
Genus Echthromorpha Holmgren, 1868
Echthromorpha Holmgren, Eng. Res. Zool., 1868, I, p. 406.
Rhynchopimpla Kriechbaumer, Berlin Entom. Zeitschr., XXXIX, 1894, p. 50.
Polyhamma Kriechbaumer, Berlin Entom. Zeitschr., XXXIX, 1894, p. 304.
Allothorinia Ashmead, Proc. U.S. Nat. Mus., 1900, XXII, p. 55.
Genotype: Echthromorpha maculipennis Holmgren (by designation Ashmead, 1900).
This genus was established by Holmgren in 1868 for some Australian species. In 1900 Ashmead separated the New Zealand form of Fabricius' species Cryptus intricatoria from the genus Echthromorpha and erected a new genus Allothorinia with the genotype A. 12-guttata. Townes (1946) examined Ashmead's type of Allothorinia 12-guttata and considered it was merely an aberrant form of Echthromorpha intricatoria (Fabr.), which differed from the typical form in having a pair of prominent tubercles on the propodeum and in having the propodeum closely punctured. I therefore follow Townes in synonymising Ashmead's genus with Echthromorpha Holmgren.
This genus may be distinguished from allied genera by the following combination of characters:
Labium exposed, plate-like, in about the same plane as the clypeus; mandibles tapering to a narrow tip and with the lower tooth shorter than upper, the tip of the mandibles not or slightly turned; propodeum punctate, striate or mat and without carinae. Abdomen punctate or somewhat mat, not polished; nervulus strongly postfurcal. This genus most closely resembles the genus Lissopimpla, from which it is readily separated by the Key given on page 2.
The genus Echthromorpha has a wide distribution. It has been recorded from Australia, New Zealand, Tahiti, Africa, Europe (?), Asia, Manilla, Ascension, Honolulu, Hawaii, Java, Sumatra, St. Helena, New Hebrides, Banks Islands and the Solomon Islands.
The genotype was described from the Hawaiian Islands, where it is said to be a common species found on most of the Islands of the group. Only one species occurs in New Zealand, Echthromorpha intricatoria (Fabr.).
Echthromorpha intricatoria (Fabr.)
Cryptus intricatorius Fabricius, Syst. Piez., 1805, p. 77.
Pimpla intricatoria Erichson, Arch. f. Naturg., VIII, 1841, p. 254.

Pimpla intricatoria Brulle. Hist. nat. Insect. Hymen. IV, 1846, p. 90.
Allotheronia 12-guttata Ashmead, Proc. U.S. Nat. Mus., XXIII, 1900, p. 55.
Echthromorpha intricatoria (Fabr.) Krieger, Sitzber. naturf. Ges. Leipzig, 1897–98, 1899, p. 59. Dalla Torre, Cat. Hymen, III, Pt. I, 1901, p. 455. Schmiedeknecht, Gener. Insect., Fasc. 62, 1907, p. 28 Morley, Rcv. Ichn., Part II, 1913, p. 39. Hudson, Ent. Mon Mag., 62, 1926, p. 40. Gourlay, Ent. Mon. Mag., 62, 1926, p. 170; Dept. Sci. & Ind. Res. Bull. 22, 1930, p. 5. Townes, Ann. Entom. Soc. Amer., XXXIII, 1940, p. 288.
This species, originally described from Australia by Fabricius in 1805, is one of the common and most conspicuous Iehneumons in New Zealand. It was first described under the genus Cryptus; Erichson, Brullè and other authors placed it in Pimpla, where it remained until 1899, when Krieger referred it to Echthromorpha. In 1900, Ashmead separated the New Zealand form, on which he based a new genus, Allotheronia, with the genotype A 12-guttata. In 1940, Townes, after examining Ashmead's type of A 12-guttata, considered it only an aberrant form of Echthromorpha intricatoria Fabricius. Ashmead's name, apparently, has not been used in the New Zealand literature and this species has been correctly referred to in literature and in collections as E. intricatoria
Schmiedeknecht (1907, p. 28) records E. intricatoria from Australia and figures it on Plate I, fig. 3 (fig. 2 sic, p. 28). This crude figure is certainly not this species, as it does not even remotely resemble it. Morley (1913, p. 39) states that he does not know any species of the present genus which at all agrees with this crude figure of Schmiedeknecht.
In 1901 Dalla Torre records Echthromorpha intricatoria Fabricius from America, New Cambria, New Hebrides, and New Zealand; but there is no record of its occurrence in America, and I cannot find any published record previous to 1900 on which this author bases its occurrence in New Zealand It is reasonable to assume, as this is quite a conspicuous and large insect, that it would have been recorded by collectors long before 1900 if it had been present; Dalla Torre's record is presumably erroneous, the first record of its occurrence in New Zealand should be that of Ashmead (1900).
Female
Head. Vertex black, red-brown between posterior ocelli and eyes, shining, impunctate, sparsely clothed with long, erect, pale-coloured hairs, posterior slope nearly vertical. Ocellar region black, rugulose, with a few fine erect hairs; Ocelli moderate size, posterior situated about their diameter from eyes and nearly 2 diameters from each other, and less than a diameter from anterior ocellus, which is slightly smaller. Frontal excavation black, deep and transversely aciculated; Face red-brown, punctate and clothed with long, light-coloured hairs; face wider (16) than long (12); internal orbits sharply indented above base of antennae; Clypeus red-brown, minutely punctate, and sparsely clothed with light-coloured hairs; wider than long, suture deep, roundly concave viewed from the front. Malar space wide, nearly equal to the length of the mandibles; Mandibles small, brown, with the tips of the teeth black, the upper tooth longer than lower tooth and both finely pointed; Cheeks black, bordered around outer orbits with red-brown, sparsely punctate, shining, clothed with long, light coloured hairs. Eyes black, with irregular streaks and blotches of dark-grey.
Antennae. Scape brown, deeply excavated dorsally; flagellum entirely brown, darkening towards apex and thickly clothed with short adpressed golden pubescence; 1st segment longest and segments decreasing in length to 11th or

12th, after which they are sub-equal, except apical segment, which is slightly longer, the 5th segment is approximately ½ as long as the 1st segment.
Thorax. Pronotum black, rugulose, sometimes with a small yellow mark on dorsal border. Mcsonotum black, coarsely punctate, clothed with fine whitish pubescence, notauli only faintly indicated anteriorly; Tegulae dark-brown with a few shallow punctures, shining, sparsely pubescent. Scutellar foveae deep, with a series of transverse ridges, lateral carinae high with a yellow mark at base of mesonotum; scutellum black with a few shallow punctures, somewhat convex, sloping posteriorly, lateral carinae reaching to about ¼ of its length, surface clothed with a few long hairs. Post-scutcllum yellow, shining, impunctate, lateral excavations at the side of post-scutellum, deep and longitudinally and finely ridged. Mesopleurae black, with a small yellow spot on sub-alar tubercle, a small yellow spot varying in size at base of hind-wing, and a larger yellow spot at the posterior ventral angle of the mesopleurae, strongly punctate, especially at the sides, punctures widely separated in the centre; pleural suture straight and transversely ridged; sub-alar tubercle distinct Sternum black, thickly punctate. Metopleurae black, a circular yellow spot in centre, coarsely punctate, the whole sclerite slightly swollen. Propodeum black with posterior lateral tubercles yellow, anteriorly rugulosely punctate, posteriorly finely transversely aciculated; propodeum spiracles elongated, slightly raised, and placed about 3 times their length from anterior border of propodeum Coxae : anterior coxae red-brown, punctate and clothed with long, light-coloured hairs; middle and posterior coxae black, irregularly punctate On dorsal surface the punctures are widely spaced, with a few light-coloured hairs scattered over the surface.
Legs. Trochantus, anterior red-brown; middle with 1st joint black, 2nd joint red-brown; posterior with both joints block. Femur all red-brown, posterior pair swollen, with an obsolete ventral longitudinal groove; Tibiae red-brown, thickening towards apex; Tibial spurs thick, about ½ the length of 1st tarsal joint; Tarsus red-brown, 1st and 5th joints sub-equal, 2nd longer than 3rd, 4th joint very small; minute spines on ventral surface, especially numerous on the 5th joint. Tarsal claws as figured (Fig. 5).
Wings (Figs. 6, 7, 8, 9). Forewing, 1st abscissa of radial straight, 2nd abscissa curved; areolet (Fig 9) petiolate; 2nd recurrent bent in centre, with 2 bullae; cubital with 1 bulla; 3rd abscissa of discordal with a bulla at its junction with anal. Stigma and veins dark brown, a clouded dark-brown spot covering the apical ⅓ of radial cell and extending slightly beyond radius into third cubital cell. Hind-wings. Abscissula nearly as long as marginal (radiella), a bulla at its lower ½, recurrent vein oblique, broken at top, nervellus (Fig. 8) continuous with 1st abscissa of median, thus strongly rechvous. Veins brown.
Abdomen Black with a large yellow spot on each of the posterial lateral angles of each of the first tergites. 1st tergite sessile, widest at posterior border, which is 1.6 times as wide between spiracles. Distance between spiracles is a little less than the distance from the spiracles to posterior border of tergite. Spiracles situated on moderately prominent tubercles, a longitudinal dorsal fovea, and an obsolete transverse ridge along posterior border, which is produced medially. 2nd tergite is 1.73 as wide apically as long and 1.4 as wide at apex as at base, on each side there is an oblique lateral groove reaching to about the middle of the segment, along the grooves are coarse punctures; the tergite convex with a few large but shallow punctures scattered over the surface. The posterior border slightly raised; in front of each of the yellow spots there is an obsolete

transverse groove. 3rd tergite, sides parallel, as wide posteriorly as anteriorly, 1·75 as long as wide, the oblique lateral grooves very short, reaching only to the 1st quarter of the length of the segment and punctate. The whole abdomen is shining and very sparsely punctured. 8th tergite fringed posteriorly with thick golden hairs. Ovipositor: Sheaths black-brown, covered with golden pubescence.
Male
Face brownish-yellow to yellow, more thickly covered by a white pubescence. Malar space usually black. Pubescence on thorax somewhat denser than in female and the mesopleurae more evenly punctate, a small yellow spot is generally present on the anterior surface of middle coxae and the fore-coxae. Tegulae yellow, the posterior ¾ black. The small yellow mark on dorsal margin of pronotum is more clearly marked than in the females. In some specimens there is a small yellow spot in centre of the anterior margin of the scutellum. Posterior tarsi with last joint infuscated with black.
Abdomen. 1st tergite, the posterior edge is more markedly protruding centrally, and a shallow and wide longitudinal depression from the spiracles which nearly reaches the posterior border. The distance between the spiracles is 1·25 the distance from the spiracles to the posterior border of the tergite. The 7th abdominal tergite occasionally has a small posterior lateral yellow spot.

