
Description
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Proportional dimensions in per cent. of total length.
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Trunk at pectoral origin: breadth 9.9; height 6.4.
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Snout length in front of: outer nostrils 6.8, mouth 8.8.
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Eye: horizontal diameter 3.9; vertical diameter 1.4.
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Mouth: breadth 7.8; length 4.0.
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Nostrils: breadth (between inner corners) 3.9.
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Labial furrow lengths: upper 1.4; lower 1.4.
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Gill-opening lengths: 1st 1.4; 2nd 1.5; 3rd 1.6; 4th 1.6; 5th 1.3.
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1st dorsal fin: vertical height 6.9; length of base 13.3.
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2nd dorsal fin: vertical height 6.9; length of base 11.6.
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Anal fin: vertical height 3.1; length of base 5.8.
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Caudal fin: upper margin 20.5; lower anterior margin 6.4.
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Pectoral fin: anterior margin 10.8; distal margin 9.4; posterior margin 5.4.
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Pelvic fin: anterior margin 4.8; distal margin 4.7; posterior margin 3.2.
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Distance from snout to: eye 8.8; 1st gill-opening 17.0; 5th gill-opening 21.3; 1st dorsal 26.7; 2nd dorsal 61.0; upper caudal 79.5; pectoral 20.2; pelvic 47.8; anal 63.3.
Interspace between: 1st and 2nd dorsals 21.7; 2nd dorsal and caudal 7.5; anal and caudal 8.8.
Distance from origin to origin of: pectoral and pelvic 27.5; pelvic and anal 16.2.
Head compact, depressed, large-eyed; trunk attenuate, slender, generally subcircular in section and gently tapered from the pectorals posteriorly. Height of trunk at origin of pectorals 1/12th of its length to origin of caudal. Dorsal and ventral profiles slightly and about equally arched. Length of body, measured to cloaca, 1/15th more than length of tail from same level. Caudal peduncle a little compressed and without keels or precaudal pits. Dermal denticles small, crowded to overlapping, with short pedicles, broad rhomboidal bases, and elongate horizontal blades. The blades tridentate, with sharply pointed teeth, the median tooth very long, the lateral teeth short and tending to point inwards. Each blade of those denticles taken from high on the sides at the level of the 1st dorsal fin bears three strong longitudinal ridges, one median and one lateral on each side, proximally flat-topped or rounded but distally sharp-edged and terminating at the end of the teeth. Denticles from the sides of the tail similar in shape but the blades bear numerous longitudinal ridges. The anterior two-thirds of the dorsal margin of the caudal with four to five irregular longitudinal rows of small

Text-Fig. 1.
Fig. A, Triakis attenuta n.sp., lateral view. Figs. B-C, dorsal and ventral views of head. Fig. D, outline of transverse section of head at level of eyes to show flange-like edges of head. Fig. E, left nostril. Fig. F, left lower tooth, 6th row from (centre of jaw. Figs. G-J, left upper teeth; G, 4th and 5th rows; H, 17th and 18th rows; I, 41st and 42nd rows; J, 47th, 48th and 49th rows. Figs. K-Q, left lower teeth; K, 1st and 2nd rows; L, 13th and 14th rows; M, 22nd and 23rd rows; N, 31st and 32nd rows; 0,44th and 45th rows; P, 51st and 52nd rows; Q, 56th and 57th rows. ANL, anterior nasal lobe; C, level of cloaca; E, eye; HE, edge of head; LJ, lower jaw; PNL, posterior nasal lobe; SBF, suborbital fold.

Text-Fig. 2. Fig. A, Triakis attenuata n. sp. dermal denticles from high on side at level of 1st dorsal Fig. B, dermal denticle showing underlying base. Figs. C–D, apical and lateral views of dermal denticle Figs. E—F, lateral and dorsal views of portion of dorsal margin of caudal fin showing upper rows of small granular denticles overlying a row of large subcircular denticles, the latter forming a prominence over a semi-naked area which ventrally is bounded by multi-ridged tridentate denticles.
granular subrhomboidal denticles, each with an incipient posterior or posterolateral tooth and numerous low ill-defined ridges Flanking these denticles on each side of the fin, a single row of large subcircular denticles, semi-erect, without distinct denticulation but weakly ridged, and forming a marked prominence along the edge of the fin. Below the row of large denticles an almost naked area, narrow in extent, with well-spaced bluntly tridentate denticles, and below these the normally shaped tridentate denticles.
Head measured to 1st gill-opening 5.8 in total length and less than interspace between 1st and 2nd dorsals. Head greatly depressed with a distinct, thin, flange-like edge, the latter especially prominent from the level of the nostrils to midway between the spiracle and the first gill-opening. Head broad, the least fleshy interorbital distance 3.2 in length of head Snout to eye 2.0 in length of head. Contour of snout tip from above broadly ovate, but just anterior to the eye, the edge of the head extends abruptly to form a distinct step in the contour.

Eye elongate and oval, 3 times as long as high, its horizontal diameter 1.5 in the interorbital. A prominent suborbital fold of skin along anterior two-thirds of eye, well separated from the lower lid anteriorly but continuous with it posteriorly. Spiracle* ovoid, large, placed slightly below level of centre of eye and behind it by a distance equal to the spiracular length. Longest diameter of spiracle directed posterolateral, its length 3.8 in horizontal diameter of eye. Gill-openings small, the anterior three concave, the 4th and 5th almost straight and above the basis of the pectoral. Length of the gill-openings increasing slightly from the 1st to the 4th the latter being 2.0 in the horizontal diameter of the eye Fifth gill-opening subequal to the length of the 1st. Interspaces between the gills 1.5 to 2.0 in the length of the gill-openings, that between the 4th and 5th the smallest Nostrils oblique, widely separated, the distance between their inner corners 1.7 in the distance from snout to tip of nostril and 4.5 in length of head. The anterior margin of each nostril expanded as a subtriangular lobe directed posteriorly. and the inner portion of the posterior margin developed as a small low folded lobe also directed posteriorly. Mouth broad and highly arched, the apex of the arch acute. Width of mouth 2.2 in length of head, and length of month, measured along midline from anterior of upper jaw to level of the outer angles 4.3 in the same ratio Upper and lower labial furrows equal in length, strongly developed but short, their length 2.5 in the horizontal diameter of the eye.
Teeth 98/114 small, closely crowded so that the successional series overlap. Serial arrangement of teeth primarily anteroposterior, but becoming increasingly oblique towards the angles of the jaws. Both jaws with 5 to 6 series of teeth functional at centre of mouth, 3 to 4 series at the angles. Teeth at angles of mouth considerably different from those at centre, but all comprise essentially a single long pointed erect or slightly oblique major cusp, flanked by one to several mall lesser cusps on each side, borne on a stout base carrying numerous well-defined longitudinal basal striae. The greatest number of lesser cusps produced on those teeth towards the angles of the jaws. In the upper jaw, the 1st to 23rd rows counted from the centre towards the angle of the jaw comprise almost symmetrical teeth with a single long pointed major cusp directed slightly obliquely, flanked on each side by two small pointed lesser cusps, the outermost lesser cusp on each side the smaller; the 24th to 44th rows similar to those medially adjacent but with the large major cusp directed more obliquely, and only on lesser cusp on the lateral side; the 45th to 48th rows with markedly asymmetrical teeth, the major cusp placed well on the lateral side of the tooth, and flanked laterally by one small lesser cusp and medially by three small lesser cusps; tie 49th row similar to those rows just medial but with the major cusp flanked on its lateral side by two small lesser cusps. The teeth in the lower jaw similar to those in the upper jaw, and showing the same general arrangement with the teeth towards the angle of the jaw carrying more medial lesser cusps than those at the centre However this pattern of increase in cusp number is not regular, as some of the paramedian rows carry fewer medial lesser cusps than either the medial or lateral rows. The teeth in the 1st row of the lower jaw are symmetrical with the major cusp flanked by one small lesser cusp on each
[Footnote] * A prominent, externally visible aperture of 1.0 mm. diameter is located on the dorsomedial well of the spiracle, and indicates the origin of a diverticulum extending towards the auditory capsule. Similar diverticula are recorded from other genera (Daniel, p. 153), but usually they are leas obvious externally.

side; the 2nd to 6th rows with a single lateral lesser cusp and two medial lesser cusps; the 7th to 18th rows similar to the 1st row with one lesser cusp on each flank; the 19th to 25th rows like the 2nd to 6th rows, with one lateral and two median lesser cusps, but with a shorter and more oblique major cusp; the 26th to 40th rows with one lateral and three medial lesser cusps; the 41st to 48th rows similar to those medially adjacent but with four medial lesser cusps; the 49th to 54th rows the same but with five medial cusps; the 55th and 56th rows with two lateral and four medial lesser cusps; and the 57th row not following the previous pattern, but with the major cusp flanked laterally by five or six lesser cusps and medially by only two.
First dorsal originates by a very shallow angle from the dorsal profile just above the insertion of the pectoral. Base of 1st dorsal 1.4 in the head or equal to the distance between anterior of eye and 5th gill-opening. Height of 1st dorsal 2.0 in its base. Anterior margin almost straight, apex acutely rounded and over posterior third of base, posterior margin concave. Posterior corner sharply pointed, free for a distance 2.3 in height of fin, and terminating anterior to pelvics by a distance equal to length of base of the latter. Second dorsal originates midway between posterior tip of pelvics and origin of anal, its base, height, and margins similar to those of 1st dorsal, but with the apex over the end of the base. Posterior free corner terminates about midway between posterior corner of anal and origin of caudal. Caudal 4.9 in total length, epiural lobe very low, terminal lobe truncate and with a distinct subterminal notch. Anterior hypural lobe narrow with a straight anterior margin, acutely rounded apex and concave posterior margin. Pectorals originate at level of 4th gill-opening, base 3.2 in the head. Anterior margin weakly convex, its length twice that of the base. Distal margin weakly concave, posterior margin convex, anterior and posterior corners acutely rounded. Pelvics small, anterior margin straight and 1.3 in the base, the latter 2.9 in the head. Distal and posterior margins almost straight. anterior free corner moderately rounded and posterior tip acutely pointed Claspers semicircular in section, sharply pointed posteriorly and projecting beyond the tips of the pelvics by a distance equal to the posterior margin of the latter. Anal originates behind origin of 2nd dorsal, with its free posterior tip terminating below end of base of 2nd dorsal. Height of fin 2.0 in its base. the latter being 2.0 in base of 2nd dorsal. Anterior margin weakly convex, distal margin concave. Apex sharply rounded, posterior corner pointed.
Colour: Generally brownish-grey, but lower surface of trunk, pectorals and caudal, and distal portions of the other fins greyish-white.
Holotype in the Dominion Museum, its catalogue number 1509.
