
Discription of Male
Antennae. First: Peduncle longer than 6-segmented flagellum; flagellar segments longer than wide, short setae on distal angles; peduncle, 2nd segment as long as 3rd, shorter than 1st, has a few stout end and marginal spines, as also 3rd; 1st segment has a few weaker end spines, a few small surface spines. Second: Peduncle slightly longer than flagellum; flagellum of 15 segments, segments wider than long, have 4 equidistant groups of strong setae distally, end segment tufted. Peduncle, 3rd segment ½ length 4th, 4th ½ 5th; 3rd and 4th have strong spines superiorly and distally; 4th has distal ½ of inferior margin also spined, 5th has strong spine groups on both margins.
Mouthparts. Upper Lip: Setose, not spined. First Maxillae: Palp of 2 segments, rudimentary. Maxilliped: Inner plate has 3 strong teeth distally; triangular field of setulose spines below teeth on both surfaces, much more extensively on inner, long setulose spines down cleft to basos; outer margin has 4 setulose spines outside distal teeth. Outer plate very bluntly pointed, reaches nearly ½ along carpus; outer margin has row of setulose spines distally; inner has very numerous herring-bone tipped spines to ischium. Basos has strong marginal and end spines; 3 or 4 strong spines on ischium distal angle; several strong spines on merus outer distal margin and end, inner margin is about ½ length of outer. Carpus as wide as, or wider than, long; inner surface expanded in flattened wing slightly farther distally than rest of carpus; wing has distally 3 or 4 strong spines; inner margin has short spines most of length. Propod much narrower, as long, narrowing somewhat to bluntly rounded tip; a few small surface spines; inner and end margins have strong spines almost entire length; dactylos rudimentary, square, scale-like, has spines distally and around base.
Gnathopods. First: Sideplate, anterior margin slightly concave, ventral rounded with a few small marginal spines; posterior margin and surface sparsely spined. Basos constricted proximally, posterior and both anterior margins spined, a few spines on posterior surface, posterodistal angle strongly spined. Ischium subrectangular, longer than wide, posterior margin spined. Merus posterior margin ½ basos length, convex and strongly spined, a few surface spines, anterior margin contiguous with proximal ½ of carpus posterior margin. Carpus ¾ basos length; anterior margin slightly convex, strongly spined; posterior margin and surface strongly spined, free margin expanded medially to small scabrous pellucid lobe guarded by spines. Propod ⅔ carpus length, widening slightly distally, greatest width ½ length; anterior margin slightly convex, both strongly spined; posterior surface spined, distal surface widened in scabrous pellucid lobe, forming slightly concave palm; group of fine spines ½ along each side of palm; palm ½ length of long curved overlapping dactylos; dactylos ½ propod length, 3 short stout marginal spines on inner margin proximally. Second: Sideplate widening greatly distally, ventrally convex; ventral and posterior margins and surface have short spines; posterior margin straight, almost imperceptibly excavate. Basos concave, margins have numerous short spines; distal width ⅓ to ¼ length; posterodistal angle spined, anterodistal naked. Ischium ¼ basos length, posterior margin spined, anterior expanded distally in shallow lobe. Merus subrectangular, slightly shorter than ischium, sparsely spined; anterior margin contiguous with small subcrescentic carpus. Propod distally as wide as convex anterior margin is long, wider than posterior margin is long; palm transverse, sinuous, concave near dactylos hinge and convex posteriorly; row of spines marginally, a second parallel row along surface below palm; long curved dactyos just reaches end of propod, has inner thickening near base; small stout characteristic projection on palm just inside dactylos hinge.
Peraeopods First: Sideplate widening so distally as wide as deep, margins more or less straight, posterior and ventral margins and surface spined, minutely excavate. Basos margins spined, constricted proximally. Ischium ¼ basos length, posteriorly spined. Merus ½ basos length, margins strongly spined, anterior convex, spines more numerous posteriorly. Carpus slightly narrower, ¾ merus length; anterior margin spined medially and distally; posterior strongly spined all along. Propod as long, narrower, has strong marginal spines, mostly posteriorly. Dactylos stout, curved, ½ propod length. Second: Posterior margin of sideplate concave; segments proportionately shorter than in Pr. 1; short, stout dactylos has spur ½ along inner margin, otherwise like Pr. 1. Third: Sideplate anterior lobe subtriangular, much the larger, reaches end of basos, posterior subtriangular but ventrally rounded, slightly longer than ½ anterior lobe, has about 3 medium-length spines on short anterior margin, minute spines on posterior becoming progressively larger distally. Basos expanded, ovate, as wide as long, anterior margin has about 5 groups of strong spines; posterior about 14 short spines. Ischium small, spined anterodistally. Merus piriform; length ⅔ basos, greatest width ¾ length, about 4 groups of strong spines on margins, more numerous anteriorly. Carpus similar, ⅔ merus length, narrower, 3 groups of strong spines posteriorly, 4 groups anteriorly Propod slightly longer, ½ width,

margins spined; dactylos as in Pr. 1 Fourth: Sideplate ovate, reaching ¾ along basos; about 13 spines anteriorly and posterodistally. Basos widest distally; nearly as wide as long; anterior margin strongly spined, a few spines on surface; about 12 small spines proximally on posterior margin. Ischium small, wider than long, anterodistal angle spined; merus subtriangular, length ¾ basos, distally nearly as wide as long and more than twice proximal width, margins strongly spined, a few surface spines. Carpus expanded, nearly as long as merus; medially as wide as long; 4 spined corrugations anteriorly, strong spines on anterior surface; posterior margin V-shaped, proximal ½ free, distal ½ has 3 groups of strong spines. Propod as long as basos, straight and narrow, strong spines on margins; dactylos slender, almost ½ propod length. Fifth: Side-plate small, has about 6 spines ventrally. Basos ovate, greatest width slightly more than length; anterior margin and surface have numerous strong spines; a longitudinal row on surface medially; posterior margin strongly convex, not markedly crenulate, has about 24 long fine marginal spines. Merus ¾ basos length, widening distally, distal width ½ length; margins strongly spined. Carpus slightly shorter and narrower, about 4 groups of strong spines on each margin. Propod as long as carpus, ½ its width; linear, margins strongly spined, dactylos as in Pr. 4.
Epimeral Plates. First: Subtriangular, anterior margin has about 5 strong spines; posterior sinuous, about 3 minute well-spaced spines. Second: Comparatively narrow, anterior margin rounding to ventral, has 7 or 8 strong spines, a few on anterior surface, posterior margin straight, has about 5 small marginal spines. Third: Subrectangular, anterodistal angle rounded broadly, anterior margin has about 5 strong spines, ventral none; posterior slightly concave with about 9 small spines, anterior surface also spined.
Pleopods. Rami as long as peduncles, which have strongly spined outer margins; inner margin and surface proximally spined in 1st and 2nd, 3rd has row of spines down inner margin; 2 coupling spines; outer margin of 2nd and to lesser degree 3rd pleopod finely bristled between spines; rami have about 6 to 8 superficial segments with paired long plumose setae; inner have 2 or 3 spines on surface proximally, outer 2 to 4 small spines proximally on outer margin.
Uropods First: About 5 spines on peduncle ventral surface. Outer ramus has about 8 strong dorsal and end spines; inner 2 dorsal rows of about 4 spines each, 2 at end. Second: Rami slightly longer than peduncle, outer has about 6 stout marginal and end spines; inner about 10 spines on margins and end. Third: Peduncle subglobular, 3 or 4 distal spines; ramus longer, cylindrical, has end row of 3 or 4 long and 2 short spines, 9 or 10 spines on upper distal surface Telson: Slightly emarginate, ovate, strongly spined.

