
Genus Novoplectron nov.
1897. Pleioplectron Hutton (in part), Trans. N. Z. Inst. 29, pp. 232–233.
Hutton erected the genus Pleioplectron in 1897 and placed in it four new species—P. simplex from N. Canterbury and Banks Peninsula, P. hudsoni from Wellington, P. pectinatum from Banks Peninsula, and P. diversum from Upper Wanganui. In 1904 he added to the genus a new species, P. serratum from Pitt Island, Chatham Islands. Comparison of specimens collected on the Chatham Islands 1954 Expedition with Hutton's generic description, has revealed five distinct points in which they differ. Hutton says, “Hind femora without apical spines”, but all the specimens I have examined have a sharp, well-defined prolateral apical spine. His description of the hind tibiae as having three pairs of apical spines “of which the superior are acicular, hairy; more than half the length of the first joint of the tarsi; the middle pair about half the length of the superior pair, the inferior pair quite small”, is

not correct as they possess a fourth pair above, equal in length to his “middle pair”, which Hutton failed to observe. Describing the hind tarsi, Hutton says, “The first joint longer than the other three together”, while the first segment is actually not quite as long as the other three together. According to Hutton the subgenital plate of the male is longer than broad, but it is really broader than long. In the female he says the subgenital plate is short and has “the apex with three points”. This does not agree with my specimens as the plate is short and widely emarginate; but in Hutton's type material for the species the plate is not visible.
From the foregoing it can be seen that the differences between P. serratum and Hutton's generic description of Pleioplectron when considered together are major ones. Thus it appears that P. serratum was not placed in the correct genus. Any one of these characters might be disregarded generically, but together they have a definite significance and so, because of this, I propose to make P. serratum the type species of a new genus Novoplectron.
The genus Novoplectron is now defined as follows:
Body rather stout, sparsely clothed with short setae. Antennae very long and tapering, three times as long as body, almost touching at their bases; scape about four times as large as pedicel, which is narrower than scape, but broader than other segments; pedicel as broad as long; third segment narrower than pedicel, on dorsal aspect one third as long again, and on ventral aspect half as long again as pedicel; from fourth segment onwards to about 0.33 length of flagellum, segments broader than long, more distally segments become unequal in length and steadily decrease in size Scape thickly clothed with setae; segments from pedicel to fourteenth segment sparsely clothed with setae; from fourteenth segment onwards all segments thickly clothed with short golden setae. Head vertical; compound eyes laterad, nearly elliptical; a single anterior, white, median ocellus only. Fastigium almost as high as long, rising abruptly, slightly sulcate, with base touching scape of antennae. Mandibles small. Maxillary palpi with third segment 0.625 length of fourth. Pronotum rounded anteriorly and produced in front over occiput, truncated posteriorly; pronotum, mesonotum and metanotum not margined. Sternum transversely narrowed, metasternum bearing a median tubercle. Fore coxae close together, but not quite touching, each armed laterally with a spine. All femora sulcate ventrally. Femora, tibiae and two proximal segments of hind tarsi armed with variable numbers of spines above and below. No spines occur on fore femora, or fore and middle tarsi. Apical spines on femora, tibiae and two proximal segments of hind tarsi constant in number–fore femur bears one spine beneath prolaterally; fore tibia bears four spines, one above and one beneath, both prolaterally and retrolaterally; fore tarsus is unarmed; middle femur bears two spines beneath, one prolateral, the other retrolateral; middle tibia bears four spines, one above and one beneath, both prolaterally and retrolaterally; middle tarsus unarmed; posterior femur bears one spine above prolaterally; posterior tibia has a pair of apical spurs above clothed with setae, two pairs of apical spines–one pair above and one beneath, and a pair of smaller subapical spines beneath, one from each pair being prolateral and the other retrolateral; posterior tarsus two proximal segments each has two spines above, one prolateral and one retrolateral; other two segments unarmed. Proximal segment of hind tarsus nearly as long as the other three segments together. Cerci long, slender, tapering, unsegmented, clothed with long and short setae. Subgenital plate of female much broader than long, with distal margin widely emarginate. Ovipositor sabre-shaped, 0.66 to 0.75 as long as body Subgenital plate of male triangular, broader than long, sides spreading slightly proximally, tapering abruptly to concave distally, with a rounded apex; lanceolate distal portion thickly clothed with short setae on both dorsal and ventral surfaces Laterally the plate bears two styli thickly clothed with short setae; the lanceolate distal portion of the plate and the two styli are joined laterally by a connective membrane.
Type species for the genus: Novoplectron serratum (Hutton).
Novoplectron serratum (Hutton), 1904.
1904. Pleioplectron serratum Hutton, Trans. N.Z. Inst 36, p. 154.
1925. Pleioplectron serratum Hutton, Rec. Cant. Mus. Vol. II, No. 5, pp. 306–307, Figs. 6–8.
Plate 24, Figs. 3, 4. Text-fig. 2, Figs. 1–4.
There are several small points in which my material differs from Hutton's description of the species Pleioplectron serratum, but none of them are as important as the generic differences. Hutton says, “Fore and middle femora unarmed below”, but in some specimens the middle femora do bear spines, although the fore femora

never have them. However, in the type material neither fore nor middle femora possess spines. The hind femora, according to Hutton, are armed “with fifteen teeth below in each row, those of the outer row smaller than those of the inner”. In 1924 Miss Lysaght found the number of spines on the hind femora very variable, and counts of the type material gave for the two females 15, 16; and 10, 13; and the two males 22, 14; and 14, 14. In 1954 I re-examined this type material and found the spines for the two females were really 17–17, 18–17; and 10–18, 12–16; and the two males 19–21, 15–24; and 13–18, 14–17. Although Miss Lysaght's figures and mine do not agree, they both differ from Hutton's and show the wide range of
Text-fig. 2–Novoplectron serratum (Hutton). Fig. 1–Female genitalia ventral view. Fig. 2–Female genitalia dorsal view. Fig. 3–Male genitalia ventral view. Fig. 4–Male gentalia dorsal view.
Scale 0.3 cm applies to Figs. 1, 2 Scale 0.2 cm applies to Figs. 3, 4. For list of abbreviations, see index at end of paper,

variability which occurs. Hutton describes the fore and middle tibiae as having three pairs of spines below and none above. This, however, is not always the case and the middle tibiae usually bear two or three prolateral spines above as well as an occasional-retrolateral one. These spines are present in the type material, but were missed by Hutton. Hutton says the spines on the hind tibiae range from 16–20, but in actual fact the amount of variation is considerably greater. In the three female types the subgenital plate is hidden and so is undescribed; however, it is visible in some of the females collected on the 1954 Expedition and is short and widely emarginate. Hutton's measurements of the type material agrees with my measurements except that the length of the pronotum in the male is 8 mm., not 15 mm.; the width of the mesonotum is 12 mm., not 9 mm. in the male, and 12 mm., not 8 mm. in the female; and the total length is 27 mm., not 32 mm. in the male, and 22 mm., not 27 mm. in the female. The measurements tend to vary among different members of the species.
In 1925 Miss Lysaght added further points to Hutton's description of P. serratum. Of the antennae she says, “The first two joints of the antennae are longer than broad, those following are broader than long till about one-third of the length from the tip, where they become longer in proportion”, but although the scape is longer than broad, the pedicel is as broad as long and the third segment is longer than broad; from the fourth segment onwards to about a third of the length of the flagellum the segments are broader than long Miss Lysaght recognised the range of variation which occurs in the spination of the legs. Describing the apical spines of the fore tibia she says, “There is one pair of apical spines below, and a single apical spine at the side”, but in all the specimens I have examined there are four apical spines, two above and two beneath. Unfortunately most of her remarks on the apical spines are so confusing that they cannot be followed, and therefore must be disregarded. With regard to the spination of the hind tibiae she says, “The number of spines on the hind tibia, which Hutton gives as 16 to 20, ranges from 18 to 28 on either margin in the females examined, and in the males from 19 to 25”. From examination of 30 specimens I can see no reason for this division of the sexes. Also she says of the apical spines, “In the female there are three pairs of apical spines, the hind tibia and one very large unpaired spine on the inside; the male has four pairs of apical spines on the hind tibia” This statement is completely without foundation, both the male and female possessing four pairs of apical spines on the hind tibia in the type material and in the specimens collected on the 1954 Expedition. Miss Lysaght was trying to draw attention to a sexual dimorphism which does not exist in the spination of the legs.
The differences observed between my specimens and the specific descriptions given by Hutton and Miss Lysaght all point to variation within the species and, although they should be recorded because of this, they are of no real taxonomic value.
From examination of 30 specimens of N. serratum, the main specific features by which it can be easily recognised appear to be.
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(1) Constancy in shape of the subgenital plate in both male and female.
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(2) Hind femora inflated in mature male only.
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(3) Constancy in number of apical spines of the legs.
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(4) Fore femora, and fore and middle tarsi without spines.
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(5) Proximal segment of hind tarsus nearly as long as other three segments together.
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(6) Maxillary palps with third segment 0.625 length of fourth.
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(7) Metasternum bearing a median tubercle.
The species Novoplectron serratum is now redefined as follows:
Colour: Basic colour of body medium to dark fuscous with lateral portions of terga paler fuscous; pronotum, mesonotum and metanotum not margined, no pale line on the pro-

| Arith. Mean. | Std. Dev. | No. | ||
| Prolat. | Fore Femur | 0 - 0 | 0 - 0 | 25 - 25 |
| Retrolat. | 0 - 0 | 0 - 0 | 25 - 25 | |
| Prolat. | Fore Tibia Inf. | 2.3 - 2.3 | 0.55 - 0.55 | 25 - 25 |
| Retrolat. | 2.9 - 2.8 | 0.34 - 0 33 | 25 - 25 | |
| Prolat. | Fore Tarsus | 0 - 0 | 0 - 0 | 25 - 25 |
| Retrolat. | 0 - 0 | 0 - 0 | 25 - 25 | |
| Prolat. | Mid. Femur Inf. | 0.2 - 0.1 | 0.42 - 0.42 | 25 - 25 |
| Retrolat. | 0.2 - 0.1 | 0.46 - 0.38 | 25 - 25 | |
| Prolat. | Mid. Tibia Sup. | 1.7 - 1.9 | 0.64 - 0.81 | 25 - 25 |
| Retrolat. | 0.2 - 0.1 | 0.46 - 0.56 | 25 - 25 | |
| Prolat. | Mid. Tibia Sup. | 1.7 - 1.9 | 0.57 - 0.5 | 25 - 25 |
| Retrolat. | 0.2 - 0.2 | 0.57 - 0.5 | 25 - 25 | |
| Prolat. | Mid. Tibia Inf. | 2.4 - 2.5 | 0.35 - 0.54 | 25 - 25 |
| Retrolat. | 2.9 - 2.7 | 0 - 0 | 25-25 | |
| Prolat. | Mid. Tarsus | 0 - 0 | 0 - 0 | 25 - 25 |
| Retrolat. | 0 - 0 | 25 - 25 | ||
| Prolat. | Post. Femur Inf. | 12.9 - 12.5 | 2.33 - 2.06 | 24 - 20 |
| Retrolat. | 16.6 - 15.9 | 2.58 - 2.42 | 24 - 20 | |
| Prolat. | Post. Tibia Sup. | 19.8 - 19.3 | 2.17 - 1.42 | 24 - 19 |
| Retrolat. | 22.1 - 22.7 | 1.46 - 2.05 | 24 - 19 | |
| Prolat. | Post. Tarsus 1 Sup. | 1.8 - 1.8 | 0.54 - 0.37 | 24 - 19 |
| Retrolat. | 2.3 - 2 4 | 0.56 - 0.51 | 24 - 19 | |
| Prolat. | Post. Tarsus 2 Sup. | 0.9 - 1 | 0.03 - 0 | 24 - 19 |
| Retrolat. | 1 - 1 | 0 - 0 | 24 - 19 | |
| For list of abbreviations, see index at end of paper. |
notum; head medium to pale fuscous; antennae pale fuscous; femora and tibiae of all legs transversely banded with broad bands of deep ochrous, interspersed with narrow bands of light brown and pale ochrous; tarsi pale ochrous; ovipositor reddish-brown.
Body: Length up to 26 mm., average 24 mm. Ovipositor 0.66 to 0.75 as long as body. Antennae three times as long as body. Compound eyes 1.25 as long as broad. Cerci. Fig. 2 (C), 0.16 length of ovipositor. Bodies of male and female subequal.
Antennae: Flagella very long, slender, tapering, thick and almost touching at their bases; scape longer than broad. Sexual dimorphism very poorly developed, the antennae in the male being a little longer and stouter than those of the female. The flagellum in the male never bears spines.
Legs: Long and slender. Fore and middle legs subequal in length, with hind leg twice as long. Sexual dimorphism present in hind leg. In mature male femur is inflated and wide and deeply sulcate; in female only slightly rounded and not so deeply sulcate. Hind tibiae spindly, but more robust in male than female. Femora, tibiae and two proximal segments of hind tarsi armed with variable numbers of spines (see Table III). Spines on hind femora increase in size from proximal to distal end and are larger on prolateral than retrolateral margin. No spines occur on fore femora or fore and middle tarsi. Ratios of length of legs to length of body are: fore leg 1.07: 1, middle leg 1 1: 1; hind leg 2.12: 1.
Genitalia: Female: Supra-anal plate, Fig. 2 (SAP), lateral margin rounded, terminal margin slightly or more deeply emarginate, clothed with short golden setae. Subgenital plate, Fig. 1 (SGP), 9.5 times as broad as long, sparsely clothed with setae, distal margin widely emarginate. Male: Supra-anal plate, Fig. 4 (SPL), slightly concave laterally, slightly emarginate terminally, thickly clothed with setae. Subgenital plate (hypandrium), Fig. 3 (H), triangular, 0.75 as long as wide, sides spreading slightly proximally, concave laterally, tapering abruptly distally with a rounded apex, lanceolate distal portion thickly clothed with short setae on dorsal and ventral surfaces. Laterally the plate bears two styli, Figs. 3, 4 (S), thickly clothed with short setae; length of styli being 0.6 the length of sternite IX. The lanceolate distal portion of the hypandrium and the two styli are joined laterally by connective membrane. Parameres, Figs. 3, 4 (P), broad at base, as long as broad with prolateral edge thickly clothed with long setae. Pseudosternite 0.73 as long as broad, tapering to a point distally. Penis two-lobed, each lobe being nearly twice as long as broad. Paraprocts, Fig. 2 (PP), small, thickly clothed with long setae on distal margin; present in female, but absent in all male specimens examined.
Localities Chatham group—Pitt Island, coll J. J. Fougère; Mangere Island, South East Island, coll. E. G. Turbott, R. R. Forster and R. K. Dell; The Sisters, coll R. R. Forster and R. K. Dell In petrel burrows and under stones.
Types. Lectotype and Paratypes in Canterbury Museum Collection.
