Go to National Library of New Zealand Te Puna Mātauranga o Aotearoa
Volume 87, 1959
This text is also available in PDF
(807 KB) Opens in new window
– 29 –

Genus Pachyrhamma. Brunner, 1888

Pachyrhamma fusca n. sp. Plate 5, Figs. 3, 4, 5. Text-fig. 1, Figs. 1–5

1930 Pleioplectron cavernae Hutton, Karny, Ann. Nat.-Hist. Mus. Wien, 44: 182–185, Fig. 143.

Colour. Basic colour medium brown, with anterior and posterior borders of pronotum and posterior borders of mesonotum, metanotum and abdominal terga dark brown; femora of all legs medium brown with transverse ochreous bands at proximal ends, tibiae mid brown proximally, changing to ochreous distally, tarsi ochreous, antennae medium brown, ovipositor reddish-brown.

Body. Length up to 30 mm in male and 26 mm in female, average length 26 mm in male and 23 mm in female. Body sparsely clothed with setae. Ovipositor subequal with length of body. Antennae approximately seven times as long as body. Fastigium as high as long, rising abruptly, with base touching scape of antennae Maxillary palps with third and fourth segments subequal in length.

Antennae. Scape about four times as large as pedicel, which is narrower than scape, but broader than other segments, third segment on dorsal aspect narrower than pedicel and twice as long, and on ventral aspect half as long again; from fourth segment onwards segments unequal in length, although steadily decreasing in size; all segments thickly clothed with short setae. Sexual dimorphism present in antennae, male possessing longer, stouter antennae than female, middle portion of flagellum in male armed with a number of short, blunt dorsal spines (Plate 18, Fig. 5), each spine borne on a swelling on the upper part of its segment, variability in number of spines present is common, but is most frequently nine, female never possesses spines.

Legs. Fore and middle legs subequal in length, with hind leg approximately twice length of fore and middle legs. Sexual dimorphism is shown by fore and middle legs of female being 0.75 as long as male, and hind legs of female 0.64 as long as male. Femora, tibiae and proximal two segments of hind tarsi armed with variable numbers of spines (Table 1). No spines occur on fore or middle femora and tarsi. Length of proximal segment of hind tarsus greater than other three together in male, subequal with other three in female. Ratios of length of legs to length of body: Fore leg, male, 2. 1:1; female, 1.8:1. Middle leg, male, 2.1:1; female, 1.8:1. Hind leg, male, 4.5:1;, female, 3.3:1.

Genitalia. Female: Suranal plate, Fig. 1 (SAP), concave laterally, rounded and notched medianly distally; distal margin clothed with short setae. Subgenital plate, Fig. 2 (SGP), deeply notched distally and sparsely clothed with short setae. Male: Suranal plate, Fig. 4 (SPL), slightly convex laterally, straight distally, bearing two groups of setae. Subgenital plate, Fig. 3 (H), triangulate, as wide as long, sides spreading slightly proximally, tapering to concave distally with a rounded apex, glabrous on dorsal side, but with apical protuberance on ventral surface thickly clothed with short setae. Two styli, Figs. 3, 4 (S), thickly clothed with short setae, length of styli being 0.2 length of sternite IX (S IX). Subgenital plate covers genitalia Parameres, Fig. 5 (P), attenuated, broad at base and tapering to a point, 2.5 longer than broad, prolateral margin thickly clothed with long setae, rest of paramere clothed with short setae. Pseudosternite, Fig. 5 (PD), 1.1 wider than long, tapering to a point distally Penis, Fig. 5 (PN), two-lobed, each lobe 1.5 longer than broad Paraprocts, Figs. 3, 5 (PP), elongate, 1.6 longer than broad.

Locality. Gold-mining tunnels, Thames (type locality), coll. A. M. Richards, 1958; tunnels Coromandel, coll. Reischek, 1891.

Types. Holotype male, Allotype female and Paratype male and female in Auckland Institute and Museum Collection. Paratype male and female in British Museum (Nat. Hist.) Collection. Paratype male and female in Vienna Museum Collection.

– 30 –

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

Table. I — Variability in Number of Linear Spines on the Legs of 21 Specimens of Pachyrhamma Fusca n. sp.
Arith. Mean Std. Dev Range
L. R. L. R. L. R.
Fore Femur Pro. 0 0
Inf. Retro. 0 0
Fore Tibia Pro. 3 3
Inf. Retro. 3 3
Fore Tarsus Pro. 0 0
Retro. 0 0
Mid Femur Pro. 0 0
Inf. Retro. 0 0
Mid Tibia Pro. 2.9 3.1 0.8 0.9 1–4 1–5
Sup Retro. 1.3 1.3 0.7 0.6 0–3 0–3
Mid Tibia Pro. 3 3
Inf. Retro. 3 3
Mid Tarsus Pro. 0 0
Retro. 0 0
Hind Femur Pro. 8.4 8.4 1.3 0.8 7–12 7–10
Inf. Retro. 2.7 2.7 0.7 0.8 2–4 2–4
Hind Tibia Pro. 31.8 31.9 3.3 3.7 26–38 25–37
Sup Retro. 34.6 34.4 2.4 2.9 30–38 30–39
Hind Tarsus Pro. 1.9 1.8 0.6 0.4 1(4), 2(16), 4 1(5), 2(15)
1 Sup Pro. 1.7 1.8 0.9 0.7 0–4 0–3
Hind Tarsus Retro. 0.1 0 0(20), 1 0(19), 1
2 Sup Retro. 0.1 0 0(20), 1 0(20)

Figures in brackets are number of specimens.

Pachyrhamma fusca is most closely related to P. waitomoensis, but differs from it in.

1 The presence of spines on antennae of mature male.

2 A greater number of linear spines on hind femora.

From the same tunnels from which P. fusca was collected, twenty-five specimens of another species, also belonging to the genus Pachyrhamma, were collected. They are considered to be new and are described as follows:

Pachyrhamma uncata n. sp. Plate 5, Figs. 1, 2 Text-fig. 2, Figs. 1–5.

Colour. Basic colour deep ochreous, with pronotum, mesonotum, metanotum and abdominal terga irregularly mottled with dark brown and medium brown. Anterior margin of pronotum and lateral margins of pronotum, mesonotum and metanotum ochreous; femora and tibiae of all legs transversely banded with broad bands of light brown interspersed with narrow bands of ochreous; tarsi pale ochreous, antennae light brown; ovipositor deep reddish-brown.

Body. Length up to 28 mm in male and 25 mm in female; average length 24 mm in male and 23 mm in female. Body sparsely clothed with setae. Ovipositor subequal in length with body. Antennae approximately 5.7 times as long as body. Fastigium rising abruptly, as high as long Maxillary palps with third and fourth joints subequal in length.

Antennae. Scape about four times as large as pedicel, which is narrower than scape, but broader than other segments; third segment on dorsal and vential aspects narrower than pedicel and twice as long. From fourth segment onwards segments unequal in length, although steadily decreasing in size, all segments thickly clothed with short setae. Sexual dimorphism poorly developed, antennae of male slightly stouter and longer than those of female; no spines present on flagellum of male or female.

Legs. Fore and middle legs subequal in length, with hind leg approximately twice length of fore and middle legs. Sexual dimorphism is shown by fore and middle legs of female

Picture icon

Fig. 1.—Pachyrhamma uncata male, lateral view
Fig. 2.—Pachyrhamma uncata female lateral view
Fig. 3.—Pachyrhamma fusca male lateral view
Fig. 4.—Pachyrhamma fusca female lateral view
Fig. 5.—Portion of antenna of adult male Pachyrhamma fusca showing blunt doisal spines
Photos J. W. Endt

– 31 –
Picture icon

Text-fig. 2Pachyrhamma uncata n. sp. Fig. 1—Female genitalia, dorsal view Fig. 2—Female genitalia, ventral view Fig. 3—Male genitalia, dorsal view Fig. 4—Male genitalia, ventral view, subgenital plate in place Fig. 5—Male genitalia, ventral view, subgenital plate removed to expose structures beneath

– 32 –

being 0.7 as long as male, and hind legs of female 0.6 as long as male Femora, tibiae and two proximal segments of hind tarsi armed with vanable numbers of spines (Table II). No spines occur on fore or middle tarsi. Length of proximal segment of hind tarsus subequal, with length of other three segments together. Sexual dimorphism is shown in the spination of the hind femora, the more proximal portion of the prolateral margin of the male bearing three laterally directed, enlarged hooked spines, while in the female they are smaller and posteriorly directed. Ratios of length of legs to length of body: Fore leg, male 2.3:1, female 2:1 Middle leg, male 2.3:1, female 1.9:1 Hind leg, male 5:1, female 3.6:1.

Genitalia. Female: Suranal plate, Fig. 1 (SAP), convex laterally, straight distally distal margin clothed with two groups of setae. Subgenital plate, Fig. 2 (SGP), concave laterally with V-shaped notch distally; sparsely clothed with short setae. Male: Suranal plate, Fig. 3 (SPL), with straight margins, the lateral ones tapering distally; distal margin with a fringe of short setae. Subgenital plate, Fig. 4 (H), triangular, 1.1 wider than long, sides spreading slightly proximally, tapering to concave distally with a rounded apex, glabrous on dorsal side, but with apical protuberance on ventral surface thickly clothed with short setae. Two styli, Figs. 3, 4 (S), thickly clothed with short setae, length of styli being 0.3 length of sternite IX (SIX). Subgenital plate covers genitalia. Parameres, Fig. 5 (P), attenuated,

[The section below cannot be correctly rendered as it contains complex formatting. See the image of the page for a more accurate rendering.]

Table. II—Variability in Number of Linear Spines on the Legs of 25 Specimens of Pachyrhamma Uncata n. sp.
Arith. Mean Std. Dev Range
L. R. L. R. L. R.
Fore Femur Pro. 2.7 3 1.7 1.9 0–6 0–7
Inf. Retro. 0.04 0.04 0(24), 1 0(24), 1
Fore Tibia Pro. 3 3
Inf. Retro. 3 3
Fore Tarsus Pro. 0 0
Retro. 0 0
Mid Femur Pro. 0.5 0.4 0.9 0.9 0–3 0–3
Inf. Retro. 1.5 1.8 1.4 1.8 0–5 0–5
Mid Tibia Pro. 3.8 3.9 1.3 1.4 2–7 2–7
Sup Retro. 2.8 2.6 1.2 1.2 1–6 1–5
Mid Tibia Pro. 3 3
Inf. Retro. 3 3
Mid Tarsus Pro. 0 0
Retro. 0 0
Hind Femur Pro. 6.4 6.2 1.0 0.7 4–8 5–8
Inf. Retro. 18.6 17.4 6.2 4.9 7–32 9–29
Hind Tibia Pro. 35.3 34.7 3.1 3.7 27–40 26–42
Sup Retro. 37.6 38.3 3.7 3.2 30–43 32–45
Hind Tarsus Pro. 3 3.2 0.8 0.8 2–4 2–4
1 Sup Retro. 3 3 0.7 1.0 2–4 1–6
Hind Tarsus Pro. 0.4 0.6 0(15), 1(10) 0(11), 1(14)
2 Sup Retro. 0.6 0.5 0(10), 1(15) 0(12), 1(13)

Figures in brackets are number of specimens. broad at base and tapering to a point, 1.2 broader than long, prolateial margin thickly clothed with long setae, rest of paramere clothed with short setae. Pseudosternite, Fig. 5 (PD), 1.1 wider than long, tapering to a point distally, Penis, Fig. 5 (PN), two-lobed, each lobe 1.8 longer than broad. Paraprocts, Figs. 4, 5 (PP), elongate, 1.8 longer than broad.

Locality. Gold-mining tunnels, Thames (type locality), coll. A. M. Richards.

Types. Holotype male, Allotype female and Paratype male and female in Auckland Institute and Museum Collection. Paratype male and female in British Museum (Nat. Hist.) Collection. Paratype male and female in Dominion Museum Collection.

– 33 –

Pachyrhamma uncata is most closely related to P. acanthocera, but differs from it in.

  • 1. The absence of spines on antennae of mature male.

  • 2. Smaller number of linear spines on middle and hind femora.

  • 3. Presence of enlarged hooked spines on prolateral margin of hind femora of adult male.