse, outer surfaces pilose; margins ciliolate; tube 2 mm. diam., equalling the calyx; limb white, 4 partite, 8 mm. long, 5–6 mm. diam. across the spreading lobes; lobes almost orbicular, the anterior one emarginate, sometimes split at the centre. Stamens 2, very slender, equally the tube; style slender, equalling the stamens. Ovary minute, globose, seated on a ringed disk. Capsule not seen.

Habitat: Debris slopes and fans on Mount McPherson, head of the Homer Valley, at 1400–1700 m. altitude.

Type specimens, collected by Mr. Owen Fletcher, in the Herbarium, Plant Research Bureau, Wellington.

This plant differs from V. muelleri by its reddish, sparsely pilose stems, its dark green or reddish leaves, widish flattened petioles, white flowers, and short, pilose calyx. The large lobe frequently splits at the centre in a manner similar to that of V. muelleri.

Veronica laxa sp. nov.

Suffrutex parvissimus, graclis parce ramosus, prostratus; ramuli 3–4 cm. longi, tenues, nonnihil purpurei, sparse pubescentes, pilis subpatentibus; folia 6–10 mm. longa, 4–6 mm. lata, late ovata vel fere orbiculata, nonnumquam obovata, 1–3 dentata vel integra; flores in axillis ramulorum, solitarii vel 2 conjunctim; bracteae 6 mm. longae, 3 mm. latae, obtusae, calyx 6 mm. longus, profunde 4-partitus, pubescens, segmentis obtusis; corolla alba, 8 mm. longa, 8 mm. diam., tubus calyce brevior, 4 mm. longus, lobis orbiculatis. Capsula obcordata, 4 mm. longa.

A slender, prostrate, laxly-branched, semi-woody plant, spreading and rooting to form loose patches 20 cm. or more diam. Branches short 3–4 cm. long, slender, purplish, thinly pubescent, with short, spreading, upturned hairs. Leaves varying greatly in size and shape 6–10 mm. long, 4–6 mm. broad, broadly ovate to almost orbicular or sometimes obovate, irregularly 1–3 toothed or lobed, or occasionally obovate spathulate and entire, rounded at the tip or subacute, rounded or narrowed to the petiole, fleshy, most minutely pubescent; petiole about half the length of the blade, narrow, flat, thickened and almost connate at the base. Flowers in the axils of the branches, solitary, or 2 together; bracts 6 mm. long, 3 mm. broad, obovate spathulate, obtuse; petiole 2 mm. long, purplish, pubescent; calyx 6 mm. long, 4 partite to the base, pubescent as the leaves; segments obovate spathulate, obtuse, entire or with one shallow notch at each side; corolla white, 8 mm. long, 8 mm. diam. with the limb spread, tube

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Fig. 1–Senerio bennettii.

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Fig. 2–Senerio cockaynei.

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shorter than the calyx, 4 mm. long, 2 mm. diam.; lobes 4, orbicular, 2 mm. broad; stamens 2, equalling the calyx; style equalling the stamens. Capsule obcordate, 4 mm. long, 4 mm. broad, laterally compressed, included in the calyx, dividing into 4 valves.

Habitat: Fine debris amongst rocks of moraine on the floor of the Homer Valley basin, near the tunnel, with Myosotis lyallii, at 800 m. altitude.

This plant was discovered by Mr. Owen Fletcher in 1939, and Mr. Coombs' specimens, now in the Herbarium, Plant Research Bureau, Wellington, have fruits and a few late flowers.