
A Skeleton of Notornis hochstetteri Meyer from Waitati, Otago, And Notes on Distribution of the Bird in the South Island, N.Z.
Department of Zoology, University of Otago*
[Read before the Nelson Philosophical Society, October 15, 1950; received by the Editor, May 28, 1951]
Summary
An incomplete skeleton of Notornis hochstetteri found at Waitati and now in the Otago Museum is described and figured. A list is given of all localities in the South Island from which Notornis remains have been recorded, also a list of localities of hitherto unrecorded bones in the Dominion, Canterbury, Otago and Invercargill museums.
A hitherto undescribed, incomplete skeleton of Notorms hochstetteri Meyer in the Otago Museum has recently come to my notice. The only available information about its origin is that it was found in sand at Waitati and was presented to the Museum by T. H. Murdoch. The skull, lower jaw, pelvis, femur and two tibiae were registered in 1913 and the tarso-metatarsus in 1914; presumably the tarso-metatarsus was overlooked in 1913 and was found in 1914 on revisiting the locality.
The skull is well preserved, but lacks the quadrate, quadrato-jugal, jugal, pterygoids and prevomers; these easily displaced bones apparently became separated from it in the sand and were overlooked by the collector. The following are the measurements of the various structures:
| Skull: | |
| Total length from post. surface of occipital condyle to ant. tip of beak | 94.0mm. |
| Greatest breadth | 39.5 |
| Least width between temporal fossae | 16.5 |
| Tip of beak to its junction with the frontals | 63.5 |
| Least width between external nostrils | 9.5 |
| Greatest depth of beak from junction of frontals to post. angle of maxilla | 30.0 |
| Lower Jaw: | |
| Total length of right ramus | 87.0 |
| Greatest vertical height at coronoid process | 22.0 |
| Length of symphysis | 36.0 |
| Greatest width at post. articular expansion | 45.0 |
| Width of right post. articular expansion | 15.5 |
| Femur: | |
| Length | 101.5 |
| Breadth of proximal end along axis of neck | 20.5 |
| Breadth of distal end | 21.5 |
| Circumference middle of shaft | 25.1 |
| Tibia: | |
| Length | 156.5 |
| Breadth of proximal end | 28.0 |
| Breadth of distal end | 17.0 |
| Circumference of shaft | 22.0 |
[Footnote] * Now at Entomological Research Station, Cawthron Institute, Nelson.

| Tarso-metatarsus: | |
| Length | 93.0 |
| Breadth of proximal end (transverse) | 17.5 |
| Breadth of proximal end (antero-posterior) | 18.0 |
| Breadth of distal end | 19.5 |
| Breadth of shaft | 8.5 |
| Pelvis: | |
| Greatest length | 115.0 |
| Greatest width | 52.5 |
| Width of sacrum | 23.5 |
The bones show no sign of fire or other indication that they may be midden material, and it appears that the bird died a natural death where it was found. This then is another record of its distribution in the South Island.
Since N. hochstetteri Meyer is considered to be confined to the South Island, and to be distinct from N. mantelli Owen, which appears to have been confined to the North Island, it is considered worthwhile to append an up-to-date list of localities from which Notornis remains have been recorded in the South Island. This is done on the assumption that all Notornis remains in the South Island are referable to N. hochstetteri.
| Date found | Reference to literature | ||
| 1849 | Duck Cove, on Resolution Island, Dusky Sound | Mantell | 1850 |
| 1851 | Secretary Island, Thompson Sound | Buller | 1873 |
| 1879 | Mararoa Flat, Te Anau | Parker | 1881 |
| * 1881 | Cascade Creek (now Cascade Burn) | Park | 1888 |
| * 1881 | Matukituki River, South branch | Park | 1888 |
| 1884 | Patience Bay, Te Anau | Parker | 1885 |
| *1888 | Mt. Hodge, Dusky Sound | Park | 1888 |
| *1888 | Docherty's Creek, Dusky Sound | Park | 1888 |
| *1888 | Mt. Pender, Dusky Sound | Park | 1888 |
| *1888 | Cooper's Island, Dusky Sound | Park | 1888 |
| *1888 | South side of Dusky Sound opposite Cooper's Island | Park | 1888 |
| 1892 | Castle Rocks, Southland | Hamilton | 1892 |
| 1892 | Earnscleugh, Central Otago | Hamilton | 1892 |
| 1892 | Long Beach, near Dunedin | Hamilton | 1892 |
| 1898 | Middle Fiord, Te Anau | Benham | 1899 |
| 1948 | Takahe Valley, Te Anau | Falla | 1948 |
Dr. R. A. Falla has furnished me with a list of Notornis bones in the collection of the Dominion Museum, from which the following new records were obtained:
Part skeleton collected by Major Paton 1872 in a limestone cave, Aniseed Valley, Nelson. This skeleton is on loan from the Nelson Museum. Various leg bones labelled by A. Hamilton as from Ngapara, North Otago, no date, and Warrington, Otago, no date.
From Dr. R. S. Duff a list of the Canterbury Museum collection, including new records from Kapua, Waimate, a tarso-metatarsus labelled by F. W. Hutton 1895. Various leg bones, a mandible and premaxilla from Pyramid Valley, North Canterbury, collected by Scarlett, Eyles, Duff and Falla between 1940 and 1949.
Dr. H. D. Skinner, Otago Museum, informs me that the recently acquired J. Murray collection of bird bones from limestone caves at Forest Hill, Southland, includes several crania, mandibles, pre-maxillae and a tarso-metatarsus of Notornis.
[Footnote] * These records of Park's were challenged by Melland (Trans N. Z. Inst, 22, pp. 295–300), but from my own experience of the habits of Notornis and its unmistakable “booming” note it seems reasonably certain that bis records are correct.

Mr. David Teviotdale supplied me with a list of the Notornis material held in the Invercargill Museum, a skull from Wakapatu, 1938, premaxilla and lower mandible from Pahia, 1933, lower mandible, Tokonui Mouth, 1939, all collected by J. H. Sorensen. A skull from Back Beach, Greenhills, 1942, collected by Robert Gibb, pelvis and sternum from caves, Limehills, 1928, collected by Messrs. George jaquiery, Fowler and Woods, and a skull and upper mandible from Wakapatu, 1946, collected by D. Teviotdale.
Falla (1948) states “Sub-fossil remains from Te Aute and Martinborough in the North Island, and Waikari, Enfield, Castle Rocks and other South Island localities …” Waikari (Pyramid Valley) and Castle Rocks are referable to records mentioned above, whilst Enfield, Dr. Falla (in litt.) informs me, refers to information gleaned from H. O. Forbes' records.
Consideration of the twenty-nine localities recorded above shows that Notornis was distributed from one end of the island to the other. The frequency of the sub-fossil remains found in an area is not necessarily a true indication of the population density of the bird, but depends rather on the suitability of the area for preservation of remains, and the work done there by collectors. The preponderance of remains and the bird's present existence in the Southland-Otago area indicates, however, that it occurred there in greater numbers than elsewhere.
Acknowledgments
My thanks are due to the Directors of the Dominion, Canterbury and Invercargill Museums for the information acknowledged in the text, and especially to the Director of the Otago Museum, Dr. H. D. Skinner, for information supplied and permission to describe the Waitati specimen, and to Mr. R. Blick, Cawthron Institute, for photographs.
References
Benham, W. B., 1899. Notes on the Fourth Skin of Notornis. Trans. N.Z. Inst., vol. 31, pp. 146–150.
Buller, W. L., 1873. History of the Birds of New Zealand. London, 1st Edition.
Falla, R. A., 1948. The Re-discovery of the Takahe. N.Z. Sci. Rev., vol. 6, pp. 123–124.
Hamilton, A., 1892. On the Fissures and Caves at the Castle Rocks, Southland; with a Description of the Remains of the Existing and Extinct Birds found in them. Trans. N.Z. Inst., vol. 25, pp. 88–106.
Mantell, G. A., 1850. Notice of the Discovery by Mr. Walter Mantell in the Middle Island of New Zealand, of a Living Specimen of the Notornis, a Bird of the Rail Family, Allied to Brachypteryx, and hitherto unknown to Naturalists except in a Fossil State. Proc. Zool. Soc. Lond., part XVIII, pp. 209–212.
Park, James. The Takahe (Notornis mantelli) in Western Otago. Trans. N.Z. Inst, vol. 21, pp. 226–230.
Parker, T. J. On the Skeleton of Notornis mantelli. Trans. N.Z. Inst, vol. 14, pp. 245–258.
Parker, T. J. Notes on a Skeleton of Notornis, recently acquired by the Otago University Museum. Trans. N.Z. Inst, vol. 18, pp. 78–82.

