Four New Curculionidae (Col.) from New Zealand
Communicated by Mr. D. Spiller.
[Read before the Auckland Institute, September 19, 1951; received by the Editor, May 29, 1952.]
A New genus Himatiodes and a new species H. setulifer, together with three other new species of Curculionidae are described.
In 1937 (Trans. Roy. Soc. N.Z., Vol. 67, p. 324) I dealt with the genera that had been referred by Broun to the Cylindrorrhminae; four of them were removed, leaving five genera in this sub-family. An examination of the typical genera of Cylindrorrhininae from South America has made it clear that they are by no means closely related to the New Zealand forms, especially as the principal genera, Cylindrorrhinus, Adioristus and the typical Listroderes, all present an unusual character—namely, the presence of moveable spurs at the end of the tibiae, normally two on the hind pair, one on the middle pair, and rarely one on the front pair. Elsewhere in the Curculionidae these spurs occur only in such primitive groups as the Belinae, Eurhynchinae and Ithycerinae.
It would thus seem preferable to restrict the sub-family Cylindrorrhininae to those genera which possess such spurs. These appear to be confined to South America, and mainly to the southern half of it.
The nearest relations of the New Zealand genera seem to be found in the South African genus Hipporrhinus Schönh., and it is therefore now proposed to include them in the Hipporrhminae.
Pparchus carinirostris, sp.n.
♀ Derm dull black, with sparse minute grey or brown scaling. Head with the eyes absolutely flat, but a little broader than in P. gourlayi Brookes (8:3·8). Rostrum somewhat shorter then the pronotum, with very shallow confluent punctation, the dorsal outline regularly curved and not angulate; the dorsal area flat, tricarinate, the median carina being broader and slightly higher towards the base than the narrow sinuous lateral carinae; scrobes not reaching the eyes. Antennae black. Prothorax as long as broad, parallel-sided in the basal fourth, rounded laterally in front, widest at two-thirds from the base, without any apical constriction; dorsum rugose, with scattered confluent granules, two large, shallow, rounded impressions on each side just behind the middle, another in the middle of the base, and a transverse one at one-third from apex; the front margin of the prosternum very shallowly sinuate. Elytra oblong-elliptical, shallowly sinuate jointly at the base, shortly dehiscent at the apex; the broad striae containing large, narrowly separated punctures without intervening granules, stria 5 not curving outwards basally, so that 6 reaches the base; the intervals scarcely wider than the striae, costate, with an irregular row of small, low granules, intervals 3 and 5 only slightly higher than 4 and 6, 7 much higher and with more numerous
granules, 5 ending at the top of the declivity, 3 and 7 meeting near the apex; the scales small and separate, but larger than those on P. gourlayi. Legs as in P. gourlayi, but the lateral setae on the tarsi shorter, stouter and straight.
Length, 12 mm.; breadth, 5.5 mm.
South Island. Mt. Peel, Nelson, 5,000 feet; 1 ♀, 15.ii. 1931 (E. S. Gourlay).
Apart from the points mentioned above P. gourlayi differs in having the antennae red, and the prothorax longer than broad, with three stripes formed of yellow and coppery scales.
P. lewisi Broun is a larger species, densely clothed with much larger grey scales, with the rostral carinae obsolescent, and a strong lateral costa (interrupted in the middle) on each side of the pronotum; on the elytra intervals 3, 5 and 7 are all strongly raised, the other intervals being flat, and 3 has a small tubercle at the top of the declivity.
Nestrius hudsoni, sp. n.
♂ ♀ Derm red-brown to piceous, with very sparse brownish setae. Head with the eyes in lateral view much nearer to the lower than to the upper surface of the rostrum; frons as wide as the base of the rostrum. Rostrum two-thirds as long as the pronotum in ♀, a little longer in ♂, gradually narrowing from the base to the middle, then widening rather strongly to the apex; dorsum moderately arcuate longitudinally, with a broad, shallow median sulcus and a fine carina on each side of it anteriorly. Antennae with the scape almost straight, gradually clavate, with long, erect setae, those on the inner (or anterior) side subclavate; funicle with joint 1 as long as 2 + 3, 3 - 7 subequal and slightly longer than broad; club not globose, but gradually narrowing basally and almost pedunculate. Prothorax a little longer than broad (8:7), strongly rounded laterally, widest much beyond the middle, rather deeply constricted at the apex, which is narrower than the base; dorsum with very large close punctures and sometimes a very shallow median sulcus, the setae fine, short and erect. Elytra elongate, ovate, obtusely acuminate apically in ♂, a little more sharply in ♀, jointly sinuate at the base, with a small granular elevation at the base of intervals 2 and 4; dorsum with rows of coarse punctures, the intervals being no wider than the septa between the punctures, each with a row of slender, erect or suberect, capitate setae which are longer posteriorly. Legs red-brown to yellowish, rather slender; hind trbiae of ♂ with a rather deep basal sinuation, followed by a small dilatation.
Length, 3.5–4.2 mm., breadth 1.2–1.5 mm.
New Zealand. Wilton's Bush, 3 ♂, 2 ♀, under dead leaves, iv. 1941 (G. V. Hudson).
The nearest ally of this species in B. prolixus Broun, which differs in having the club of the antennae globular and not gradually narrowing at the base; further, the prothorax is much narrower (8:6) and only slightly rounded laterally; and the elytra lack the basal granular elevations.
Unas conirostris, sp. n.
♂ ♀ Derm piceous, bare, the antennae and legs paler. Head separated from the rostrum by a very shallow transverse impression, the occiput with small sparse
punctures; frons about as broad as the base of the rostrum, with large close punctures, the interspaces being narrower than the punctures; eyes strongly convex. Rostrum much shorter than the pronotum (3:5), stout, somewhat curved, distinctly narrowed from base to apex, the punctures fairly close, gradually diminishing from base to apex, rather stronger in ♂ than in ♀. Prothorax about as long as broad, widest near the base, gradually narrowing in front and rather deeply constricted at the apex, the constriction not continued across the disk; dorsum with close, coarse punctures without any smooth median line. Elytra cylindrical, with comparatively deep striae containing strong punctures that excise the intervals, the lateral striae about as deep as the dorsal ones; the intervals hardly wider than the striae, with a row of minute granules towards the sides and apex. Legs with joint 3 of the tarsi not wider than 2 and not bilobate.
Length, 2.0–2.5 mm., breadth, 0.5–0.7 mm.
New Zealand. Titirangi, Auckland, 1 ♀, ix. 1916 (Brookes, type); Whangarei, 7 ♂, 5 ♀, from timber, x. 1940 (K. Harrow); Titirangi, 1 ♀, in flooring boards attacked by Ambeodontus and Torostoma, ii. 1945 (M. W. Carter).
It has seemed desirable to include this species provisionally in the genus Unas despite the narrow non-bilobate third tarsal joint, because in other respects it is closely allied to the unique genotype, piceus Broun. While, in general, this tarsal character seems to have generic value in the Cossoninae, there are a few cases (notably in Euophryum) in which it has been regarded as having specific value only.
Broun (Trans. N.Z. Inst., xli, 1908 (1909), p. 197) pointed out that the description of Microtribus pictonensis Sharp (1886) indicated that this species could not belong to the genus to which Sharp had assigned it. An examination of the unique type has shown that Broun was quite correct, for the species clearly belongs to Broun's genus Unas. The three species now included in this genus can be distinguished as follows:—
1 (4). Third tarsal joint bilobed and somewhat wider than the second; rostrum parallel-sided; elytra without granules; front tibiae with the mucro rudimentary.
2 (3). Head with the punctures behind the eyes finer and much sparser than those on the frons; pronotum more rounded laterally, with the interspaces as wide as or wider than the small punctures, the apical constriction not continued across the dorsum pictonensis Sharp.
3 (2). Head with the punctures behind the eyes as large and as dense as those on the frons; pronotum widest near the base, with the punctures larger and mostly wider than the inter-spaces, the apical constriction continued across the dorsum piceus Broun.
4 (1). Third tarsal joint not bilobate and not wider than the second; rostrum gradually narrowing from base to apex; elytra with extremely minute granules on the intervals towards the sides and apex; front tibiae with the mucro more distinct conirostris, sp. n.
Himatiodes, gen. n.
Head subglobose, short, separated from the rostrum by a transverse impression, the frons as wide as the base of the rostrum, the eyes quite flat, widely separated beneath. Rostrum directed downwards, parallel-sided to the antennae
and slightly wider in the subapical part; scrobes oblique, the upper edge reaching the lower margin of the eye, invisible from above. Antennae inserted close to the apex of the rostrum in ♂ and only slightly further back in ♀; scape reaching the eye, almost straight, gradually clavate; funicle 7-jointed; club broadly ovate, as long as joints 2–7, densely pubescent. Prothorax very nearly as long as broad, narrower at the apex than at the truncate base. Scutellum distinct. Elytra much broader at the shoulders than the base of the prothorax (3:2), with a distinct subapical depression and a callus above it; the striae broader than the intervals, stria 10 abbreviated, stria 6 reaching the base inside the shoulder. Legs with a long tibial uncus, but the mucro comparatively small and inconspicuous; tarsi with joint 3 much broader than 2 and deeply bilobate. Sternum with the front coxae not widely separated, the interspace about as wide as two-thirds of a coxa; the mesosternum depressed below the level of the metasternum, the intercoxal space a little wider than a coxa. Venter with the intercoxal process broadly truncate, much wider than a coxa.
Genotype: Himatiodes setulifer, sp. n.
This genus has no very close ally in New Zealand. From the comparatively few genera that have seven funicular joints Himatiodes differs in the following combination of characters: the unusually short and broad elytra, the deeply bilobate third tarsal joint, and the marked reduction in the size of the tibial mucro.
Himatiodes setulifer, sp. n.
♂ ♀ Derm red-brown, dull, with sparse short recumbent yellowish setae. Head with close shallow punctures and sparse setae, the flat eyes placed rather low down on the head. Rostrum a little longer than the pronotum in both sexes, moderately curved, rather closely punctate and setose in ♂, a little more sparsely so in ♀. Antennae with joint 1 of the funicle longer and thicker than 2, 3–7 transverse and closely approximated. Prothorax gently rounded laterally, widest beyond the middle, rather deeply constricted at the apex, the constriction continued across the dorsum, which is flat on the disk, closely and strongly punctate, the intervals narrower than the punctures, without any smooth median line. Elytra parallel from the roundly prominent shoulders to beyond the middle, rather broadly rounded at the apex; the broad striae containing large close punctures, the intervals narrower than the striae and subcostate, each with a row of numerous short recumbent setae. Underside coarsely and closely punctate throughout.
Length, 2.0–2.5 mm; breadth, 0.6–0.8 mm.
New Zealand. Titirangi, Auckland, 4 ♂, 3 ♀, from an Araliaceous tree, Nothopanax arboreum, v–vi. 1942 (M. W. Carter).