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Volume 75, 1945-46
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Genus Propesineuda nov.
1935 Parasinella Carpenter.

The name Parasinella was first used for a genus of Collembola in 1934 by Bonet (Arch. Zool. Exp. gén., Paris, 76, p. 365) when he erected the sub-genus Parasinella for the species P. cavernarum

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(Packard). Bonet's use of the name takes priority over that of Carpenter, and I propose the new name Propesinella for Parasinella Carpenter (Bull. B.P. Bishop Mus. 113, p. 138) of which the Genotype is: Propesinella (Parasinella) adamsoni (Carpenter).

The genera Ptenura Templeton, 1842; and SEIRA Lubbock, 1869.

Considerable confusion appears to have arisen amongst workers on the Collembola regarding the true identities of these two genera. The genus Ptenura was erected in 1842 by Templeton for several species previously included in the genus Podura. Templeton included in Ptenura his species nitida, nigromaculata and albocincta together with P. grisea Fabr. The genus Seira (often mis-spelt Sira by later authors) was erected by Lubbock in 1869 for the species Degeeria domestica of Nicolet and a new species, S. buskii. Later Lubbock added a third species, S. nigromaculata, and it was this species that Börner confused with P. nigromaculata of Templeton when he stated that Seira was a synonym of Ptenura (Das System der Collembolen, Mitt. Naturf. Mus., Hamburg, 23, p. 157). The species P. nigromaculata Templeton and S. nigromaculata Lubbock are not the same thing. Börner could not have studied Templeton's description of P. nigromaculata very closely or he would not have made this statement. Börner's error herein was first pointed out by Axelson (Die Apterygotenfauna Finlands, 1907–12, p. 215), but this appears to have been overlooked by subsequent authors. From a study of his description and figures of the species P. nigromaculata Temp., it seems almost certain that Templeton had before him, when he made this description, a species of what is now known as Mydonius. This is further borne out by the fact that Lubbock, in his monograph on the Order (Ray Society, 1873), placed P. nigromaculata Temp, in the genus Degeeria as a synonym of D. annulata Fabr. This issue is, however, further complicated by the fact that Templeton did not designate as the type of the genus Ptenura any particular species of the four he named. Of the other three species named by Templeton, albocincta is also probably a Mydonius, grisea an Isotoma and nitida belongs to the genus Heteromurus (Templetonia of Lubbock). The only possible way of fixing a type for the genus Ptenura is to take the first species mentioned in order of page priority, which, in this case, happens to be the species P. nitida. Accordingly the genus Heteromurus Wankel becomes a synonym of Ptenura Templeton, 1842, and the Genotype of the genus Ptenura must be Ptenura nitida Temp.

Turning to the genus Seira Lubbock, this was read before the Linnean Society of London in 1869, but it did not appear in print until the Transactions of the Society for that year were published in 1871. Here, on page 277, in his opening remarks, Lubbock expressly states that Nicolet's species Degeeria domestica forms the type of his new genus Seira: to quote—-“In placing on record the existence of Nicolet's Degeeria domestica as a British species…. I cannot regard it as belonging to the genus Degeeria, but rather as constituting the type of a separate genus of much interest as …” On page 279 of the same work the genus Seira is described and includes Seira domestica Nicolet and Seira buskii n.sp. In the year 1893 Schött split tip the

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genus Seira forming Pseudosira for those species with a falciform mucro and scaled dens, which definition included the species S. domestica Nic. However, as Lubbock had expressly stated that the species S. domestica formed the type of his genus Seira the name Pseudosira must fall as a primary synonym of Seira. Schött should have proposed his new name for these forms like nigromaculata and buskii with a bidentate mucro and scaleless dens; but Schött did not do this and, therefore, the name Willowsia proposed by Shoebotham in 1917 [Ann. Mag. Nat. Hist. (8), 19, p. 431] must stand for these latter forms. The Genotype of the genus Willowsia is Willowsia (Seira) nigromaculata (Lubbock).

This synonymy of the genus Pseudosira was pointed out by J. W. Shoebotham when he erected the genus Wilowsia in 1917, but it has apparently been overlooked by later authors. Denis (Bull. Soc. Ent. France, 1923, p. 54) recognised the genus Willowsia and in the same paper stated that he considered Seira trouessarti Moniez as identical with Seira domestica (Nic.) and published a full description of Seira domestica taken from the type specimens of S. trouessarti. In this description he states that Ant. IV is annulated; but, as Lubbock expressly stated in his description of the genus Seira, of which domestica is the type, that the antennae were not annulated, the species S. trouessarti Moniez cannot be the same as S. domestica (Nic.), Lubb. S. trouessarti is properly a species of the genus Lepidocyrtinus and there seems no doubt that the species referred to as Lepidocyrtinus domesticus (Nicolet) by more recent workers is Lepidocyrtinus trouessarti (Moniez). In his Monograph (1873) Lubbock figures the species Seira domestica from specimens apparently captured in England by a Mr. McIntyre. The antennae in these figures are not annulated.

The following synonymy should, therefore, be recognised in connection with these genera:—

  • Genus Seira Lubbock 1869.

  • 1872 Sira Tullberg.

  • 1893 Pseudosira Schott.

  • Genus Ptenura Templeton 1842.

  • 1801 Heteromurus Wankel.

  • 1802 Templetonia Lubbock.

  • * 1942 Propemesira Salmon.

[Footnote] * In a recent letter to me F. Bonet suggested that my Propemesira duo-oculata might be the cosmopolitan P. nitida. Re-examination of my material shows this to be correct, the fifth antennal segment on my material being somewhat obscured through the antennae sinking into the top of the head, and my genus Propemesira becomes, therefore, a synonym of Ptenura.