Go to National Library of New Zealand Te Puna Mātauranga o Aotearoa
Volume 84, 1956-57
This text is also available in PDF
(4 MB) Opens in new window
– 417 –

Redescription

♀ Dorsal surface and appendages rather shining, impunctate. pronotum and scutellum minutely roughened, head smooth and highly polished, body, appendages, and veins and margins of fore wings with a covering of fine, pale brown hairs, those of head, pronotum and appendages rather long, suberect, not closely down-like.

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

Head From above (excluding rostrum, labrum and “neck”) about ⅕ longer than pronotum (95 80), anterior lobe a little wider across eyes than posterior lobe (38:35) and 5/7 as long again (60:35); posterior lobe as long as wide, subglobose, widest at middle, dorsal surface convex; both anterior and posterior constrictions well marked, but the sides of the head not incised at the level of the former and gradually convex behind the eyes, the dorsal surface of anterior and posterior lobes rather gradually declivous into anterior constriction, base of posterior lobe similarly declivous into posterior constriction and the sides only slightly incised at its level (at anterior margin of “neck”) Eyes moderate, from above each about ⅓ as wide as interocular space (7 5:23), from below subequal to it (13 12), in side view convexly subovoid, with the postero-ventral margin shallowly concave, nearly as wide across greatest diameter as high (20:22), occupying about ¾ of height of head (22.28). from above about half as long as posterior lobe (18:35). Ocelli moderate, set towards sides of head and about three ocellus-widths apart Rostrum with total length subequal to that of head, third segment with sides only shallowly incurved just beyond base, slightly and gradually widened

– 418 –

toward apex, which is not greatly incrassate ventrad, in side view only slightly deeper than base (10:8); relative length of segments I-IV, 14:27:30:21. Antennae clothed with suberect hairs longer than width of the segments; over ¾ as long again as head and very nearly as long as head and pronotum together; relative length of segments I-IV, 30:54:48:38; first segment the stoutest, gradually thickening toward apex, second and third segments cylindrical, the second stouter than the third: fourth slenderly fusiform.

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

Pronotum ¼ as wide again across humeral angles as long in median line (100:80), middle lobe ⅜ as long again as posterior lobe (37:27) and 2⅓ times as long as anterior lobe (37:16), posterior lobe three times as wide as anterior lobe (100:33) and 3/7 as wide again as middle lobe (100:70); anterior lobe with a raised transverse callus posteriorly; middle lobe with a distinct median longitudinal impression deepest and widest just before base, continued as a less distinct impression on anterior lobe and as a low, narrow carina on posterior lobe; middle lobe with a small sublateral fovea on each side at about half its length, and behind it a rounded tuberosity, posterior margin distinctly though rather thallowly emarginate, obtusely angulate in the middle, humeral angles broadly rounded Scutellum with disc flattened; apical lobe over ⅓ as long as whole scutellum (15:43), convexly carinate, apex acutely rounded.

Fore wings fuscous, veins brown, venation as for genus (no basal cell, discal cell open, veins not conspicuous nor strongly raised).

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

Legs. Front legs moderately thickened, middle and hind legs rather slender; front and middle coxae subglobose, hind coxae more linear Front legs: length of coxa, femur, tibia, tarsus, as 34:105:90:26; tarsus one-segmented, nearly cylindrical, ⅖ as wide as long, with two pairs of very short, thorn-like spines ventrally on apical third, posterior (outer) claw equal in length to tarsus, anterior (inner) claw longer (31.26), tibia gradually expanded toward apex, which is as deep as tarsus is long, and bears, in addition to long pale hairs, a well developed pecten along inner margin and a group of six spines on ventral angle, arranged in rows of 1, 3, and 2, the single more dorsal spine slender, the others stout, the three middle ones the longest, femur nearly ¼ as long as deep in middle as long (25.105). Middle and hind tibiae slender, apex with a pair of ventro-lateral pectines and, mid-ventrally between them, a pair of slender spines longer than the pectines Middle and hind tarsi two-segmented, the basal segment very short, longest ventrally; apical segment subcylindrical, somewhat flattened laterally, without spines, with a pair of nearly equal claws 1/1 as long as tarsus Hind legs, length of coxa, femur, tibia, tarsus, as 45:115:120:45, femur ⅙ as deep as long (19:115), inner claw ½ as long as tarsus (15:45), outer claw scarcely shorter (14:15).

Colour. Head, pronotum, scutellum, and venter of thorax fuscous reddish brown, fore wings fuscous brown with veins brown, not conspicuous, antennae, rostrum, and legs testaceous, the legs more or less infuscated; abdomen duller, paler, yellowish brown, except for the dark last visible sternum (eighth), the posterior margin of which is strongly convex, the infuscated venter of the basal sternum (second), a dark, median, transverse spot near posterior margin of seventh sternum, two ventro-lateral rows of small dark spots near anterior margin of 4th–7th sterna, and above and laterad of them on each side a row of much larger dark patches on 2nd–7th sterna, not reaching their posterior margins, two sublateral rows of similar patches on the corresponding terga, eighth tergum dark, except for posterior margin, ninth tergum and anal tube dark, the posterior margin of the former concave.

Length (to apex of closed wings), 6.2–7.0 mm (length of abdomen varies with state of ovarian development and in pregnant female may greatly exceed wings).

Width (across eyes) 0.51–0.55 mm, (across base of pronotum) 1.33–1.47 mm (across closed wings) 1.33–1.70 mm.

: There is a marked sexual dimoiphism, and the males are readily distinguished from the females, particularly by the more slender legs, especially the front pair, the larger and more prominent eyes and ocelli, the more yellowish veins of the fore wings, the generally smaller and more slender form, and the presence of the capsule-like pygophor.

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

Differences from ♀: From above head (excluding rostrum, labrum, and “neck”) nearly ¼ as long again as pronotum (68.55); anterior lobe equal or subequal in width across eyes to posterior lobe (31:30); ratio of length, anterior:posterior lobe.: 43.25, posterior lobe rather wider than long, somewhat more globose than in ♀, with the anterior constriction more abrupt. Eyes relatively larger and more prominent than in ♀, from above each about ½ as wide as interocular space (7 5:16), from below about half as wide again (11.5:8); ratio of dorsal length of eye. posterior lobe ·· 13:25. Ocelli larger and more prominent than in ♀, less than two ocellus-widths apart Rostrum a little shorter than head (62:68); third segment gradually and moderately incrassate near apical third, in side view ratio of depth subapically:depth at base :: 7.5:6; relative length of segments I-IV, 7:15:25:15. Antennae twice as long as head and 1/7 as long again as head and pronotum together; apical segment with four whorls of very long, erect hairs about four times as long as width of segment; relative length of segments I-IV, 20:46.42:32. Pronotum ⅔ as wide again across humeral angles as long in median line (77.55), 2 ½ times as wide as anterior lobe (77:30) and ⅝ as wide again as middle lobe (77:47), middle lobe ⅝ as long again as posterior lobe (27.17) and 2 ½ times as long as

– 419 –

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

anterior lobe (27.11), constriction between anterior and middle lobes much longer anteroposteriorly than in ♀, forming a definite trough (this has been included with the middle lobe in the measurements of length above), posterior margin more deeply emarginate than in ♀ (posterior lobe thus relatively shorter in mid-line). Fore wings subfuscous, veins distinct, though not strongly raised. Legs, especially tibiae, more slender than in ♀. Tibiae, especially hind tibiae, curved Front legs: length of coxa, femur, tibia, tarsus, inner claw, as 23:76: 70:16.20, front tibiae less expanded than in ♀ at apex, where only ⅕ as deep as long (3/10 in ♀), front femora only about ⅙ as deep in middle as long (13:76) Hind legs. length of coxa, femur, tibia, tarsus as 35:90:110:35, femur ⅛ as deep as long. Abdomen: posterior margin of eighth sternum nearly straight; pygophor dark shining brown, open posteriorly; rest of abdomen yellowish brown, eighth sternum rather darker Length 5.2–5.5 mm Width (across eyes) 0.41–0.44 mm, (across base of pronotum) 1.03–1.09 mm.

Specimens Examined.1 ♂, Karori Reservoir Reserve (Wellington), 27.12.1934 (No. 104a) Wainui-o-mata (Well) 1 ♂, Christmas, 1936 (104b); 4 ♂ ♂, Christmas, 1939 (104c); 1 ♂, 16.1 1944 (104d), all G. V. Hudson Coll. (D.M.).1 ♂, Akatarawa (Well.), ex lichens, 10.2.1948, J. T. Salmon (D M) 1 ♀, Vinegar Hill.

Picture icon

Fig. 32—Systelloderes maclachlani (Kirkaldy) a ♂ head and pronotum, b ♂ fore leg, posterior (outer) aspect, c, 5th instar nymph

– 420 –

Reserve (Upper Rangitikei R), 12.12.1948, R. R. Forster (C M.). 2 ♀ ♀, Feilding (Well.), leaf mould, 16.1.1952, R. R. Forster (C.M.) 1 ♀, Mt Moehau-Te Hope Track (1,500–2,400ft), Coromandcl Peninsula (Auck.), leaf mould, rain forest, 17.1.1952, T. E. Woodward. 1 ♀, Coromandel-Whangapoua Rd. (Auck.), leaf mould, bush on summit, 18.1 1952, E. T. Giles 1 ♀, Tararua Ranges (ex leaf mould from Judd Ridge), 12.1.1954, T. E. Woodward.