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Volume 83, 1955-56
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New Zealand Cretaceous Fossil Plant Localities.

N.Z. Geol. Surv. Age and Locality.
Register.
Albian (Clarence Series).
B470 Tributary of Cover Creek, Coverham, from concretion containing
Albian Mollusca (S35/617, grid reference 155463).
? Turonian (Raukumara Series)
The following collections are from the Seymour Coal Measures conformably overlain by marine sediments containing basal Raukumar [ unclear: ] (? lower Senonian) mollusca:
B125 Quail Flat, Clarence River (Gs560 = S41/500).
B166 Quail Flat, Clarence River (Gs568 = S41/492).
B351 West bank, Seymour River (S41/549, grid reference 743147).
B356 East bank, Seymour River (S41/537, grid reference 734091).
B358 Quail Flat, Clarence River (S41/550, grid reference 722145).
Lower Senonian (Paparoa Coal Measures)
B23 Ford Formation, drillhole 241 (S44/731, grid reference 832973).
B172 Waiomo Formation, Paparoa mine (S44/483, grid reference 878001)
B52
B428 Rewanui Formation, Strongman mine dump (S44/752, grid reference 800000).
B19 Goldlight Formation, drillhole 266, Brunner Bridge (S44/727, gird reference 830904).
Upper Senonian (Sub-Piripauan)
B160 Shag Point (Gs414 = S146/519, additional specimens in Otago Museum).
B161 Pakawau (Gs410 = S3/492, grid reference 910070; additional spec [ unclear: ] mens in Canterbury Museum).
B530 Mikonui Stream, coast south of Oaro River, Marlborough (S56/510, grid reference 799762).
B160 and B530 are from coal measures underlying fossiliferous Piripauan
Maestrichtian.
B182 Bulls Point and Batley, Kaipara Harbour, from concretions containing Maestrichtian ammonites (N28/626, grid references 806390 and 843365).
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The collections from Coverham (Albian) and Kaipara (about Maestrichtian) are dated from associated marine fossils. Those from Shag Point and Mikonui Stream are from coal measures conformably underlying Piripauan (upper Senonian) marine sediments and are probably themselves upper Senonian. The Pakawau coal measures have been correlated with those at Shag Point on the evidence of plant microfossils (R. A. Couper, pers. comm.).

The collections from Seymour River and Quail Flat (middle Clarence Valley) are from coal measures conformably underlying marine sediments containing Trigonia glyptica, T. meridiana (Woods, 1917) and fragments of a giant Inoceramus which is the zone fossil of the basal Raukumara Series as redefined by Wellman (1955). Wellman (1955) has correlated the base of the Raukumara Series with the base of the Senonian. The Seymour River coal measures are thus either basal Senonian or probably Turonian. They are here classed as ? Turonian.

The Paparoa Coal Measures have never been dated from satisfactory paleontological evidence and are variously classed as lower Cretaceous (?) (Wellman, 1950), possibly pre-Cretaceous (Cotton, 1951), probably upper Cretaceous (Gage, 1952) and lower Cretaceous (Couper, 1953).

The fossil cycads and ferns described below provide the following additional evidence for dating the rocks in which they occur:

The Seymour River coal measures contain leaves of Ptilophyllum and Pterophyllum, genera recorded in New Zealand from the Jurassic but not previously from the Cretaceous. Pterophyllum does not range above the Jurassic in other parts of the world (Seward, 1933: 332) and is apparently an archaic element in the New Zealand Cretaceous. Ptilophyllum ranges up to the middle Cretaceous in other countries (Seward, 1933, 413).

In New Zealand, Taeniopteris stipulata apparently replaced the Jurassic and Neocomian Taeniopteris spatulata during the Lower Cretaceous. No beds are known to contain both species. T. spatulata is present in the Styx River Series of Queensland, tentatively classed as Cenomanian by David and Browne (1950), but T. stipulata is unknown in Australia. If a change from T. spatulata to T. stipulata occurred at the same time in Australia and New Zealand, the Seymour River coal measures (which contain T. stipulata) are probably post-Cenomanian and may well be Turonian as suggested above.

Four species from the Paparoa Coal Measures are unknown in the upper Senonian of New Zealand, but range back to middle Cretaceous or older strata (Sphenopteris mackayi, Phyllopteris lanceolata, P. expansa, and Coniopteris ? lobata). On the other hand, the Paparoa Coal Measures lack the genera Pterophyllum and Ptilophyllum that persist into the Seymour River coal measures and contain more angiosperms than known pre-Senonian floras in New Zealand. They are thus probably older than upper Senonian but younger than Turonian, that is, lower or middle Senonian.

The Shag Point and Pakawau Coal Measures have been correlated from their plant microfossils (R. A. Couper, pers comm.). They seem to be younger than the Paparoa Coal Measures because they lack the older Mesozoic species of Phyllopteris, Coniopteris and Sphenopteris (mackayi) and contain three species of fern unknown in Paparoa or older beds.

As the Shag Point and Mikonui coal measures underlie Piripauan (upper Senonian), they and the Pakawau coal measures may be slightly older than the plants from Kaipara, collected from concretions containing ammonites probably of Maestrichtian age (Mason, 1953: 356). The Kaipara beds apparently lack the

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common upper Cretaceous species Taeniopters stipulata and contain Taemopteris batleyensis which has not been found at other horizons and thus may be an index species for the Maestrichtian, although it is a pteridophyte (Edwards. 1934) and thus not the descendant of T. stipulata.

The known ranges of pteridophytes and cycads in the Cretaceous of Australia and New Zealand, on which the above conclusions are based, are summarised in Table I.

  • Cladophlebis australis (Morr.)

  • Coniopteris? lobata (Oldham)

  • Taeniopteris spatulata MacClell.

  • Phyllopteris lanceolata Walk.

  • P. expansa Walk.

  • Zamites takuraensis Walk.

  • Lycopodium of volubile Forst. f.

  • Sphenopteris mackayi n. sp.

  • Taeniopteris stipulata Hect.

  • Pterophyllum (clarencianium n. sp.)

  • Ptilophyllum (seymouricum n.sp.)

  • Cladophlebis wellmanii n. sp.

  • C. prisca (Ett.)

  • C. obscura (Ett.)

  • Sphenopteris pteriodies (Ett.)

  • Sphenopteris sp. (Edwards 1926)

  • Taeniopteris batlcyensis Edw.

Waikato Heads1 Neocomian
Burrum, Queensland2 Aptian-Albian
Coverham Albian
Styx. Queensland3 Cenomanian
Seymour R. coal measures Turonian
Ford
Waiomo
Rewanui
Goldlight Paparoa Coal Measures
Lower Senonian
Pakawau4
Shag Point4
Mikonui Upper Senonian
Kaipara5 Maestrichtian
Table I—Stratigraphic Distribution of New Zealand and Australian Cretaceous Pte [ unclear: ] dophytes and Cycads. 1Arber 1917, Edwards 1934; 2Walkom 1919 David and Browne 1950, 3Walkom 1919; 4Ettingshausen 1887, Suggate and Couper 1952, 5Edwards 1926.