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Genus Oiophysa Drake and Salmon, 1950.
Oiophysa cumberi sp. nov. Fig. 3.
Sub-brachypterous: Median length 2.41 mm (♂), 2.44 mm, 2.51 mm (♀); total length 2.53 mm (♂), 2.56 mm, 2.63 mm (♀); greatest width across tegmina 1.44 mm (♂), 1.44 mm, 1.52 mm (♀).
Head anteriorly deeply emarginate, broadly U-shaped, the median incision over one-third total length of head (0.35–0.37); anterior margin biarcuate, the right and left halves nearly evenly convex, extending anterior to level of front margin of eyes. Width across eyes 1.02–1.05 mm, median length 0.21–0.24 mm; ratio, width:length 4.94 (♂), 4.50, 4.33 (♀). Frontal membrane about 0.3 times as long in middle as total median length of head; anterior margin micropunctate and very shortly ciliate; posterior margin with a row of punctures, small and obscure mesially, larger and more distinct laterally. Crown with a low median carina extending on to frontal membrane; laterally and posteriorly rather coarsely punctate. Posterior margin nearly straight, carinate. Postero-lateral margins and eyes distant from paranota; the former very shortly serrate-ciliate, sinuate, more or less convex basally. Antennae not or scarcely extending laterally beyond mesial margin of eyes; apical segment 0.13–0.15 mm long. Eyes extending posteriorly beyond lateral margins of head. Rostrum reaching hind coxae.
Pronotum more than 3.5 times as wide across paranota as long in middle (♂ 1.36 mm, 0.37 mm; ♀ 1.44 mm: 0.37 mm, 1.45 mm: 0.39 mm). Median carina narrow in middle, not reaching posterior margin, before which it expands laterally for a short distance; in front expanding to form a triangular raised area with base along entire anterior margin. Disc strongly declivous on each side of carina and sub-circularly depressed, but raised anteriorly and posteriorly. A row of punctures along each side of carina; distinct punctures behind carina; postero-lateral angles of disc coarsely punctate. Anterior margin between paranota nearly straight; posterior margin broadly concave. Paranota raised anteriorly; lateral and postero-lateral margins nearly straight, the postero-lateral angles between them usually more or less acutely rounded (ca. 60°–70°). The main veins irregularly branched and anastomosed, dividing all or most areolae into areolets.
Scutellum triangular, anteriorly depressed in middle as in distincta.
Tegmina very similar to those of distincta in shape and venation. Costal margin bisinuate both from above and from side, concave and elevated before middle and again shortly before
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Fig. 1.—Outline of head, to show variations. a, b, Oiophysa distincta Woodward; c, d, O. cumberi sp. nov.
Fig. 2.—Oiophysa cumberi sp. nov., right paranotum. a, ♂; b, ♀.
apex, the subapical emargination being much the deeper, so that apices appear narrowly produced. Veins mesad of R less strongly elevated than in distincta. Cell 2 m nearly equal to apical radial cell. 9–11 costal cells; 4–5 radial cells; costal cells and the subcostal cell wide; 2 m much larger than subapical radial. Anterior width across tegmina greater than across middle (♂, 1.44 mm: 1.39 mm; ♀, 1.44 mm: 1.35 mm; 1.52 mm: 1.41 mm).
Abdomen relatively broader than in distincta, about half as wide again as long between anterior margin of sternum III and apex of sternum VII in female, of sternum VIII in male Segment III as in distincta. Female: Postero-lateral angles of tergum VIII rounded, but little produced. Posterior margin of sternum VII less prominently produced posteriorly in middle than in distincta but strongly and convexly produced ventrad. Venter of segment VII relatively broader than in either distincta or fuscata pendergrasti, about 2.6–2.7 times as wide at base as long in middle. Segment IX with outer ventral margin, viewed from side, shallowly concave before apex; posterior lateral extension broadly rounded as seen from side. Valvifers and valvulae similar to those of distincta, though smaller. Anal tube nearly cylindrical, tapering only slightly to base and apex; no pronounced dorsal swelling. Male: Tergum VIII similar to that of distincta, except that posterior margin nearly straight; 1.5 times as long as tergum VII. Venter of segment VIII, basal width 2.85 times median length, 2.1 times length of sternum VII. Pygophor 1.2 times as long as wide, proportionately shorter than in distincta, particularly the part basad of the lateral shoulders, which are more evenly rounded than in distincta or pendergrasti; sides basad of shoulders more concave than in distincta; posterior median process
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Fig. 4—Abdomen and terminalia, ♀. a, Oiophysa cumberi sp. nov., abdomen, ventral, sterna 3–7, b, O. cumberi, anal tube; c, O. distincta Woodward, anal tube; d, O. cumberi, left 2nd valvula, lateral, e, O. cumberi, 9th segment (lateral), with left 1st valvifer and 1st valvula and left 2nd valvifer and 3rd valvula.
a, apical lobe of 9th segment; v.1, 1st valvula; v.3, 3rd valvula; vf. 1, 1st valvifer; vf.2, 2nd valvifer; 9, 9th segment.
resembles that of distincta and differs from that of pendergrasti in being subacute at apex. Claspers (harpagones) smaller, thinner, more laterally flattened than in distincta; apex, seen from inner aspect, not truncate as in distincta, but rounded, though more narrowly so than in pendergrasti. Aedeagus similar to that of these two forms. Anal tube short, stout, ovate with both ends truncated, widest at basal quarter; sides more convex than in distincta; narrowing more abruptly at apex than in pendergrasti; dorsally swollen at base.
Colour: Ochraceous, varyingly infuscated, especially on scutellum, base of tegmina and costal veinlets, sometimes extensively melanic; membranes of head and paranota and at least costal and subcostal cells and apex of tegmina pale. Eyes reddish brown.
Localities. North Island: Holotype female, paratype female, 15.2.1957. Pohokura (Taupo-Napier Road, between Rangitaiki and Tarawera, mixed forest. ca 2,000ft). Allotype male, 15.2.1957, Kaimai Range (saddle, Tauranga-Matamata Road, mixed forest, 1,700ft). All collected R. A. Cumber.
Types. Holotype and allotype deposited in the collections of the Entomological Research Station, D.S.I.R., Nelson; paratype female at Entomology Division. D. S. I. R., Palmerston North.